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<document id="2FCF6A26313830515C8DF8E9742E44F4" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFF15FF48FDEDFD41AF1AED90" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia pseudopleiopetala Tebbitt" docType="treatment" docVersion="7" lastPageNumber="178" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="177" updateTime="1707123162844" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="79062F28C4605498C6159B758314F260">The genus Begonia (Begoniaceae) in Peru</mods:title>
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<mods:namePart id="2868682196785C474BD4A69954118388">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="B16A47572518DD60414A5CB0F10A4645">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
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<mods:namePart id="7BA3B6051A3A17507A4A78B80732C3D3">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="D1E102E0304D98A233BB0A2F965AFC8C">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
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<mods:namePart id="3E278BA44743EFBFBC733A290485B146">Purvis, David A.</mods:namePart>
<mods:affiliation id="4C72EB83D4A2933D692ADC48E81751EC">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
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<mods:namePart id="7440458DDE2CDDD69C800829ADFAC324">Delves, Jay</mods:namePart>
<mods:affiliation id="670F260CC6D072C60EB3B954EECAE89F">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
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<mods:namePart id="209E17C990C2A0100F8623D22DC105AC">Allen, Josh P.</mods:namePart>
<mods:affiliation id="BC44BBB2130B6D283C837403F70CC377">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="C0CF0A759778BC696F8334D476FC9DF8" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
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<mods:namePart id="20E4022DCCFB2E8C13E032AC52094951">Reynel, Carlos</mods:namePart>
<mods:affiliation id="FE274BD963C2BA22BB057236107E7717">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
<mods:nameIdentifier id="DD18ADF4D401C14DD62C435DE48DA2AC" type="email">reynel@lamolina.edu.pe</mods:nameIdentifier>
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<mods:title id="95CE917D362779617177C5573745D82F">European Journal of Taxonomy</mods:title>
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<mods:date id="BA8539D2E93B6B429270FA99F6E72605">2023</mods:date>
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<treatment id="03B26B4BFF15FF48FDEDFD41AF1AED90" LSID="urn:lsid:plazi:treatment:03B26B4BFF15FF48FDEDFD41AF1AED90" httpUri="http://treatment.plazi.org/id/03B26B4BFF15FF48FDEDFD41AF1AED90" lastPageId="177" lastPageNumber="178" pageId="176" pageNumber="177">
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39.
<taxonomicName id="4C1BA1DEFF15FF49FDB9FD42AED8E96E" ID-CoL="87Z4K" ID-ENA="2184944" authority="Tebbitt" authorityName="Tebbitt" box="[608,1023,765,792]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="176" pageNumber="177" phylum="Tracheophyta" rank="species" species="pseudopleiopetala">
<emphasis id="B96F064FFF15FF49FDB9FD42AE81E961" bold="true" box="[608,934,765,792]" italics="true" pageId="176" pageNumber="177">Begonia pseudopleiopetala</emphasis>
Tebbitt
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<figureCitation id="1320C6D8FF15FF49FD3FFC9EAE6AE945" box="[742,845,801,828]" captionStart="Fig" captionStartId="160.[189,232,1822,1848]" captionTargetBox="[194,1391,558,1776]" captionTargetId="figure-14@160.[189,1398,548,1784]" captionTargetPageId="160" captionText="Fig. 52. Distribution of the octopetala group of Begonia sect. Eupetalum (Lindl.) A.DC. in Peru and surrounding countries. A. B. anemoniflora Irmsch. (red) and B. tumbezensis Irmsch. (blue). B. B. octopetala L.Hér. subsp. octopetala (red) and B. octopetala subsp. ovatiformis Irmsch. (blue). C. B. pleiopetala A.DC. (red) and B. pseudopleiopetala Tebbitt (blue). D. B. polypetala A.DC. (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178413" httpUri="https://zenodo.org/record/8178413/files/figure.png" pageId="176" pageNumber="177">Fig. 52C</figureCitation>
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<bibCitation id="5E31D231FF15FF49FF64FCDBAF73E907" box="[189,596,867,894]" issue="4" journal="Novon" page="486" pageId="176" pageNumber="177" score="4.864797368871433" volume="23" year="2015">
<emphasis id="B96F064FFF15FF49FF64FCDBAC2DE904" box="[189,266,868,893]" italics="true" pageId="176" pageNumber="177">Novon</emphasis>
23 (4): 486 (
<bibRefCitation id="EF8AA7ACFF15FF49FE74FCDBAF6BE907" author="Tebbitt M. C." box="[429,588,867,894]" journalOrPublisher="Novon" pageId="176" pageNumber="177" pagination="479 - 489" part="23" refId="ref232441" refString="Tebbitt M. C. 2015. Two new species of Andean tuberous Begonia in the B. octopetala group (Begoniaceae). Novon 23 (4): 479 - 489. https: // doi. org / 10.3417 / 2013027" title="Two new species of Andean tuberous Begonia in the B. octopetala group (Begoniaceae)" type="journal article" year="2015">Tebbitt 2015</bibRefCitation>
)
</bibCitation>
.
</paragraph>
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<paragraph id="8BA4DA5DFF15FF49FDBBFCDBACEAE993" blockId="176.[189,1398,867,1002]" pageId="176" pageNumber="177">
<emphasis id="B96F064FFF15FF49FDA3FCDBAF9FE907" bold="true" box="[634,696,868,894]" pageId="176" pageNumber="177">Type</emphasis>
:
<materialsCitation id="3B73D000FF15FF49FD13FCDBA902E9BF" collectingDate="1986-04-14" collectionCode="F" collectorName="M. O. Dillon &amp; D. Dillon &amp; A. Sagastegui A." country="Peru" elevation="2180" latitude="-7.4166665" location="Contumaza" longLatPrecision="1303" longitude="-78.416664" municipality="Contumaza" pageId="176" pageNumber="177" specimenCode="V0360785F" specimenCount="1" stateProvince="Cajamarca Region" typeStatus="holotype">
<collectingCountry id="F30C9ACDFF15FF49FD13FCDBAE32E907" box="[714,789,868,894]" name="Peru" pageId="176" pageNumber="177">PERU</collectingCountry>
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<collectingRegion id="49DF14BFFF15FF49FCEEFCDCA90CE904" box="[823,1067,867,893]" country="Peru" name="Cajamarca" pageId="176" pageNumber="177">Cajamarca Region</collectingRegion>
:
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ca
<quantity id="4CE377B8FF15FF49FAC9FCDBA87BE907" box="[1296,1372,868,894]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="176" pageNumber="177" unit="km" value="20.0">20 km</quantity>
S of Contumazá, ca 18jm above Cascas
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;
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,
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;
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<quantity id="4CE377B8FF15FF49FC44FC38A967E9DB" box="[925,1088,903,930]" metricMagnitude="3" metricUnit="m" metricValue="2.18" metricValueMax="2.2" metricValueMin="2.16" pageId="176" pageNumber="177" unit="m" value="2180.0" valueMax="2200.0" valueMin="2160.0">21602200 m</quantity>
a.s.l.
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;
<date id="FFA5FC9DFF15FF49FB56FC37A80AE9D8" box="[1167,1325,903,930]" pageId="176" pageNumber="177" value="1986-04-14">
<collectingDate id="EFE10575FF15FF49FB56FC37A80AE9D8" box="[1167,1325,903,930]" pageId="176" pageNumber="177" value="1986-04-14">14 Apr. 1986</collectingDate>
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;
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<collectorName id="26EEBF8BFF15FF49FAE3FC37AC1EE9BC" pageId="176" pageNumber="177">M.O. Dillon</collectorName>
,
<collectorName id="26EEBF8BFF15FF49FE9CFC13AC92E9BC" box="[325,437,939,965]" pageId="176" pageNumber="177">D. Dillon</collectorName>
&amp;
<collectorName id="26EEBF8BFF15FF49FE03FC13AFB9E9BF" box="[474,670,940,966]" pageId="176" pageNumber="177">A. Sagástegui A.</collectorName>
4528
</emphasis>
;
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:
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[
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<materialsCitation id="3B73D000FF15FF49FBF4FC14ACEAE993" collectingDate="1986-04-14" collectionCode="GB, US" collectorName="M. O. Dillon &amp; D. Dillon &amp; A. Sagastegui A." country="Peru" elevation="2180" latitude="-7.4166665" location="Contumaza" longLatPrecision="1303" longitude="-78.416664" municipality="Contumaza" pageId="176" pageNumber="177" specimenCode="GB0058014, US00431510" specimenCount="1" stateProvince="Cajamarca Region" typeStatus="isotypes">
<typeStatus id="54A064FFFF15FF49FBF4FC14A9B5E9BC" box="[1069,1170,939,965]" pageId="176" pageNumber="177">isotypes</typeStatus>
:
<collectionCode id="ED0A4298FF15FF49FB45FC13A9E2E9BF" box="[1180,1221,940,966]" country="Sweden" name="University of Gothenburg" pageId="176" pageNumber="177" type="Herbarium">GB</collectionCode>
[
<specimenCode id="DBBD7226FF15FF49FB0AFC14A84CE9BF" box="[1235,1387,939,966]" collectionCode="GB" country="Sweden" name="University of Gothenburg" pageId="176" pageNumber="177" type="Herbarium">GB0058014</specimenCode>
],
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[
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].
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<emphasis id="B96F064FFF15FF49FF64FBADAC62EE55" bold="true" box="[189,325,1042,1068]" pageId="176" pageNumber="177">Etymology</emphasis>
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This species is easily confused with
<taxonomicName id="4C1BA1DEFF15FF49FDB1FBFDAE29EE22" box="[616,782,1089,1115]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="176" pageNumber="177" phylum="Tracheophyta" rank="species" species="pleiopetala">
<emphasis id="B96F064FFF15FF49FDB1FBFDAE29EE22" box="[616,782,1089,1115]" italics="true" pageId="176" pageNumber="177">B. pleiopetala</emphasis>
</taxonomicName>
and the epithet reflects this.
</paragraph>
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<subSubSection id="C30189D6FF15FF49FF64FB3CA82CEF95" pageId="176" pageNumber="177" type="materials_examined">
<paragraph id="8BA4DA5DFF15FF49FF64FB3CAF09EEE4" blockId="176.[189,558,1155,1181]" box="[189,558,1155,1181]" pageId="176" pageNumber="177">
<heading id="D0EC6D31FF15FF49FF64FB3CAF09EEE4" bold="true" box="[189,558,1155,1181]" fontSize="11" level="3" pageId="176" pageNumber="177" reason="3">
<emphasis id="B96F064FFF15FF49FF64FB3CAF09EEE4" bold="true" box="[189,558,1155,1181]" pageId="176" pageNumber="177">Selected specimens examined</emphasis>
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</paragraph>
<paragraph id="8BA4DA5DFF15FF49FF64FB0CA82CEF95" blockId="176.[189,1399,1201,1516]" pageId="176" pageNumber="177">
<materialsCitation id="3B73D000FF15FF49FF64FB0CAF4BEE89" collectingDate="1948-03-22" collectionCode="USM" collectorName="R. Ferreyra" country="Peru" county="Prov. Huancabamba" elevation="1400" latitude="-5.366667" location="Canchaque" longLatPrecision="1305" longitude="-79.6" municipality="Canchaque" pageId="176" pageNumber="177" specimenCount="1" stateProvince="Piura Region">
<collectingCountry id="F30C9ACDFF15FF49FF64FB0CAC2FEEB4" box="[189,264,1203,1229]" name="Peru" pageId="176" pageNumber="177">PERU</collectingCountry>
<emphasis id="B96F064FFF15FF49FEF9FB0DAFF0EEB5" bold="true" box="[288,727,1202,1228]" pageId="176" pageNumber="177">
<collectingRegion id="49DF14BFFF15FF49FEF9FB0DACEDEEB5" box="[288,458,1202,1228]" country="Peru" name="Piura" pageId="176" pageNumber="177">Piura Region</collectingRegion>
:
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a.s.l.
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;
<date id="FFA5FC9DFF15FF49FA80FB0DAC1EEE89" pageId="176" pageNumber="177" value="1948-03-22">
<collectingDate id="EFE10575FF15FF49FA80FB0DAC1EEE89" pageId="176" pageNumber="177" value="1948-03-22">22 Mar. 1948</collectingDate>
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3134
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;
<collectionCode id="ED0A4298FF15FF49FDFDFB69AF40EE89" box="[548,615,1238,1264]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="176" pageNumber="177">USM</collectionCode>
.
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:
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;
<date id="FFA5FC9DFF15FF49FDB3FB45AE27EF6C" box="[618,768,1274,1301]" pageId="176" pageNumber="177" value="1965-05-06">
<collectingDate id="EFE10575FF15FF49FDB3FB45AE27EF6C" box="[618,768,1274,1301]" pageId="176" pageNumber="177" value="1965-05-06">6 May 1965</collectingDate>
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<collectorName id="26EEBF8BFF15FF49FCD7FB45AEF0EF6D" box="[782,983,1274,1300]" pageId="176" pageNumber="177">A. Sagástegui A.</collectorName>
&amp;
<collectorName id="26EEBF8BFF15FF49FBD8FB44A988EF6D" box="[1025,1199,1274,1300]" pageId="176" pageNumber="177">M. Fukushima</collectorName>
5070
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;
<collectionCode id="ED0A4298FF15FF49FADFFB44A864EF6C" box="[1286,1347,1275,1301]" country="Japan" httpUri="http://grbio.org/cool/gtk4-xv5b" name="HUT Culture Collection" pageId="176" pageNumber="177">HUT</collectionCode>
,
<collectionCode id="ED0A4298FF15FF49FA88FB45A851EF6D" box="[1361,1398,1274,1300]" pageId="176" pageNumber="177">US</collectionCode>
[
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<materialsCitation id="3B73D000FF15FF49FE55FAA1AEE7EF24" collectingDate="1985-04-06" collectionCode="HUT" collectorName="A. Sagastegui A. &amp; M. Guzman &amp; S. Leiva G. &amp; C. Tellez A." elevation="2700" latitude="-7.4" location="El Tunel" longLatPrecision="1303" longitude="-78.8" pageId="176" pageNumber="177" specimenCount="1">
<location id="8EC48C86FF15FF49FE55FAA1ACDFEF41" LSID="urn:lsid:plazi:treatment:03B26B4BFF15FF48FDEDFD41AF1AED90:8EC48C86FF15FF49FE55FAA1ACDFEF41" box="[396,504,1310,1336]" latitude="-7.4" longLatPrecision="1303" longitude="-78.8" name="El Tunel" pageId="176" pageNumber="177">El Túnel</location>
,
<location id="8EC48C86FF15FF49FDD1FAA1AFCAEF41" LSID="urn:lsid:plazi:treatment:03B26B4BFF15FF48FDEDFD41AF1AED90:8EC48C86FF15FF49FDD1FAA1AFCAEF41" box="[520,749,1310,1336]" latitude="-7.4" longLatPrecision="1303" longitude="-78.8" name="Cascas-Contumaza" pageId="176" pageNumber="177">Cascas-Contumazá</location>
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<quantity id="4CE377B8FF15FF49FBDDFAA1A946EF40" box="[1028,1121,1310,1337]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="176" pageNumber="177" unit="m" value="2700.0">2700 m</quantity>
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;
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<collectorName id="26EEBF8BFF15FF49FA84FAA1AC47EF25" pageId="176" pageNumber="177">A. Sagástegui A.</collectorName>
,
<collectorName id="26EEBF8BFF15FF49FEB2FAFCACD0EF25" box="[363,503,1346,1372]" pageId="176" pageNumber="177">M. Guzmán</collectorName>
,
<collectorName id="26EEBF8BFF15FF49FDD9FAFDAFA7EF25" box="[512,640,1346,1372]" pageId="176" pageNumber="177">S. Leiva G.</collectorName>
&amp;
<collectorName id="26EEBF8BFF15FF49FD7AFAFDAE00EF25" box="[675,807,1346,1372]" pageId="176" pageNumber="177">C. Tellez A.</collectorName>
12629
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;
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<materialsCitation id="3B73D000FF15FF49FC0CFAFCA992EFF9" collectingDate="1994-06-16" collectionCode="NY" collectorName="A. Sagastegui A. &amp; S. Leiva G. &amp; P. Lezama" elevation="2500" latitude="-7.4333334" location="Bosque Cachil" longLatPrecision="1303" longitude="-78.816666" pageId="176" pageNumber="177" specimenCount="1">
<location id="8EC48C86FF15FF49FC0CFAFCA9A5EF25" LSID="urn:lsid:plazi:treatment:03B26B4BFF15FF48FDEDFD41AF1AED90:8EC48C86FF15FF49FC0CFAFCA9A5EF25" box="[981,1154,1346,1373]" latitude="-7.4333334" longLatPrecision="1303" longitude="-78.816666" name="Bosque Cachil" pageId="176" pageNumber="177">Bosque Cachil</location>
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<quantity id="4CE377B8FF15FF49FF64FAD9AC3EEFF9" box="[189,281,1382,1409]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="176" pageNumber="177" unit="m" value="2500.0">2500 m</quantity>
a.s.l.
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;
<date id="FFA5FC9DFF15FF49FEB2FAD9AF2DEFF9" box="[363,522,1382,1408]" pageId="176" pageNumber="177" value="1994-06-16">
<collectingDate id="EFE10575FF15FF49FEB2FAD9AF2DEFF9" box="[363,522,1382,1408]" pageId="176" pageNumber="177" value="1994-06-16">16 Jun. 1994</collectingDate>
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,
<collectorName id="26EEBF8BFF15FF49FD2AFAD9AE5BEFF9" box="[755,892,1382,1408]" pageId="176" pageNumber="177">S. Leiva G.</collectorName>
&amp;
<collectorName id="26EEBF8BFF15FF49FC71FAD8A904EFF9" box="[936,1059,1383,1408]" pageId="176" pageNumber="177">P. Lezama</collectorName>
15405
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;
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.
<materialsCitation id="3B73D000FF15FF49FB03FAD9AE24EFB1" collectingDate="1962-05-19" collectionCode="HUT" collectorName="A. Lopez &amp; A. Sagastegui A. &amp; I. Sanchez" elevation="2250" latitude="-7.4166665" location="Cascas-Contumaza" longLatPrecision="1303" longitude="-78.78333" municipality="Prov. Gran Chimu" pageId="176" pageNumber="177" specimenCount="1" stateProvince="La Libertad Region">
<emphasis id="B96F064FFF15FF49FB03FAD9AF36EFDD" bold="true" pageId="176" pageNumber="177">
<collectingRegion id="49DF14BFFF15FF49FB03FAD9AC3DEFDD" country="United States of America" name="La Libertad Region" pageId="176" pageNumber="177">La Libertad Region</collectingRegion>
:
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<location id="8EC48C86FF15FF49FDF0FA35AE29EFDD" LSID="urn:lsid:plazi:treatment:03B26B4BFF15FF48FDEDFD41AF1AED90:8EC48C86FF15FF49FDF0FA35AE29EFDD" box="[553,782,1418,1444]" latitude="-7.4166665" longLatPrecision="1303" longitude="-78.78333" municipality="Prov. Gran Chimu" name="Cascas-Contumaza" pageId="176" pageNumber="177" stateProvince="La Libertad Region">Cascas-Contumazá</location>
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,
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a.s.l.
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;
<date id="FFA5FC9DFF15FF49FB6AFA35A876EFDD" box="[1203,1361,1418,1444]" pageId="176" pageNumber="177" value="1962-05-19">
<collectingDate id="EFE10575FF15FF49FB6AFA35A876EFDD" box="[1203,1361,1418,1444]" pageId="176" pageNumber="177" value="1962-05-19">19 May 1962</collectingDate>
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<collectorName id="26EEBF8BFF15FF49FA84FA35AC20EFB1" pageId="176" pageNumber="177">A. López</collectorName>
,
<collectorName id="26EEBF8BFF15FF49FECBFA11ACF2EFB1" box="[274,469,1454,1480]" pageId="176" pageNumber="177">A. Sagástegui A.</collectorName>
&amp;
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3678
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;
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,
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a.s.l.
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;
<date id="FFA5FC9DFF15FF49FD41FA6DAE1FEF95" box="[664,824,1489,1516]" pageId="176" pageNumber="177" value="2015-05-23">
<collectingDate id="EFE10575FF15FF49FD41FA6DAE1FEF95" box="[664,824,1489,1516]" pageId="176" pageNumber="177" value="2015-05-23">23 May 2015</collectingDate>
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;
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&amp;
<collectorName id="26EEBF8BFF15FF49FC27FA6DA97AEF92" box="[1022,1117,1490,1516]" pageId="176" pageNumber="177">A. Daza</collectorName>
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;
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,
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</paragraph>
</subSubSection>
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<emphasis id="B96F064FFF15FF49FF64F9ABAC68EC57" bold="true" box="[189,335,1556,1582]" pageId="176" pageNumber="177">Description</emphasis>
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Acaulescent, tuberous herb, to
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high.
<emphasis id="B96F064FFF15FF49FD15F9FCAE36EC24" box="[716,785,1603,1629]" italics="true" pageId="176" pageNumber="177">Tuber</emphasis>
ellipsoid, 0.81.3 ×
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, with 1 growing point.
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persistent, triangular, 23 ×
<quantity id="4CE377B8FF15FF49FD58F9D8AE27ECFB" box="[641,768,1639,1666]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="176" pageNumber="177" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
, apex acuminate, aristate, opaque, brown, glabrous, margin entire, aciliate.
<emphasis id="B96F064FFF15FF49FE1FF933AF3FECDC" box="[454,536,1676,1701]" italics="true" pageId="176" pageNumber="177">Leaves</emphasis>
15, alternate, basifixed; petiole 3.511(50) cm long, pale pink, glabrous to densely pubescent; blade sub-symmetric, ovate, to 6 ×
<quantity id="4CE377B8FF15FF49FC50F910AEC6ECB0" box="[905,993,1711,1738]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="176" pageNumber="177" unit="cm" value="6.5">6.5 cm</quantity>
, succulent, apex acuminate, base cordate, basal lobes not overlapping, sinus to
<quantity id="4CE377B8FF15FF49FD38F96CAE11EC94" box="[737,822,1747,1774]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="176" pageNumber="177" unit="mm" value="15.0">15 mm</quantity>
deep, margin denticulate to dentate, glabrous to ciliate, upper surface green, sparsely to densely pilose, lower surface pale green, glabrous, pubescent on the major veins, veins palmate with one major vein, 56 veined from the base, with 13 secondary veins on the larger side, 13 secondary veins on the smaller side.
<emphasis id="B96F064FFF15FF49FC17F880A951ED20" box="[974,1142,1855,1881]" italics="true" pageId="176" pageNumber="177">Inflorescences</emphasis>
1 per plant, bisexual, axillary, erect, a dichasial or monochasial cyme, with up to 3 branches, bearing up to 3 staminate flowers and 2 pistillate flowers, protandrous; peduncle to
<quantity id="4CE377B8FF15FF49FCC6F839AE4BEDD8" box="[799,876,1926,1953]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" pageId="176" pageNumber="177" unit="cm" value="65.0">65 cm</quantity>
long, pink, sparsely to densely pubescent, bracts persistent, elliptic to ovate, 2.511 ×
<quantity id="4CE377B8FF15FF49FD62F815AE12EDBC" box="[699,821,1962,1989]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="8.0" metricValueMin="1.5" pageId="176" pageNumber="177" unit="mm" value="4.75" valueMax="8.0" valueMin="1.5">1.58 mm</quantity>
, opaque, pink, glabrous, pubescent beneath, apex acute, margin entire, aciliate.
<emphasis id="B96F064FFF15FF49FDCDF870AFC0ED91" box="[532,743,1998,2025]" italics="true" pageId="176" pageNumber="177">Staminate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF15FF49FCA2F871AEEAED90" box="[891,973,1998,2025]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="176" pageNumber="177" unit="mm" value="40.0">40 mm</quantity>
long, sparsely to densely pubescent; tepals 78, spreading, obovate to elliptic, 1319 ×
<quantity id="4CE377B8FF14FF48FCCEFEB6AEADEB5A" box="[791,906,265,292]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="11.0" metricValueMin="5.0" pageId="177" pageNumber="178" unit="mm" value="8.0" valueMax="11.0" valueMin="5.0">511 mm</quantity>
, apex obtuse to rounded, white, glabrous, outer surfaces sparsely pubescent, margin entire, aciliate; stamens ca 100, spreading, yellow, filaments
<quantity id="4CE377B8FF14FF48FF64FEEDAC1FEB15" box="[189,312,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="177" pageNumber="178" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.53 mm</quantity>
long, fused at the base, anthers obovate, ca 1 ×
<quantity id="4CE377B8FF14FF48FC4BFEEDAED7EB15" box="[914,1008,338,365]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="177" pageNumber="178" unit="mm" value="0.5">0.5 mm</quantity>
long, dehiscing via lateral slits, connectives extended, symmetrically basifixed.
<emphasis id="B96F064FFF14FF48FD2EFEC9AEE6EBE9" box="[759,961,374,400]" italics="true" pageId="177" pageNumber="178">Pistillate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF14FF48FB85FEC9A996EBE8" box="[1116,1201,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="177" pageNumber="178" unit="mm" value="32.0">32 mm</quantity>
long; bracteoles lacking; tepals 5, subequal, persistent in fruit, spreading, ovate to obovate, 817 ×
<quantity id="4CE377B8FF14FF48FBA6FE24A9C4EBCC" box="[1151,1251,411,437]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="177" pageNumber="178" unit="mm" value="4.5" valueMax="7.0" valueMin="2.0">27 mm</quantity>
, apex obtuse to rounded, white, glabrous, margin entire, aciliate; ovary body obovoid, 38 ×
<quantity id="4CE377B8FF14FF48FB5DFE00A9CCEBA3" box="[1156,1259,447,474]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="177" pageNumber="178" unit="mm" value="4.5" valueMax="7.0" valueMin="2.0">27 mm</quantity>
, pale green to pink, pubescent, unequally 3-winged, largest wing rectangular, 815 ×
<quantity id="4CE377B8FF14FF48FBF6FE5CA985EB87" box="[1071,1186,483,510]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="11.0" metricValueMin="5.0" pageId="177" pageNumber="178" unit="mm" value="8.0" valueMax="11.0" valueMin="5.0">511 mm</quantity>
, smallest rib-like,
<quantity id="4CE377B8FF14FF48FF66FDB7AC06E85A" box="[191,289,520,547]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="177" pageNumber="178" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
wide; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, free,
<quantity id="4CE377B8FF14FF48FF21FD93AC7FE83E" box="[248,344,556,583]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="177" pageNumber="178" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 mm</quantity>
long, irregularly 3 to 5 times-divided, stigmatic papillae in a spirally twisted band.
<emphasis id="B96F064FFF14FF48FACBFD92AC34E812" italics="true" pageId="177" pageNumber="178">Fruiting pedicel</emphasis>
to
<quantity id="4CE377B8FF14FF48FEE1FDEEACACE812" box="[312,395,593,620]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="177" pageNumber="178" unit="mm" value="35.0">35 mm</quantity>
long.
<emphasis id="B96F064FFF14FF48FE0CFDEDAF75E812" box="[469,594,593,619]" italics="true" pageId="177" pageNumber="178">Fruit body</emphasis>
obovoid, to 8 ×
<quantity id="4CE377B8FF14FF48FCCCFDEEAE7BE812" box="[789,860,593,619]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="177" pageNumber="178" unit="mm" value="7.0">7 mm</quantity>
, drying brown, wings same shape as in ovary, the largest expanding to 17 ×
<quantity id="4CE377B8FF14FF48FDC0FDC9AF48E8E9" box="[537,623,630,656]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="177" pageNumber="178" unit="mm" value="10.0">10 mm</quantity>
, the smallest expanding to
<quantity id="4CE377B8FF14FF48FC74FDC9AED6E8E9" box="[941,1009,630,656]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="177" pageNumber="178" unit="mm" value="3.0">3 mm</quantity>
wide.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF14FF48FF64FD07AE3DE97B" pageId="177" pageNumber="178" type="conservation">
<paragraph id="8BA4DA5DFF14FF48FF64FD07AF4AE8AB" blockId="177.[189,621,696,722]" box="[189,621,696,722]" pageId="177" pageNumber="178">
<heading id="D0EC6D31FF14FF48FF64FD07AF4AE8AB" bold="true" box="[189,621,696,722]" fontSize="11" level="3" pageId="177" pageNumber="178" reason="3">
<emphasis id="B96F064FFF14FF48FF64FD07AF4AE8AB" bold="true" box="[189,621,696,722]" pageId="177" pageNumber="178">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF14FF48FF64FD57AE3DE97B" blockId="177.[189,794,744,771]" box="[189,794,744,771]" pageId="177" pageNumber="178">
Assessed by
<bibRefCitation id="EF8AA7ACFF14FF48FE8DFD57ACDBE97A" author="Tebbitt M. C." box="[340,508,744,771]" journalOrPublisher="Novon" pageId="177" pageNumber="178" pagination="479 - 489" part="23" refId="ref232441" refString="Tebbitt M. C. 2015. Two new species of Andean tuberous Begonia in the B. octopetala group (Begoniaceae). Novon 23 (4): 479 - 489. https: // doi. org / 10.3417 / 2013027" title="Two new species of Andean tuberous Begonia in the B. octopetala group (Begoniaceae)" type="journal article" year="2015">Tebbitt (2015)</bibRefCitation>
as Data Deficient (DD).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF14FF48FF64FC94AF78ED71" pageId="177" pageNumber="178" type="discussion">
<paragraph id="8BA4DA5DFF14FF48FF64FC94AC23E93C" blockId="177.[189,260,811,837]" box="[189,260,811,837]" pageId="177" pageNumber="178">
<heading id="D0EC6D31FF14FF48FF64FC94AC23E93C" bold="true" box="[189,260,811,837]" fontSize="11" level="3" pageId="177" pageNumber="178" reason="3">
<emphasis id="B96F064FFF14FF48FF64FC94AC23E93C" bold="true" box="[189,260,811,837]" pageId="177" pageNumber="178">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF14FF48FF64FCE5AF0DEC7D" blockId="177.[188,1401,858,1541]" pageId="177" pageNumber="178">
<bibRefCitation id="EF8AA7ACFF14FF48FF64FCE5AC42E90C" author="Tebbitt M. C." box="[189,357,858,885]" journalOrPublisher="Novon" pageId="177" pageNumber="178" pagination="479 - 489" part="23" refId="ref232441" refString="Tebbitt M. C. 2015. Two new species of Andean tuberous Begonia in the B. octopetala group (Begoniaceae). Novon 23 (4): 479 - 489. https: // doi. org / 10.3417 / 2013027" title="Two new species of Andean tuberous Begonia in the B. octopetala group (Begoniaceae)" type="journal article" year="2015">Tebbitt (2015)</bibRefCitation>
described
<taxonomicName id="4C1BA1DEFF14FF48FE3FFCE4AFC6E90D" box="[486,737,858,884]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="pseudopleiopetala">
<emphasis id="B96F064FFF14FF48FE3FFCE4AFC6E90D" box="[486,737,858,884]" italics="true" pageId="177" pageNumber="178">B. pseudopleiopetala</emphasis>
</taxonomicName>
to encompass specimens from
<collectingRegion id="49DF14BFFF14FF48FB8DFCE5A9B2E90D" box="[1108,1173,858,884]" country="Peru" name="Piura" pageId="177" pageNumber="178">Piura</collectingRegion>
,
<collectingRegion id="49DF14BFFF14FF48FB78FCE5A839E90D" box="[1185,1310,858,884]" country="Peru" name="Cajamarca" pageId="177" pageNumber="178">Cajamarca</collectingRegion>
and
<collectingRegion id="49DF14BFFF14FF48FA81FCE4AC07E9E0" country="Peru" name="La Libertad" pageId="177" pageNumber="178">La Libertad</collectingRegion>
Regions that superficially resembled
<taxonomicName id="4C1BA1DEFF14FF48FD35FC3FAEB1E9E0" box="[748,918,895,921]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="pleiopetala">
<emphasis id="B96F064FFF14FF48FD35FC3FAEB1E9E0" box="[748,918,895,921]" italics="true" pageId="177" pageNumber="178">B. pleiopetala</emphasis>
</taxonomicName>
but with several consistent differences.
<taxonomicName id="4C1BA1DEFF14FF48FF64FC1BAF23E9C4" box="[189,516,931,957]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="pseudopleiopetala">
<emphasis id="B96F064FFF14FF48FF64FC1BAF23E9C4" box="[189,516,931,957]" italics="true" pageId="177" pageNumber="178">Begonia pseudopleiopetala</emphasis>
</taxonomicName>
had acuminate (vs rounded to acute) leaf apices; obovate (vs elliptic to oblong) anthers lacking extended connectives; and 5 tepals on the pistillate flowers (vs 79). We have since collected several specimens of
<taxonomicName id="4C1BA1DEFF14FF48FDA0FC52AE06EE7F" box="[633,801,1004,1030]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="pleiopetala">
<emphasis id="B96F064FFF14FF48FDA0FC52AE06EE7F" box="[633,801,1004,1030]" italics="true" pageId="177" pageNumber="178">B. pleiopetala</emphasis>
</taxonomicName>
from central
<collectingCountry id="F30C9ACDFF14FF48FC1FFC52AEDBEE7E" box="[966,1020,1005,1031]" name="Peru" pageId="177" pageNumber="178">Peru</collectingCountry>
(
<collectingRegion id="49DF14BFFF14FF48FBD4FC53A974EE7F" box="[1037,1107,1004,1030]" country="Peru" name="Pasco" pageId="177" pageNumber="178">Pasco</collectingRegion>
and
<collectingRegion id="49DF14BFFF14FF48FB48FC53A9F6EE7F" box="[1169,1233,1004,1030]" country="Peru" name="Junin" pageId="177" pageNumber="178">Junín</collectingRegion>
Regions) that share several of these characteristics. All were collected from &lt;
<quantity id="4CE377B8FF14FF48FC75FBAEA923EE52" box="[940,1028,1041,1067]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="177" pageNumber="178" unit="m" value="2000.0">2000 m</quantity>
a.s.l. in elevation on seasonally wet cliffs in lower and middle montane forest rather than the species usual high elevation grassland habitat. These specimens variously have acuminate leaf apices (
<emphasis id="B96F064FFF14FF48FC6DFBE5A80FEE0A" box="[948,1320,1113,1140]" italics="true" pageId="177" pageNumber="178">P.W. Moonlight &amp; A. Daza 236</emphasis>
,
<emphasis id="B96F064FFF14FF48FAE1FBE5A843EE0D" box="[1336,1380,1114,1140]" italics="true" pageId="177" pageNumber="178">257</emphasis>
); anthers lacking extended connectives (
<emphasis id="B96F064FFF14FF48FD4AFBC0A932EEE1" box="[659,1045,1150,1176]" italics="true" pageId="177" pageNumber="178">P.W. Moonlight &amp; A. Daza 236</emphasis>
,
<emphasis id="B96F064FFF14FF48FBF1FBC1A973EEE1" box="[1064,1108,1150,1176]" italics="true" pageId="177" pageNumber="178">257</emphasis>
); as few as 4 tepals on the pistillate flower (
<emphasis id="B96F064FFF14FF48FE67FB1CAE65EEC5" box="[446,834,1186,1213]" italics="true" pageId="177" pageNumber="178">P.W. Moonlight &amp; A. Daza 295</emphasis>
,
<emphasis id="B96F064FFF14FF48FC8FFB1DAEA5EEC5" box="[854,898,1186,1212]" italics="true" pageId="177" pageNumber="178">297</emphasis>
); and other unusual characters including two-locular ovaries (
<emphasis id="B96F064FFF14FF48FE60FB77AE1EEE98" box="[441,825,1223,1249]" italics="true" pageId="177" pageNumber="178">P.W. Moonlight &amp; A. Daza 295</emphasis>
,
<emphasis id="B96F064FFF14FF48FC95FB78AE5FEE98" box="[844,888,1223,1249]" italics="true" pageId="177" pageNumber="178">297</emphasis>
); but no individual has more than two of these characters. These characters may be the result of stress in marginal habitats, but limited evidence suggests this is not the case. Seed collected from the collection
<emphasis id="B96F064FFF14FF48FC44FAAFA82EEF50" box="[925,1289,1295,1322]" italics="true" pageId="177" pageNumber="178">P.W. Moonlight &amp; A. Daza 236</emphasis>
grown at the Royal Botanic Gardens Edinburgh retained the acuminate leaf apices and anthers lacking extended connectives of its parent plant (see
<figureCitation id="1320C6D8FF14FF48FDB1FAE7AFE7EF0A" box="[616,704,1368,1395]" captionStart="Fig" captionStartId="170.[189,232,1752,1778]" captionTargetBox="[283,1303,265,1708]" captionTargetId="figure-13@170.[283,1303,265,1708]" captionTargetPageId="170" captionText="Fig. 55. Begonia pleiopetala A.DC. A. Habit. B. Section of inflorescence and bracts. C. Leaf, adaxial surface. D. Leaf, abaxial surface. E. Tuber. F. Staminate flower, side view. G. Staminate flower, front view. H. Staminate flower, back view. I. Smallest tepal of staminate flower. J. Largest tepal of staminate flower. K. Androecium, side view. L. Pistillate flower, side view. M. Pistillate flower, front view. N. Pistillate flower, back view. O. Cross section of ovary. P. Smallest tepal of pistillate flower. Q. Largest tepal of pistillate flower. R. Pistils, side view. S. Pistil, side view. All photographs taken by D.A. Purvis in the living collections of the Royal Botanic Garden Edinburgh (Accession 20160140, grown from seeds collected as part of P.W. Moonlight &amp; A. Daza 236)." figureDoi="http://doi.org/10.5281/zenodo.8178419" httpUri="https://zenodo.org/record/8178419/files/figure.png" pageId="177" pageNumber="178">Fig. 55</figureCitation>
), suggesting these characters are not simply a response to atypical growing conditions in the wild. None of these specimens have all four of the characteristics used by Tebbitt to separate
<taxonomicName id="4C1BA1DEFF14FF48FE2CFA1DAFC9EFC2" box="[501,750,1441,1467]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="pseudopleiopetala">
<emphasis id="B96F064FFF14FF48FE2CFA1DAFC9EFC2" box="[501,750,1441,1467]" italics="true" pageId="177" pageNumber="178">B. pseudopleiopetala</emphasis>
</taxonomicName>
from
<taxonomicName id="4C1BA1DEFF14FF48FCECFA1DAEFDEFC2" box="[821,986,1441,1467]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="pleiopetala">
<emphasis id="B96F064FFF14FF48FCECFA1DAEFDEFC2" box="[821,986,1441,1467]" italics="true" pageId="177" pageNumber="178">B. pleiopetala</emphasis>
</taxonomicName>
so we retain it as a different species but suggest further genetic and morphological work may be required to determine whether they should be retained as separate species.
</paragraph>
<paragraph id="8BA4DA5DFF14FF48FF64F992AC97EC3E" blockId="177.[189,432,1581,1607]" box="[189,432,1581,1607]" pageId="177" pageNumber="178">
<heading id="D0EC6D31FF14FF48FF64F992AC97EC3E" bold="true" box="[189,432,1581,1607]" fontSize="11" level="3" pageId="177" pageNumber="178" reason="3">
<emphasis id="B96F064FFF14FF48FF64F992AC97EC3E" bold="true" box="[189,432,1581,1607]" pageId="177" pageNumber="178">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF14FF48FF64F9E1AF78ED71" blockId="177.[188,1399,1629,1800]" pageId="177" pageNumber="178">
<taxonomicName id="4C1BA1DEFF14FF48FF64F9E1ACD8EC0E" box="[189,511,1629,1655]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="pseudopleiopetala">
<emphasis id="B96F064FFF14FF48FF64F9E1ACD8EC0E" box="[189,511,1629,1655]" italics="true" pageId="177" pageNumber="178">Begonia pseudopleiopetala</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C1BA1DEFF14FF48FD4EF9E1AE1BEC0E" box="[663,828,1629,1655]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="pleiopetala">
<emphasis id="B96F064FFF14FF48FD4EF9E1AE1BEC0E" box="[663,828,1629,1655]" italics="true" pageId="177" pageNumber="178">B. pleiopetala</emphasis>
</taxonomicName>
in its leaves with an acuminate apex (vs rounded to acute and rarely acuminate), its pistillate flowers with five tepals (vs [5]79 tepals), and its obovate (vs elliptic to oblong) anthers with non-extended (vs non-extended to extended) connectives. Some specimens of
<taxonomicName id="4C1BA1DEFF14FF48FEBEF974AF36EC9D" box="[359,529,1738,1764]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="pleiopetala">
<emphasis id="B96F064FFF14FF48FEBEF974AF36EC9D" box="[359,529,1738,1764]" italics="true" pageId="177" pageNumber="178">B. pleiopetala</emphasis>
</taxonomicName>
share one or rarely two of these characters (see Notes) but no known specimens share all four characters.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF14FF48FF64F88EAF1AED90" pageId="177" pageNumber="178" type="distribution">
<paragraph id="8BA4DA5DFF14FF48FF64F88EACD2ED32" blockId="177.[189,501,1841,1867]" box="[189,501,1841,1867]" pageId="177" pageNumber="178">
<heading id="D0EC6D31FF14FF48FF64F88EACD2ED32" bold="true" box="[189,501,1841,1867]" fontSize="11" level="3" pageId="177" pageNumber="178" reason="3">
<emphasis id="B96F064FFF14FF48FF64F88EACD2ED32" bold="true" box="[189,501,1841,1867]" pageId="177" pageNumber="178">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF14FF48FF64F8DEAF1AED90" blockId="177.[189,1399,1889,2025]" pageId="177" pageNumber="178">
Endemic to
<collectingCountry id="F30C9ACDFF14FF48FE8FF8DDACABED05" box="[342,396,1890,1916]" name="Peru" pageId="177" pageNumber="178">Peru</collectingCountry>
and known from
<collectingRegion id="49DF14BFFF14FF48FDAAF8DEAF93ED02" box="[627,692,1889,1915]" country="Peru" name="Piura" pageId="177" pageNumber="178">Piura</collectingRegion>
,
<collectingRegion id="49DF14BFFF14FF48FD1CF8DEAE61ED02" box="[709,838,1889,1915]" country="Peru" name="Cajamarca" pageId="177" pageNumber="178">Cajamarca</collectingRegion>
, and
<collectingRegion id="49DF14BFFF14FF48FC57F8DDA93AED02" box="[910,1053,1889,1916]" country="Peru" name="La Libertad" pageId="177" pageNumber="178">La Libertad</collectingRegion>
Regions (
<figureCitation id="1320C6D8FF14FF48FB79F8DEA835ED05" box="[1184,1298,1889,1916]" captionStart="Fig" captionStartId="160.[189,232,1822,1848]" captionTargetBox="[194,1391,558,1776]" captionTargetId="figure-14@160.[189,1398,548,1784]" captionTargetPageId="160" captionText="Fig. 52. Distribution of the octopetala group of Begonia sect. Eupetalum (Lindl.) A.DC. in Peru and surrounding countries. A. B. anemoniflora Irmsch. (red) and B. tumbezensis Irmsch. (blue). B. B. octopetala L.Hér. subsp. octopetala (red) and B. octopetala subsp. ovatiformis Irmsch. (blue). C. B. pleiopetala A.DC. (red) and B. pseudopleiopetala Tebbitt (blue). D. B. polypetala A.DC. (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178413" httpUri="https://zenodo.org/record/8178413/files/figure.png" pageId="177" pageNumber="178">Fig. 52C</figureCitation>
). Found within northwest Peruvian relict montane forest at elevations from
<quantity id="4CE377B8FF14FF48FBCCF83AA998EDD9" box="[1045,1215,1925,1952]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.5" metricValueMin="1.3" pageId="177" pageNumber="178" unit="m" value="1900.0" valueMax="2500.0" valueMin="1300.0">13002500 m</quantity>
a.s.l.
<taxonomicName id="4C1BA1DEFF14FF48FACDF839ACB1EDBD" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="pseudopleiopetala">
<emphasis id="B96F064FFF14FF48FACDF839ACB1EDBD" italics="true" pageId="177" pageNumber="178">Begonia pseudopleiopetala</emphasis>
</taxonomicName>
is a geophytic species and dies down to its tuber in the dry season. It flowers during the wet season in March to June.
</paragraph>
</subSubSection>
</treatment>
</document>