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<document id="BEC7F5287619A8F9EB823D11DE947A6E" ID="10.11646/zootaxa.4247.5.5SLASH10678" ID-CLB-Dataset="33326" ID-DOI="10.11646/zootaxa.4247.5.5" ID-GBIF-Dataset="92957f37-f810-4bdf-a6e6-fcf99047b816" ID-ISSN="1175-5326" ID-Zenodo-Dep="439185" ID-ZooBank="427F743A-5286-4023-80A6-66659E5B7B49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490859898841" checkinUser="plazi" docAuthor="Bezark, Larry G. &amp; Santos-Silva, Antonio" docDate="2017" docId="03CE9B411F7CFD17FF1D5222FB395CF7" docLanguage="en" docName="zootaxa.4247.5.5SLASH10678.pdf" docOrigin="Zootaxa 4247 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lygrocharis setosus Bezark &amp; Santos-Silva, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="592" masterDocId="FFF7E3391F79FD10FF8A5145FFF25D27" masterDocTitle="Three new species of Ecuadorian Rhinotragini (Coleoptera, Cerambycidae)" masterLastPageNumber="592" masterPageNumber="585" pageNumber="590" updateTime="1698438568535" updateUser="ExternalLinkService">
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<mods:title id="E3EA6EE851C6FBC896E2BD4792C9CEE9">Three new species of Ecuadorian Rhinotragini (Coleoptera, Cerambycidae)</mods:title>
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<mods:namePart id="00C2915C12AAC17A1D9F5A0F8B5F6954">Bezark, Larry G.</mods:namePart>
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<mods:namePart id="A2C3685C193DE36A81B140E98B2CC2EE">Santos-Silva, Antonio</mods:namePart>
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<mods:title id="C517E96F51AE8A5FAD1192491D1CC763">Zootaxa</mods:title>
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<mods:part id="BB1D54ABC7E2746488FC91FECE87F4D8">
<mods:date id="ABAEEC35EFC48923820CC4B03DBDB9FB">2017</mods:date>
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<mods:number id="1035957A31EA0FBED97930B1ABA60303">4247</mods:number>
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<mods:classification id="0E746574E579E06F17FD7F809C81F4B3">journal article</mods:classification>
<mods:identifier id="4B7B151CBB26A27A6C3C4E3AE0B7482D" type="CLB-Dataset">33326</mods:identifier>
<mods:identifier id="C551E581DC1BFD9AA211D3654500B92C" type="DOI">10.11646/zootaxa.4247.5.5</mods:identifier>
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<treatment id="03CE9B411F7CFD17FF1D5222FB395CF7" ID-DOI="http://doi.org/10.5281/zenodo.6033250" ID-GBIF-Taxon="128090210" ID-Zenodo-Dep="6033250" LSID="urn:lsid:plazi:treatment:03CE9B411F7CFD17FF1D5222FB395CF7" httpUri="http://treatment.plazi.org/id/03CE9B411F7CFD17FF1D5222FB395CF7" lastPageId="7" lastPageNumber="592" pageId="5" pageNumber="590">
<subSubSection id="C37D79DC1F7CFD15FF1D5222FED25E84" pageId="5" pageNumber="590" type="nomenclature">
<paragraph id="8BD82A571F7CFD15FF1D5222FED25E84" blockId="5.[151,487,871,931]" pageId="5" pageNumber="590">
<taxonomicName id="4C6751D41F7CFD15FF1D5222FE775EA6" ID-CoL="BLKNN" box="[151,389,871,897]" class="Insecta" family="Cerambycidae" genus="Lygrocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="590" phylum="Arthropoda" rank="species" species="setosus" status="sp. nov.">
<emphasis id="B913F6451F7CFD15FF1D5222FE775EA6" bold="true" box="[151,389,871,897]" italics="true" pageId="5" pageNumber="590">Lygrocharis setosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2204B3E1F7CFD15FE07522DFE155EA6" box="[397,487,872,897]" pageId="5" pageNumber="590" rank="species">
<emphasis id="B913F6451F7CFD15FE07522DFE155EA6" bold="true" box="[397,487,872,897]" pageId="5" pageNumber="590">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="135C36D21F7CFD15FF1552CFFEEA5E84" box="[159,280,906,931]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1804,1827]" captionTargetBox="[167,1419,561,1775]" captionTargetId="figure@6.[151,1436,553,1783]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Lygrocharis setosus sp. nov., holotype female: 9. Dorsal habitus; 10. Ventral habitus; 11. Lateral hatibus; 12. Head, frontal view." httpUri="https://zenodo.org/record/439187/files/figure.png" pageId="5" pageNumber="590">Figs. 9–12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37D79DC1F7CFD17FF1D5297FE195DF3" lastPageId="7" lastPageNumber="592" pageId="5" pageNumber="590" type="description">
<paragraph id="8BD82A571F7CFD15FF1D5297FC35582C" blockId="5.[151,1437,978,2011]" pageId="5" pageNumber="590">
<emphasis id="B913F6451F7CFD15FF1D5297FE8B5ECC" bold="true" box="[151,377,978,1003]" pageId="5" pageNumber="590">Description. Male.</emphasis>
Head and prosternum black; mouthparts yellowish to reddish-brown; scape, pedicel and antennomeres III–VI reddish-brown; antennomeres VI–XI yellowish-brown; pronotum dark reddish-brown; sides of prothorax dark reddish-brown, darker brown on basal and anterior regions; most of ventral side of mesothorax dark-brown; most of ventral side of metathorax brown, more reddish toward apex; elytra reddish-brown with some regions more brownish, mainly on distal half; femoral peduncle yellowish on base, gradually reddish-brown toward femoral club; femoral club brown; pro- and mesotibiae light reddish-brown on base, gradually brown toward apex; metatibiae yellowish on basal 2/3, brown on distal third; pro- and mesotarsi brown; metatarsomere I mostly yellowish-brown, with base and apex more reddish; metatarsomeres II-V reddish-brown; abdomen mostly brown. Pubescence yellowish, more whitish depending on angle of light.
</paragraph>
<paragraph id="8BD82A571F7CFD15FF4D5453FB805A48" blockId="5.[151,1437,978,2011]" pageId="5" pageNumber="590">
<emphasis id="B913F6451F7CFD15FF4D5453FEF55808" bold="true" box="[199,263,1302,1327]" pageId="5" pageNumber="590">Head</emphasis>
. Basal area not elongate behind eyes (distance between posterior margin of eyes and prothoracic margin equal to width of upper eye lobes); rostrum (area between lower eye lobe and apex of gena), viewed from front, short (about 0.4 times length of lower eye lobe). Frons finely, abundantly punctate, except for smooth longitudinal region in middle (punctures slightly sparser toward antennal tubercles); with short, moderately sparse setae. Area between antennal tubercles and upper eyes lobes finely, abundantly punctate; with short, abundant (primarily in middle) setae. Area between upper eye lobes and prothorax finely, sparsely punctate close to eyes, punctate-striate close to prothorax; with short, abundant setae on punctate region, very sparse on punctate-striate region. Antennal tubercles with sculpture and setae as on frons (smoother toward upper eye lobes). Clypeus finely, abundantly punctate, with short setae, except for smooth, glabrous at sides and area closer to labrum; with one long, erect seta on each side of punctate region. Area behind eyes finely, abundantly punctate close to eyes, sparsely punctate close to prothorax, smooth between these two areas on region behind lower eye lobes; with short, abundant setae interspersed with long, erect setae close to lower eye lobes. Gula smooth, glabrous. Submentum rugose-punctate; with short, sparse setae interspersed with long, erect setae. Genae very finely punctate; with short, dense setae close to eyes, glabrous near apex. Distance between upper eye lobes 0.75 times length of scape; in frontal view, distance between lower eye lobes in frontal view 0.95 times length of scape. Antennae 1.2 times length of elytral, reaching middle of abdominal ventrite III; antennal formula (ratio) based on antennomere III: scape = 0.65; pedicel = 0.22; IV = 0.58; V = 0.63; VI = 0.74; VII = 0.61; VIII = 0.52; IX = 0.48; X = 0.39; XI = 0.43.
</paragraph>
<paragraph id="8BD82A571F7CFD15FF4D563FFA6F5AFC" blockId="5.[151,1437,978,2011]" pageId="5" pageNumber="590">
<emphasis id="B913F6451F7CFD15FF4D563FFED25AB4" bold="true" box="[199,288,1914,1939]" pageId="5" pageNumber="590">Thorax</emphasis>
. Prothorax subcylindrical, distinctly widened about middle, slightly longer than wide (1.1 times). Pronotum with five gibbosities on basal half, one longitudinal, at middle, one on each side near base, and one oblique on each side near middle of pronotum; lateral gibbosities on each side of longitudinal one sub-fused.
</paragraph>
<paragraph id="8BD82A571F7FFD16FF1D51D2FD1D5CD2" blockId="6.[151,1437,151,501]" pageId="6" pageNumber="591">
Pronotal surface finely, abundantly punctate, except for sub-smooth area on gibbosities; pubescents, nearly glabrous on gibbosities (primarily on central and anterior gibbosities), and with long, erect, sparse setae. Sides of prothorax finely, abundantly punctate, pubescent, except for transversely striate and subglabrous anterior region. Prosternum finely, abundantly punctate in middle at base, sparsely punctate on remaining surface; pubescent laterally, and with setae distinctly sparser in middle triangular area (setae more abundant basally). Procoxal cavities closed behind. Metepisterna and metasternum pubescent but not obscuring integument. Scutellum pubescent.
<emphasis id="B913F6451F7FFD16FF1D502AFF115CAF" bold="true" box="[151,227,367,392]" pageId="6" pageNumber="591">Elytra</emphasis>
. Slightly narrowed from base to about middle; apex rounded; with distinct carina at center of distal fifth; coarsely, moderately abundantly punctate, punctures slightly finer toward apex; with short, sparse setae, denser at apex and on each side of distal carina, interspersed with long, erect, sparse setae on basal third.
<emphasis id="B913F6451F7FFD16FAD750F2FA675CF7" bold="true" box="[1373,1429,439,464]" pageId="6" pageNumber="591">Legs</emphasis>
. Metatarsomere I 1.3 times longer than II–III together.
</paragraph>
<caption id="DF187ADF1F7FFD16FF1D5649FEA85A66" httpUri="https://zenodo.org/record/439187/files/figure.png" pageId="6" pageNumber="591" targetBox="[167,1419,561,1775]" targetPageId="6">
<paragraph id="8BD82A571F7FFD16FF1D5649FEA85A66" blockId="6.[151,1435,1804,1857]" pageId="6" pageNumber="591">
<emphasis id="B913F6451F7FFD16FF1D5649FEB25A05" bold="true" box="[151,320,1804,1827]" pageId="6" pageNumber="591">FIGURES 9–12</emphasis>
.
<taxonomicName id="4C6751D41F7FFD16FEC75648FDE55A05" box="[333,535,1805,1826]" class="Insecta" family="Cerambycidae" genus="Lygrocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="591" phylum="Arthropoda" rank="species" species="setosus" status="sp. nov.">
<emphasis id="B913F6451F7FFD16FEC75648FDE55A05" box="[333,535,1805,1826]" italics="true" pageId="6" pageNumber="591">Lygrocharis setosus</emphasis>
</taxonomicName>
<emphasis id="B913F6451F7FFD16FD975648FD9C5A05" bold="true" box="[541,622,1805,1826]" pageId="6" pageNumber="591">
<taxonomicNameLabel id="A2204B3E1F7FFD16FD975648FD9C5A05" box="[541,622,1805,1826]" pageId="6" pageNumber="591" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female:
<emphasis id="B913F6451F7FFD16FCA65648FCCA5A05" bold="true" box="[812,824,1805,1826]" pageId="6" pageNumber="591">9</emphasis>
. Dorsal habitus;
<emphasis id="B913F6451F7FFD16FC6C5648FC0D5A05" bold="true" box="[998,1023,1805,1826]" pageId="6" pageNumber="591">10</emphasis>
. Ventral habitus;
<emphasis id="B913F6451F7FFD16FB395648FB3B5A05" bold="true" box="[1203,1225,1805,1826]" pageId="6" pageNumber="591">11</emphasis>
. Lateral hatibus;
<emphasis id="B913F6451F7FFD16FAF65648FA675A05" bold="true" box="[1404,1429,1805,1826]" pageId="6" pageNumber="591">12</emphasis>
. Head, frontal view.
</paragraph>
</caption>
<paragraph id="8BD82A571F7FFD16FF4D5628FAC05AA1" blockId="6.[151,1436,1901,2035]" box="[199,1330,1901,1926]" pageId="6" pageNumber="591">
<emphasis id="B913F6451F7FFD16FF4D5628FECE5AA1" bold="true" box="[199,316,1901,1926]" pageId="6" pageNumber="591">Abdomen</emphasis>
. Ventrites pubescent, interspersed with some long, erect setae. Apex of ventrite V truncate
</paragraph>
<paragraph id="8BD82A571F7FFD16FF4D56D7FABD5AE8" blockId="6.[151,1436,1901,2035]" pageId="6" pageNumber="591">
<emphasis id="B913F6451F7FFD16FF4D56D7FEEC5A8C" bold="true" box="[199,286,1938,1963]" pageId="6" pageNumber="591">Female</emphasis>
. Head mostly dark reddish-brown dorsally; all antennal segments reddish-brown. In frontal view, distance between lower eye lobes 1.1 times length of scape. Apex of abdominal ventrite V slightly rounded.
</paragraph>
<paragraph id="8BD82A571F7FFD17FF4D569FFE195DF3" blockId="6.[151,1436,1901,2035]" lastBlockId="7.[151,1436,151,464]" lastPageId="7" lastPageNumber="592" pageId="6" pageNumber="591">
<emphasis id="B913F6451F7FFD16FF4D569FFCC55AD4" bold="true" box="[199,823,2010,2035]" pageId="6" pageNumber="591">
Dimensions in mm (
<typeStatus id="54DC94F51F7FFD16FE32569FFDD25AD4" box="[440,544,2010,2035]" pageId="6" pageNumber="591" type="holotype">holotype</typeStatus>
male/
<typeStatus id="54DC94F51F7FFD16FDE6569FFD245AD4" box="[620,726,2010,2035]" pageId="6" pageNumber="591" type="paratype">paratype</typeStatus>
female)
</emphasis>
. Total length, 4.20/4.90; prothorax: length, 0.95/1.00; anterior width, 0.65/0.70; posterior width, 0.60/0.65; widest prothoracic width, 0.80/0.80; humeral width, 0.80/ 0.90; elytral length, 2.70/3.10.
</paragraph>
</subSubSection>
<subSubSection id="C37D79DC1F7EFD17FF4D519AFBB85C3A" pageId="7" pageNumber="592" type="materials_examined">
<paragraph id="8BD82A571F7EFD17FF4D519AFBB85C3A" blockId="7.[151,1436,151,464]" pageId="7" pageNumber="592">
<emphasis id="B913F6451F7EFD17FF4D519AFE815DDF" bold="true" box="[199,371,223,248]" pageId="7" pageNumber="592">Type material</emphasis>
. Holotype female (CASC),
<materialsCitation id="3B0F200A1F7EFD17FD4E51A5FBB85C3A" ID-GBIF-Occurrence="1454954778" collectingDate="2006-02-22" collectionCode="LGBC" collectorName="F. T. Hovore &amp; I. Swift" country="Ecuador" latitude="-4.1356945" location="Gonzanama" longLatPrecision="1" longitude="-79.39344" pageId="7" pageNumber="592" specimenCount="1" stateProvince="Loja" typeStatus="paratype">
<typeStatus id="54DC94F51F7EFD17FD4E51A5FCD65DDF" box="[708,804,224,248]" pageId="7" pageNumber="592" type="paratype">paratype</typeStatus>
female (
<collectionCode id="ED76B2921F7EFD17FC1851A5FC2C5DDF" box="[914,990,224,248]" pageId="7" pageNumber="592">LGBC</collectionCode>
<emphasis id="B913F6451F7EFD17FC54519AFC155DDF" bold="true" box="[990,999,223,248]" pageId="7" pageNumber="592">)</emphasis>
from
<collectingCountry id="F3706AC71F7EFD17FBBD51A5FB335DDF" box="[1079,1217,224,248]" name="Ecuador" pageId="7" pageNumber="592">ECUADOR</collectingCountry>
,
<collectingRegion id="49A3E4B51F7EFD17FB5851A5FAF75DD0" box="[1234,1285,224,247]" country="Ecuador" name="Loja" pageId="7" pageNumber="592">
<emphasis id="B913F6451F7EFD17FB5851A5FAF75DD0" box="[1234,1285,224,247]" italics="true" pageId="7" pageNumber="592">Loja</emphasis>
</collectingRegion>
:
<quantity id="4C9F87B21F7EFD17FA90519AFA895DDF" box="[1306,1403,223,248]" metricMagnitude="4" metricUnit="m" metricValue="1.85" pageId="7" pageNumber="592" unit="km" value="18.5">18.5 km</quantity>
N
<location id="8EB87C8C1F7EFD17FF1D5041FEEF5C3B" LSID="urn:lsid:plazi:treatment:03CE9B411F7CFD17FF1D5222FB395CF7:8EB87C8C1F7EFD17FF1D5041FEEF5C3B" box="[151,285,260,284]" country="Ecuador" latitude="-4.1356945" longLatPrecision="1" longitude="-79.39344" name="Gonzanama" pageId="7" pageNumber="592" stateProvince="Loja">Gonzanama</location>
(
<geoCoordinate id="EE534C901F7EFD17FEA65041FE365C3A" box="[300,452,260,285]" direction="south" orientation="latitude" pageId="7" pageNumber="592" precision="1" value="-4.1356945">04°08’08.5”S</geoCoordinate>
/
<geoCoordinate id="EE534C901F7EFD17FE505041FD725C3A" box="[474,640,260,285]" direction="west" orientation="longitude" pageId="7" pageNumber="592" precision="1" value="-79.39344">79°23’36.4”W</geoCoordinate>
),
<collectingDate id="EF9DF57F1F7EFD17FD195041FCF85C3A" box="[659,778,260,285]" pageId="7" pageNumber="592" value="2006-02-22">
<date id="FFD90C971F7EFD17FD195041FCF85C3A" box="[659,778,260,285]" pageId="7" pageNumber="592" value="2006-02-22">22.II.2006</date>
</collectingDate>
,
<collectorName id="26924F811F7EFD17FC9C5040FC505C3B" box="[790,930,261,284]" pageId="7" pageNumber="592">F. T. Hovore</collectorName>
&amp;
<collectorName id="26924F811F7EFD17FC4C5040FBE85C3A" box="[966,1050,260,285]" pageId="7" pageNumber="592">I. Swift</collectorName>
col.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37D79DC1F7EFD17FF4D5062FB635C18" box="[199,1169,295,320]" pageId="7" pageNumber="592" type="etymology">
<paragraph id="8BD82A571F7EFD17FF4D5062FB635C18" blockId="7.[151,1436,151,464]" box="[199,1169,295,320]" pageId="7" pageNumber="592">
<emphasis id="B913F6451F7EFD17FF4D5062FEB45C67" bold="true" box="[199,326,295,320]" pageId="7" pageNumber="592">Etymology</emphasis>
. The specific epithet
<taxonomicName id="4C6751D41F7EFD17FDB2506CFD7B5C67" box="[568,649,297,320]" class="Insecta" family="Cerambycidae" genus="Lygrocharis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="592" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="B913F6451F7EFD17FDB2506CFD7B5C67" box="[568,649,297,320]" italics="true" pageId="7" pageNumber="592">setosus</emphasis>
</taxonomicName>
, refers to the abundant setae on the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C37D79DC1F7EFD17FF4D5009FB395CF7" pageId="7" pageNumber="592" type="discussion">
<paragraph id="8BD82A571F7EFD17FF4D5009FB395CF7" blockId="7.[151,1436,151,464]" pageId="7" pageNumber="592">
<emphasis id="B913F6451F7EFD17FF4D5009FEC65C42" bold="true" box="[199,308,332,357]" pageId="7" pageNumber="592">Remarks</emphasis>
.
<taxonomicName id="4C6751D41F7EFD17FEC85009FDD35C43" box="[322,545,332,356]" class="Insecta" family="Cerambycidae" genus="Lygrocharis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="592" phylum="Arthropoda" rank="species" species="setosus" status="sp. nov.">
<emphasis id="B913F6451F7EFD17FEC85009FDD35C43" box="[322,545,332,356]" italics="true" pageId="7" pageNumber="592">Lygrocharis setosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2204B3E1F7EFD17FDA25009FD705C42" box="[552,642,332,357]" pageId="7" pageNumber="592" rank="species">
<emphasis id="B913F6451F7EFD17FDA25009FD705C42" bold="true" box="[552,642,332,357]" pageId="7" pageNumber="592">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C6751D41F7EFD17FC9C5008FBE55C43" authority="Melzer, 1927" authorityName="Melzer" authorityYear="1927" box="[790,1047,332,357]" class="Insecta" family="Cerambycidae" genus="Lygrocharis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="592" phylum="Arthropoda" rank="species" species="neivai">
<emphasis id="B913F6451F7EFD17FC9C5008FC8B5C43" box="[790,889,333,356]" italics="true" pageId="7" pageNumber="592">L. neivai</emphasis>
Melzer, 1927
</taxonomicName>
, and
<taxonomicName id="4C6751D41F7EFD17FBD85008FA6A5C43" authority="Mendes, 1938" authorityName="Mendes" authorityYear="1938" box="[1106,1432,332,357]" class="Insecta" family="Cerambycidae" genus="Lygrocharis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="592" phylum="Arthropoda" rank="species" species="nigripennis">
<emphasis id="B913F6451F7EFD17FBD85008FB1D5C43" box="[1106,1263,333,356]" italics="true" pageId="7" pageNumber="592">L. nigripennis</emphasis>
<bibRefCitation id="EFF657A61F7EFD17FB7C5009FA6A5C43" author="Mendes" box="[1270,1432,332,357]" pageId="7" pageNumber="592" refString="Mendes, D. (1938) Tres especies novas de cerambycideos do Brasil, dos generos Lygrocharis, Rhathymoscelis e Alphus (Coleoptera). Revista de Entomologia, 9, 117 - 121." type="journal article" year="1938">Mendes, 1938</bibRefCitation>
</taxonomicName>
, by the prothorax being proportionally larger and wider anteriorly than basally (see photographs of the
<typeStatus id="54DC94F51F7EFD17FAB55035FA885CAF" box="[1343,1402,368,392]" pageId="7" pageNumber="592">types</typeStatus>
of these species at
<bibRefCitation id="EFF657A61F7EFD17FEDB50D1FE145C8A" author="Bezark" box="[337,486,404,429]" pageId="7" pageNumber="592" refString="Bezark, L. G. (2016) A photographic Catalog of the Cerambycidae of the New World. Available from: https: // apps 2. cdfa. ca. gov / publicApps / plant / bycidDB / wsearch. asp? w = n (accessed 31 March 2016)" type="journal article" year="2016">Bezark 2016</bibRefCitation>
). However, the new species does share the distinctly separated lower eyes lobes of males, the closed procoxal cavities, and the carinate apical fifth of the elytra, with these species.
</paragraph>
</subSubSection>
</treatment>
</document>
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