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<mods:title id="24ED0086DC5231BDF0E70185CC919D76">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<treatment id="03D7FF1CFF0CFF2AFF48F9983373FA0C" ID-DOI="http://doi.org/10.5281/zenodo.3682429" ID-GBIF-Taxon="162246880" ID-Zenodo-Dep="3682429" LSID="urn:lsid:plazi:treatment:03D7FF1CFF0CFF2AFF48F9983373FA0C" httpUri="http://treatment.plazi.org/id/03D7FF1CFF0CFF2AFF48F9983373FA0C" lastPageNumber="198" pageId="199" pageNumber="198">
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<taxonomicName id="4C7E3589FF0CFF2AFF48F9983309FCFE" authority="(Linnaeus, 1767)" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[192,629,758,782]" class="Insecta" family="Dynastidae" genus="Coelosis" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="198" phylum="Arthropoda" rank="species" species="biloba">
<emphasis id="B90A9218FF0CFF2AFF48F9983309FCFE" bold="true" box="[192,629,758,782]" pageId="199" pageNumber="198">
<emphasis id="B90A9218FF0CFF2AFF48F99830F0FCFE" bold="true" box="[192,396,758,782]" italics="true" pageId="199" pageNumber="198">Coelosis biloba</emphasis>
(
<bibRefCitation id="EFEF33FBFF0CFF2AFE14F9983310FCFE" author="Linnaeus" box="[412,620,758,782]" firstAuthor="Linnaeus" pageId="199" pageNumber="198" pagination="533 - 1327" refId="ref212835" refString="Linnaeus, C. 1767. Systema naturae. Editio duodecima reformata. Tom. I. Pars II. Laur. Salvii; Stockholm. p. 533 - 1327." type="book chapter" year="1767">Linnaeus, 1767</bibRefCitation>
)
</emphasis>
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<treatmentCitation id="0ADF681BFF0CFF2AFF60F87333E2FCC3" author="Linnaeus" box="[232,670,797,819]" page="544" pageId="199" pageNumber="198" year="1767">
<taxonomicName id="4C7E3589FF0CFF2AFF60F87333E2FCC3" ID-CoL="79VT4" authority="Linnaeus 1767: 544" authorityName="Linnaeus" authorityPageNumber="544" authorityYear="1767" box="[232,670,797,819]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="198" phylum="Arthropoda" rank="species" species="bilobus">
<emphasis id="B90A9218FF0CFF2AFF60F87330BCFCC2" box="[232,448,797,818]" italics="true" pageId="199" pageNumber="198">Scarabaeus bilobus</emphasis>
<bibRefCitation id="EFEF33FBFF0CFF2AFE4EF87333E2FCC3" author="Linnaeus" box="[454,670,797,819]" firstAuthor="Linnaeus" pageId="199" pageNumber="198" pagination="533 - 1327" refId="ref212835" refString="Linnaeus, C. 1767. Systema naturae. Editio duodecima reformata. Tom. I. Pars II. Laur. Salvii; Stockholm. p. 533 - 1327." type="book chapter" year="1767">Linnaeus 1767: 544</bibRefCitation>
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(none -
<typeStatus id="54C5F0A8FF0CFF2AFD7EF87332E8FCC3" box="[758,916,797,819]" pageId="199" pageNumber="198" type="lectotype">LECTOTYPE</typeStatus>
)
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</paragraph>
</subSubSection>
<subSubSection id="C3641D81FF0CFF2AFF48F82D3373FA0C" pageId="199" pageNumber="198" type="distribution">
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<emphasis id="B90A9218FF0CFF2AFF48F82D3099FCAB" bold="true" box="[192,485,835,859]" pageId="199" pageNumber="198">
Distribution.
<collectingCountry id="F3690E9AFF0CFF2AFEFDF82D309CFCAB" box="[373,480,835,859]" name="Guyana" pageId="199" pageNumber="198">Guyana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF0CFF2AFE63F82D33B1FCAB" author="Erichson, W. F." box="[491,717,835,860]" editor="R. Schomburgk" journalOrPublisher="J. J. Weber; Leipzig, Germany" pageId="199" pageNumber="198" pagination="553 - 1260" refId="ref201522" refString="Erichson, W. F. 1848. Insecten. p. 553 - 617. In: R. Schomburgk. Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil. J. J. Weber; Leipzig, Germany. p. 553 - 1260." title="Insecten" type="book chapter" volumeTitle="Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil" year="1848">Erichson 1848: 561</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0CFF2AFD73F82D32C4FCAB" box="[763,952,835,859]" class="Insecta" family="Dynastidae" genus="Coelosis" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="198" phylum="Arthropoda" rank="species" species="bilobus">
<emphasis id="B90A9218FF0CFF2AFD73F82D32C4FCAB" box="[763,952,835,859]" italics="true" pageId="199" pageNumber="198">Coelosis bilobus</emphasis>
</taxonomicName>
).
<emphasis id="B90A9218FF0CFF2AFC44F82D3525FCAB" bold="true" box="[972,1113,835,859]" pageId="199" pageNumber="198">
<collectingCountry id="F3690E9AFF0CFF2AFC44F82D352EFCAB" box="[972,1106,835,859]" name="Suriname" pageId="199" pageNumber="198">Suriname</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF0CFF2AFBD6F82D3457FCAB" author="Endrodi, S." box="[1118,1323,835,859]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="199" pageNumber="198" pagination="9 - 174" part="29" refId="ref201147" refString="Endrodi, S. 1976. Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae). Folia Entomologica Hungarica (N. S.) 29 (2): 9 - 174." title="Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae)" type="journal article" year="1976">Endrödi 1976: 46</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFABCF82D300BFC8B" author="Dechambre, R. - P." journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="199" pageNumber="198" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFE0DF80D3259FC8B" author="Iannuzzi, L. &amp; R. C. Marinoni" box="[389,805,867,891]" journalOrPublisher="Revista Brasileira de Zoologia" pageId="199" pageNumber="198" pagination="93 - 121" part="12" refId="ref209058" refString="Iannuzzi, L., and R. C. Marinoni. 1995. Revisao do genero neotropical Coelosis Hope (Coleoptera, Scarabaeidae, Dynastinae). Revista Brasileira de Zoologia 12 (1): 93 - 121." title="Revisao do genero neotropical Coelosis Hope (Coleoptera, Scarabaeidae, Dynastinae)" type="journal article" year="1995">Iannuzzi and Marinoni 1995: 115</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0CFF2AFCD3F80A3519FC8B" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[859,1125,867,892]" class="Insecta" family="Dynastidae" genus="Coelosis" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="198" phylum="Arthropoda" rank="species" species="biloba" subGenus="Eucoelosis">
<emphasis id="B90A9218FF0CFF2AFCD3F80A320AFC8B" box="[859,886,868,891]" italics="true" pageId="199" pageNumber="198">C.</emphasis>
(
<emphasis id="B90A9218FF0CFF2AFC00F80D357BFC8B" box="[904,1031,867,891]" italics="true" pageId="199" pageNumber="198">Eucoelosis</emphasis>
)
<emphasis id="B90A9218FF0CFF2AFB91F80D3519FC8B" box="[1049,1125,867,891]" italics="true" pageId="199" pageNumber="198">biloba</emphasis>
</taxonomicName>
); MHCN, NZCS; RMNH.
<emphasis id="B90A9218FF0CFF2AFF48F8ED30EBFC6B" bold="true" box="[192,407,899,923]" pageId="199" pageNumber="198">
<collectingCountry id="F3690E9AFF0CFF2AFF48F8ED30EEFC6B" box="[192,402,899,923]" name="French Guiana" pageId="199" pageNumber="198">French Guiana</collectingCountry>
:
</emphasis>
H. W.
<bibRefCitation id="EFEF33FBFF0CFF2AFE63F8EA33CCFC6B" author="Bates, H. W." bookContentInfo="432 p." box="[491,688,899,924]" journalOrPublisher="Frederick Ducane Godman and Osbert Salvin; London" pageId="199" pageNumber="198" pagination="161 - 336" refId="ref193515" refString="Bates, H. W. 1888. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. p. 161 - 336. In: Biologia Centrali-Americana. Frederick Ducane Godman and Osbert Salvin; London. 432 p." title="Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia" type="book chapter" volumeTitle="Biologia Centrali-Americana" year="1888">Bates 1888: 326</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFD34F8ED32AFFC6B" author="Blackwelder, R. E." box="[700,979,899,924]" journalOrPublisher="United States National Museum Bulletin" pageId="199" pageNumber="198" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFC57F8ED35CFFC6B" author="Endrodi, S." box="[991,1203,899,923]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="199" pageNumber="198" pagination="9 - 174" part="29" refId="ref201147" refString="Endrodi, S. 1976. Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae). Folia Entomologica Hungarica (N. S.) 29 (2): 9 - 174." title="Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae)" type="journal article" year="1976">Endrödi 1976: 46</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFB37F8ED3193FC4B" author="Dechambre, R. - P." journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="199" pageNumber="198" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFF75F8CD33E0FC4B" author="Iannuzzi, L. &amp; R. C. Marinoni" box="[253,668,931,955]" journalOrPublisher="Revista Brasileira de Zoologia" pageId="199" pageNumber="198" pagination="93 - 121" part="12" refId="ref209058" refString="Iannuzzi, L., and R. C. Marinoni. 1995. Revisao do genero neotropical Coelosis Hope (Coleoptera, Scarabaeidae, Dynastinae). Revista Brasileira de Zoologia 12 (1): 93 - 121." title="Revisao do genero neotropical Coelosis Hope (Coleoptera, Scarabaeidae, Dynastinae)" type="journal article" year="1995">Iannuzzi and Marinoni 1995: 115</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0CFF2AFD5AF8CA32A7FC4B" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[722,987,931,956]" class="Insecta" family="Dynastidae" genus="Coelosis" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="198" phylum="Arthropoda" rank="species" species="biloba" subGenus="Eucoelosis">
<emphasis id="B90A9218FF0CFF2AFD5AF8CA3391FC4B" box="[722,749,932,955]" italics="true" pageId="199" pageNumber="198">C.</emphasis>
(
<emphasis id="B90A9218FF0CFF2AFD76F8CD3201FC4B" box="[766,893,931,955]" italics="true" pageId="199" pageNumber="198">Eucoelosis</emphasis>
)
<emphasis id="B90A9218FF0CFF2AFC07F8CD32A7FC4B" box="[911,987,931,955]" italics="true" pageId="199" pageNumber="198">biloba</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0CFF2AFC7CF8CD346FFC4C" author="Brule, S. &amp; J. Touroult &amp; P. - H. Dalens" bookContentInfo="120 p." box="[1012,1299,931,956]" journalOrPublisher="Rapport de la Societe entomologique Antilles-Guyane, SEAG, ONF" pageId="199" pageNumber="198" refId="ref195373" refString="Brule, S., J. Touroult, and P. - H. Dalens (coord.). 2011 a. Resultats de l'inventaire entomologique du site de Saut Parare, reserve des Nouragues (Guyane), 2009 - 2010. Rapport de la Societe entomologique Antilles-Guyane, SEAG, ONF. 120 p. + annexes." title="Resultats de l'inventaire entomologique du site de Saut Parare, reserve des Nouragues (Guyane)" type="book" year="2011">Brûlé et al. 2011a: 191</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF0CFF2AFAA9F8CA34E7FC4C" author="Brule, S. &amp; J. Touroult &amp; E. Poirier &amp; P. - H. Dalens" bookContentInfo="144 p." box="[1313,1435,932,956]" journalOrPublisher="SEAG, PAG" pageId="199" pageNumber="198" refId="ref195498" refString="Brule, S., J. Touroult, E. Poirier, and P. - H. Dalens. 2014. Resultats de l'etude-inventaire entomologique du site de la Montagne Pelee, Saul (Guyane), 2010 - 2012. Rapport de la Societe entomologique Antilles-Guyane. SEAG, PAG. 144 p. + annexes." title="Resultats de l'etude-inventaire entomologique du site de la Montagne Pelee, Saul (Guyane), 2010 - 2012. Rapport de la Societe entomologique Antilles-Guyane" type="book" year="2014">2014: 181</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFF48F8AA30D9FC2C" author="Duranton, M." box="[192,421,964,988]" journalOrPublisher="Coleopteres, numero special" pageId="199" pageNumber="198" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton 2011: 21</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFE38F8AD3303FC2B" author="Ponchel" box="[432,639,963,988]" firstAuthor="Ponchel" pageId="199" pageNumber="198" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011: 61</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFD01F8AD32CCFC2C" author="Brule, S. &amp; P. - H. Dalens" bookContentInfo="23 p." box="[649,944,963,988]" journalOrPublisher="Rapport de la Societe entomologique Antilles-Guyane" pageId="199" pageNumber="198" refId="ref195220" refString="Brule, S., and P. - H. Dalens. 2012 a. Etude et inventaire entomologique du Mont Itoupe. Mission 2010 pour le compte du Parc Amazonien de Guyane. Rapport de la Societe entomologique Antilles-Guyane. 23 p. + annexes." title="Etude et inventaire entomologique du Mont Itoupe. Mission 2010 pour le compte du Parc Amazonien de Guyane" type="book" year="2012">Brûlé and Dalens 2012a</bibRefCitation>
: annex 4: 18;
<bibRefCitation id="EFEF33FBFF0CFF2AFBD9F8AD344AFC2C" author="Dupuis, F." box="[1105,1334,963,988]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="199" pageNumber="198" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b: 110</bibRefCitation>
; MHCN; NZCS.
<emphasis id="B90A9218FF0CFF2AFE91F88D30D0FC0B" bold="true" box="[281,428,995,1019]" pageId="199" pageNumber="198">“Guianas”:</emphasis>
<bibRefCitation id="EFEF33FBFF0CFF2AFE3CF88A33F8FC0B" author="Arrow, G. J." bookContentInfo="124 p." box="[436,644,995,1020]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="199" pageNumber="198" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 35</bibRefCitation>
.
<emphasis id="B90A9218FF0CFF2AFD18F88D324FFC0B" bold="true" box="[656,819,995,1019]" pageId="199" pageNumber="198">
<collectingCountry id="F3690E9AFF0CFF2AFD18F88D3398FC0B" box="[656,740,995,1019]" name="Brazil" pageId="199" pageNumber="198">Brazil</collectingCountry>
(NA):
</emphasis>
<bibRefCitation id="EFEF33FBFF0CFF2AFCB3F88A3575FC0B" author="Arrow, G. J." bookContentInfo="124 p." box="[827,1033,995,1020]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="199" pageNumber="198" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 35</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFB9EF88D344CFC0B" author="Blackwelder, R. E." box="[1046,1328,995,1020]" journalOrPublisher="United States National Museum Bulletin" pageId="199" pageNumber="198" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFAB5F88D3050FBEB" author="Endrodi, S." journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="199" pageNumber="198" pagination="9 - 174" part="29" refId="ref201147" refString="Endrodi, S. 1976. Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae). Folia Entomologica Hungarica (N. S.) 29 (2): 9 - 174." title="Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae)" type="journal article" year="1976">Endrödi 1976: 46</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFEB3FF6D3324FBEB" author="Dechambre, R. - P." box="[315,600,1027,1052]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="199" pageNumber="198" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFDEFFF6D3575FBEB" author="Iannuzzi, L. &amp; R. C. Marinoni" box="[615,1033,1027,1051]" journalOrPublisher="Revista Brasileira de Zoologia" pageId="199" pageNumber="198" pagination="93 - 121" part="12" refId="ref209058" refString="Iannuzzi, L., and R. C. Marinoni. 1995. Revisao do genero neotropical Coelosis Hope (Coleoptera, Scarabaeidae, Dynastinae). Revista Brasileira de Zoologia 12 (1): 93 - 121." title="Revisao do genero neotropical Coelosis Hope (Coleoptera, Scarabaeidae, Dynastinae)" type="journal article" year="1995">Iannuzzi and Marinoni 1995: 115</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0CFF2AFBC9FF6A3430FBEB" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[1089,1356,1027,1052]" class="Insecta" family="Dynastidae" genus="Coelosis" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="198" phylum="Arthropoda" rank="species" species="biloba" subGenus="Eucoelosis">
<emphasis id="B90A9218FF0CFF2AFBC9FF6A3520FBEB" box="[1089,1116,1028,1051]" italics="true" pageId="199" pageNumber="198">C.</emphasis>
(
<emphasis id="B90A9218FF0CFF2AFBE6FF6D3591FBEB" box="[1134,1261,1027,1051]" italics="true" pageId="199" pageNumber="198">Eucoelosis</emphasis>
)
<emphasis id="B90A9218FF0CFF2AFA88FF6D3430FBEB" box="[1280,1356,1027,1051]" italics="true" pageId="199" pageNumber="198">biloba</emphasis>
</taxonomicName>
) (NA);
<bibRefCitation id="EFEF33FBFF0CFF2AFF48FF4D332EFBCB" author="Andreazze, R. &amp; C. R. V. da Fonseca" box="[192,594,1059,1084]" journalOrPublisher="Acta Amazonica" pageId="199" pageNumber="198" pagination="59 - 66" part="28" refId="ref190842" refString="Andreazze, R., and C. R. V. da Fonseca. 1998. Dinastineos (Coleoptera, Scarabaeoidea, Melolonthidae) em uma area de terra firme na Amazonia central, Brasil. Acta Amazonica 28 (1): 59 - 66." title="Dinastineos (Coleoptera, Scarabaeoidea, Melolonthidae) em uma area de terra firme na Amazonia central, Brasil" type="journal article" year="1998">Andreazze and Fonseca 1998: 62</bibRefCitation>
(NA);
<bibRefCitation id="EFEF33FBFF0CFF2AFD2CFF4E3280FBCC" author="Gasca-Alvarez, H. J. &amp; N. O. Aguiar" box="[676,1020,1056,1084]" journalOrPublisher="Acta Zoologica Mexicana (N. S.)" pageId="199" pageNumber="198" pagination="221 - 224" part="24" refId="ref203674" refString="Gasca-Alvarez, H. J., and N. O. Aguiar. 2008. Primer registro de Amblyodus taurus Westwood, 1878 (Scarabaeidae: Dynastinae: Phileurini) en Brasil. Acta Zoologica Mexicana (N. S.) 24 (3): 221 - 224." title="Primer registro de Amblyodus taurus Westwood, 1878 (Scarabaeidae: Dynastinae: Phileurini) en Brasil" type="journal article" year="2008">Gasca-Álvarez et al. 2008: 8</bibRefCitation>
(NA);
<bibRefCitation id="EFEF33FBFF0CFF2AFBC5FF4E3048FBAC" author="Gasca-Alvarez, H. J. &amp; C. R. V. da Fonseca" journalOrPublisher="Acta Amazonica" pageId="199" pageNumber="198" pagination="719 - 722" part="39" refId="ref203726" refString="Gasca-Alvarez, H. J., and C. R. V. da Fonseca. 2009. Oryctini (Coleoptera: Scarabaeidae: Dynastinae) da colecao de invertebrados do Instituto Nacional de Pesquisas da Amazonia (INPA), Manaus, Amazonas, Brasil. Acta Amazonica 39 (3): 719 - 722." title="Oryctini (Coleoptera: Scarabaeidae: Dynastinae) da colecao de invertebrados do Instituto Nacional de Pesquisas da Amazonia (INPA), Manaus, Amazonas, Brasil" type="journal article" year="2009">Gasca-Álvarez and Fonseca 2009: 721</bibRefCitation>
(NA);
<bibRefCitation id="EFEF33FBFF0CFF2AFE09FF2E323DFBAC" author="Gasca-Alvarez, H. J. &amp; B. C. Ratcliffe" box="[385,833,1088,1116]" journalOrPublisher="Zootaxa" pageId="199" pageNumber="198" pagination="21 - 40" part="3090" refId="ref203834" refString="Gasca-Alvarez, H. J., and B. C. Ratcliffe. 2011. An annoted checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms. Zootaxa 3090: 21 - 40." title="An annoted checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms" type="journal article" year="2011">Gasca-Álvarez and Ratcliffe 2011: 24</bibRefCitation>
.
<emphasis id="B90A9218FF0CFF2AFCC3FF2D329CFBAB" bold="true" box="[843,992,1091,1115]" pageId="199" pageNumber="198">
<collectingCountry id="F3690E9AFF0CFF2AFCC3FF2D32A7FBAB" box="[843,987,1091,1115]" name="Venezuela" pageId="199" pageNumber="198">Venezuela</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF0CFF2AFC6DFF2D35CAFBAB" author="Endrodi, S." box="[997,1206,1091,1115]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="199" pageNumber="198" pagination="9 - 174" part="29" refId="ref201147" refString="Endrodi, S. 1976. Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae). Folia Entomologica Hungarica (N. S.) 29 (2): 9 - 174." title="Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae)" type="journal article" year="1976">Endrödi 1976: 46</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFB48FF2D3193FB8B" author="Dechambre, R. - P." journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="199" pageNumber="198" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF0CFF2AFF72FF0D33F2FB8B" author="Iannuzzi, L. &amp; R. C. Marinoni" box="[250,654,1123,1147]" journalOrPublisher="Revista Brasileira de Zoologia" pageId="199" pageNumber="198" pagination="93 - 121" part="12" refId="ref209058" refString="Iannuzzi, L., and R. C. Marinoni. 1995. Revisao do genero neotropical Coelosis Hope (Coleoptera, Scarabaeidae, Dynastinae). Revista Brasileira de Zoologia 12 (1): 93 - 121." title="Revisao do genero neotropical Coelosis Hope (Coleoptera, Scarabaeidae, Dynastinae)" type="journal article" year="1995">Iannuzzi and Marinoni 1995: 115</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF0CFF2AFD37FF0A32BEFB8B" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[703,962,1123,1148]" class="Insecta" family="Dynastidae" genus="Coelosis" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="198" phylum="Arthropoda" rank="species" species="biloba" subGenus="Eucoelosis">
<emphasis id="B90A9218FF0CFF2AFD37FF0A33A5FB8B" box="[703,729,1124,1147]" italics="true" pageId="199" pageNumber="198">C.</emphasis>
(
<emphasis id="B90A9218FF0CFF2AFD60FF0D321BFB8B" box="[744,871,1123,1147]" italics="true" pageId="199" pageNumber="198">Eucoelosis</emphasis>
)
<emphasis id="B90A9218FF0CFF2AFCFEFF0D32BEFB8B" box="[886,962,1123,1147]" italics="true" pageId="199" pageNumber="198">biloba</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0CFF2AFC50FF0E34E0FB8C" author="Gasca-Alvarez, H. J. &amp; B. C. Ratcliffe" box="[984,1436,1120,1148]" journalOrPublisher="Zootaxa" pageId="199" pageNumber="198" pagination="21 - 40" part="3090" refId="ref203834" refString="Gasca-Alvarez, H. J., and B. C. Ratcliffe. 2011. An annoted checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms. Zootaxa 3090: 21 - 40." title="An annoted checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms" type="journal article" year="2011">Gasca-Álvarez and Ratcliffe 2011: 24</bibRefCitation>
.
<emphasis id="B90A9218FF0CFF2AFF48FFED3064FB6B" bold="true" box="[192,280,1155,1179]" pageId="199" pageNumber="198">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF0CFF2AFE97FFED3373FB6B" author="Linnaeus" box="[287,527,1155,1179]" firstAuthor="Linnaeus" pageId="199" pageNumber="198" pagination="533 - 1327" refId="ref212835" refString="Linnaeus, C. 1767. Systema naturae. Editio duodecima reformata. Tom. I. Pars II. Laur. Salvii; Stockholm. p. 533 - 1327." type="book chapter" year="1767">Linnaeus 1767: 544</bibRefCitation>
(southern Europe); H. W.
<bibRefCitation id="EFEF33FBFF0CFF2AFCDAFFEA3568FB6B" author="Bates, H. W." bookContentInfo="432 p." box="[850,1044,1155,1180]" journalOrPublisher="Frederick Ducane Godman and Osbert Salvin; London" pageId="199" pageNumber="198" pagination="161 - 336" refId="ref193515" refString="Bates, H. W. 1888. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. p. 161 - 336. In: Biologia Centrali-Americana. Frederick Ducane Godman and Osbert Salvin; London. 432 p." title="Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia" type="book chapter" volumeTitle="Biologia Centrali-Americana" year="1888">Bates 1888: 326</bibRefCitation>
(Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama, South America, West Indies);
<bibRefCitation id="EFEF33FBFF0CFF2AFB8AFFCA35B9FB4B" author="Arrow, G. J." bookContentInfo="124 p." box="[1026,1221,1187,1212]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="199" pageNumber="198" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 35</bibRefCitation>
(Mexico, Panama, Trinidad and Tobago);
<bibRefCitation id="EFEF33FBFF0CFF2AFE55FFAD338AFB2B" author="Blackwelder, R. E." box="[477,758,1219,1244]" journalOrPublisher="United States National Museum Bulletin" pageId="199" pageNumber="198" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
(Argentina, Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama, Trinidad and Tobago, West Indies);
<bibRefCitation id="EFEF33FBFF0CFF2AFC51FF8D35D4FB0B" author="Endrodi, S." box="[985,1192,1251,1275]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="199" pageNumber="198" pagination="9 - 174" part="29" refId="ref201147" refString="Endrodi, S. 1976. Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae). Folia Entomologica Hungarica (N. S.) 29 (2): 9 - 174." title="Monographie der Dynastinae. 5. Tribus: Oryctini (die Arten von Amerika) (Coleoptera: Melolonthidae)" type="journal article" year="1976">Endrödi 1976: 46</bibRefCitation>
(Argentina, Bolivia, Chile, Colombia, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru), 1985a: 573, 575 (ArgentinaMexico);
<bibRefCitation id="EFEF33FBFF0CFF2AFDF4FE4D32F7FACB" author="Dechambre, R. - P." box="[636,907,1315,1340]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="199" pageNumber="198" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
(Argentina, Bolivia, Central America, Chile, Colombia, Mexico, Paraguay, Peru);
<bibRefCitation id="EFEF33FBFF0CFF2AFD0CFE2D320AFAAC" author="Lachaume" box="[644,886,1347,1372]" firstAuthor="Lachaume" pageId="199" pageNumber="198" pagination="7 - 56" refId="ref211183" refString="Lachaume, G. 1992. Dynastidae americains Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14: 7 - 56." type="journal article" year="1992">Lachaume 1992: 39</bibRefCitation>
(ArgentinaMexico);
<bibRefCitation id="EFEF33FBFF0CFF2AFB0AFE2D3044FA8B" author="Iannuzzi, L. &amp; R. C. Marinoni" journalOrPublisher="Revista Brasileira de Zoologia" pageId="199" pageNumber="198" pagination="93 - 121" part="12" refId="ref209058" refString="Iannuzzi, L., and R. C. Marinoni. 1995. Revisao do genero neotropical Coelosis Hope (Coleoptera, Scarabaeidae, Dynastinae). Revista Brasileira de Zoologia 12 (1): 93 - 121." title="Revisao do genero neotropical Coelosis Hope (Coleoptera, Scarabaeidae, Dynastinae)" type="journal article" year="1995">Iannuzzi and Marinoni 1995: 115</bibRefCitation>
(Argentina, Belize, Bolivia, Chile, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru - as
<taxonomicName id="4C7E3589FF0CFF2AFCAFFEEA3551FA6B" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[807,1069,1411,1436]" class="Insecta" family="Dynastidae" genus="Coelosis" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="198" phylum="Arthropoda" rank="species" species="biloba" subGenus="Eucoelosis">
<emphasis id="B90A9218FF0CFF2AFCAFFEEA323DFA6B" box="[807,833,1412,1435]" italics="true" pageId="199" pageNumber="198">C.</emphasis>
(
<emphasis id="B90A9218FF0CFF2AFCDAFEED32ADFA6B" box="[850,977,1411,1435]" italics="true" pageId="199" pageNumber="198">Eucoelosis</emphasis>
)
<emphasis id="B90A9218FF0CFF2AFC69FEED3551FA6B" box="[993,1069,1411,1435]" italics="true" pageId="199" pageNumber="198">biloba</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0CFF2AFBCDFEED358FFA6C" author="Krell" box="[1093,1267,1411,1436]" firstAuthor="Krell" pageId="199" pageNumber="198" pagination="13 - 20" refId="ref210957" refString="Krell, F. - T. 1996. Type catalogue of some Scarabaeoidea species described by Linnaeus (Insecta: Coleoptera). The Linnean 11 (4): 13 - 20." type="journal article" year="1996">Krell 1996: 17</bibRefCitation>
(none - designation
<typeStatus id="54C5F0A8FF0CFF2AFE9FFECA30C3FA4C" box="[279,447,1444,1468]" pageId="199" pageNumber="198" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="4C7E3589FF0CFF2AFE4EFECD33CFFA4B" authorityName="Linnaeus" authorityYear="1767" box="[454,691,1443,1467]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="199" pageNumber="198" phylum="Arthropoda" rank="species" species="bilobus">
<emphasis id="B90A9218FF0CFF2AFE4EFECD33CFFA4B" box="[454,691,1443,1467]" italics="true" pageId="199" pageNumber="198">Scarabaeus bilobus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF0CFF2AFD43FECD32CCFA4C" author="Ratcliffe" box="[715,944,1443,1468]" firstAuthor="Ratcliffe" pageId="199" pageNumber="198" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 292</bibRefCitation>
(ArgentinaMexico);
<bibRefCitation id="EFEF33FBFF0CFF2AFB31FECE30E2FA2C" author="Gasca-Alvarez, H. J. &amp; B. C. Ratcliffe" journalOrPublisher="Zootaxa" pageId="199" pageNumber="198" pagination="21 - 40" part="3090" refId="ref203834" refString="Gasca-Alvarez, H. J., and B. C. Ratcliffe. 2011. An annoted checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms. Zootaxa 3090: 21 - 40." title="An annoted checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms" type="journal article" year="2011">Gasca-Álvarez and Ratcliffe 2011: 24</bibRefCitation>
(Belize, Bolivia, Colombia, Costa Rica, Ecuador, El Salvador, Honduras, Mexico, Nicaragua, Panama, Peru).
</paragraph>
</subSubSection>
</treatment>
</document>