treatments-xml/data/03/B0/87/03B087E7FFB96860CEB98F03A0A7FD9D.xml

<document id="9ABA5A3A0D9F4F7930EF0474B7828CB0" ID-DOI="10.11646/zootaxa.3948.3.9" ID-GBIF-Dataset="bdfc1a75-65c6-4294-895b-eb952625ebe5" ID-ISSN="1175-5326" ID-Zenodo-Dep="239206" ID-ZooBank="BA8ED874-6B20-43B4-B728-132122B1821A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461044970624" checkinUser="plazi" docAuthor="González-Moreno, Alejandra, Bordera, Santiago &amp; Sääksjärvi, Ilari Eerikki" docDate="2015" docId="03B087E7FFB96860CEB98F03A0A7FD9D" docLanguage="en" docName="zt03948p586.pdf" docOrigin="Zootaxa 3948 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Labena littoralis Gonzalez-Moreno &amp; Bordera, sp. nov." docType="treatment" docVersion="8" lastPageNumber="580" masterDocId="FF89FF9FFFB86868CE2E8B5AA370FFFD" masterDocTitle="Description of three new species of Labena Cresson from Mexico (Hymenoptera, Ichneumonidae, Labeninae), with notes on tropical species richness" masterLastPageNumber="586" masterPageNumber="573" pageNumber="574" updateTime="1698576490452" updateUser="plazi">
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<mods:title id="61B703344EE42D796E45A49A60C6801B">Description of three new species of Labena Cresson from Mexico (Hymenoptera, Ichneumonidae, Labeninae), with notes on tropical species richness</mods:title>
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<mods:namePart id="EE539948F44A3D5258691A50A24376B5">González-Moreno, Alejandra</mods:namePart>
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<mods:namePart id="D2F39CF46F0816F11FD46BB792C7CB7F">Bordera, Santiago</mods:namePart>
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<mods:namePart id="F0AA58F4C81178FCE07A511FFC1718DB">Sääksjärvi, Ilari Eerikki</mods:namePart>
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<mods:date id="F0ED5CBF5AA3C62B8EE6542A684C625D">2015</mods:date>
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<treatment id="03B087E7FFB96860CEB98F03A0A7FD9D" ID-DOI="http://doi.org/10.5281/zenodo.6121267" ID-GBIF-Taxon="119610359" ID-Zenodo-Dep="6121267" LSID="urn:lsid:plazi:treatment:03B087E7FFB96860CEB98F03A0A7FD9D" httpUri="http://treatment.plazi.org/id/03B087E7FFB96860CEB98F03A0A7FD9D" lastPageId="8" lastPageNumber="580" pageId="1" pageNumber="574">
<subSubSection id="C303657AFFB96869CEB98F03A1E3FB68" pageId="1" pageNumber="574" type="nomenclature">
<paragraph id="8BA636F1FFB96869CEB98F03A031FB8E" blockId="1.[151,833,1113,1173]" box="[151,833,1113,1139]" pageId="1" pageNumber="574">
<heading id="D0EE819DFFB96869CEB98F03A031FB8E" bold="true" box="[151,833,1113,1139]" fontSize="11" level="1" pageId="1" pageNumber="574" reason="1">
<emphasis id="B96DEAE3FFB96869CEB98F03A031FB8E" bold="true" box="[151,833,1113,1139]" pageId="1" pageNumber="574">
<taxonomicName id="4C194D72FFB96869CEB98F03A1A7FB8E" authority="Gonzalez-Moreno &amp; Bordera" authorityName="Gonzalez-Moreno &amp; Bordera" box="[151,727,1113,1139]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="574" phylum="Arthropoda" rank="species" species="littoralis" status="sp. nov.">
<emphasis id="B96DEAE3FFB96869CEB98F03A22EFB8E" bold="true" box="[151,350,1113,1139]" italics="true" pageId="1" pageNumber="574">Labena littoralis</emphasis>
González-Moreno &amp; Bordera
</taxonomicName>
, 
<taxonomicNameLabel id="A25E5798FFB96869CCCA8F03A031FB8E" box="[740,833,1113,1139]" pageId="1" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA636F1FFB96869CEB98F26A1E3FB68" blockId="1.[151,833,1113,1173]" box="[151,659,1148,1173]" pageId="1" pageNumber="574">
(
<figureCitation id="13222A74FFB96869CEB18F26A391FB69" box="[159,225,1148,1173]" captionStart="FIGURE 1" captionStartId="2.[151,250,1882,1904]" captionTargetBox="[151,1436,195,1857]" captionTargetId="figure@2.[151,1436,195,1858]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Labena spp.: A— L. littoralis (♀ holotype); B— L. marginata (♀); C— L. littoralis (♂ paratype); D— L. marginata (♂); E— L. tekalina (♂ paratype); F— L. madoricola (♂ holotype); G— L. schausi (♂)." httpUri="https://zenodo.org/record/239207/files/figure.png" pageId="1" pageNumber="574">Figs 1</figureCitation>
A, 1C, 2A, 2C, 2E, 3A, 3C, 3E, 3G, 6)
</paragraph>
</subSubSection>
<subSubSection id="C303657AFFB96869CEB98F9EA1F2F9B9" pageId="1" pageNumber="574" type="diagnosis">
<paragraph id="8BA636F1FFB96869CEB98F9EA1F2F9B9" blockId="1.[151,1437,1220,2037]" pageId="1" pageNumber="574">
<emphasis id="B96DEAE3FFB96869CEB98F9EA261FB20" bold="true" box="[151,273,1220,1245]" pageId="1" pageNumber="574">Diagnosis.</emphasis>
<taxonomicName id="4C194D72FFB96869CF338F9EA2AEFB21" box="[285,478,1220,1244]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="574" phylum="Arthropoda" rank="species" species="littoralis">
<emphasis id="B96DEAE3FFB96869CF338F9EA2AEFB21" box="[285,478,1220,1244]" italics="true" pageId="1" pageNumber="574">Labena littoralis</emphasis>
</taxonomicName>
can be distinguished from all New world species by the following characters in combination: metapleuron without longitudinal striae (
<figureCitation id="13222A74FFB96869CCD28FBDA034FAFD" box="[764,836,1255,1280]" captionStart="FIGURE 2" captionStartId="3.[151,250,1770,1792]" captionTargetBox="[152,1434,193,1747]" captionTargetId="figure@3.[152,1435,193,1749]" captionTargetPageId="3" captionText="FIGURE 2. Propodeum and tergite I of Labena spp., female: A— L. littoralis, propodeum, dorsal view (holotype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (holotype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (holotype); F— L. marginata, tergite I, dorsal view." httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="1" pageNumber="574">Figs 2</figureCitation>
C, 3C); flagellum, in dorsal view, orange, basally and apically dark brown to black; fore wing hyaline, yellowish tinged, without a subapical black spot (
<figureCitation id="13222A74FFB96869CAD08E56A632FAD9" box="[1278,1346,1292,1317]" captionStart="FIGURE 1" captionStartId="2.[151,250,1882,1904]" captionTargetBox="[151,1436,195,1857]" captionTargetId="figure@2.[151,1436,195,1858]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Labena spp.: A— L. littoralis (♀ holotype); B— L. marginata (♀); C— L. littoralis (♂ paratype); D— L. marginata (♂); E— L. tekalina (♂ paratype); F— L. madoricola (♂ holotype); G— L. schausi (♂)." httpUri="https://zenodo.org/record/239207/files/figure.png" pageId="1" pageNumber="574">Figs 1</figureCitation>
A, 1C); sternite I without a central transverse ridge; propodeum yellow with most part of carinae black (
<figureCitation id="13222A74FFB96869CAF68E75A650FAB5" box="[1240,1312,1327,1352]" captionStart="FIGURE 2" captionStartId="3.[151,250,1770,1792]" captionTargetBox="[152,1434,193,1747]" captionTargetId="figure@3.[152,1435,193,1749]" captionTargetPageId="3" captionText="FIGURE 2. Propodeum and tergite I of Labena spp., female: A— L. littoralis, propodeum, dorsal view (holotype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (holotype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (holotype); F— L. marginata, tergite I, dorsal view." httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="1" pageNumber="574">Figs 2</figureCitation>
A, 2C, 3A, 3C). Metasomal tergites from third to the apex, dark brown to black anteriorly, yellow banded posteriorly, in males tending to be light brown centrally (
<figureCitation id="13222A74FFB96869CC048E2DA11CFA6D" box="[554,620,1399,1424]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Propodeum and tergites of Labena spp., male: A— L. littoralis, propodeum, dorsal view (paratype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (paratype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (paratype); F— L. marginata, tergite I, dorsal view G— L. littoralis, tergites III—IV, dorsal view (paratype); H— L. marginata, tergites III—IV, dorsal view." httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="1" pageNumber="574">Fig. 3</figureCitation>
G). Propodeum of female (
<figureCitation id="13222A74FFB96869CDB78E2DA0ABFA6D" box="[921,987,1399,1424]" captionStart="FIGURE 2" captionStartId="3.[151,250,1770,1792]" captionTargetBox="[152,1434,193,1747]" captionTargetId="figure@3.[152,1435,193,1749]" captionTargetPageId="3" captionText="FIGURE 2. Propodeum and tergite I of Labena spp., female: A— L. littoralis, propodeum, dorsal view (holotype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (holotype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (holotype); F— L. marginata, tergite I, dorsal view." httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="1" pageNumber="574">Fig. 2</figureCitation>
A) with area basalis very transverse, 2.7 × as broad as long; area superomedia hexagonal, not strongly narrowed posteriorly; area petiolaris not defined by median lateral carinae so confluent with area posteroexterna and dentipara; area externa longer than broad; area lateralis conspicuously shorter than broad, very transverse (
<figureCitation id="13222A74FFB96869CD6B8EBEA0FBFA01" box="[837,907,1508,1533]" captionStart="FIGURE 2" captionStartId="3.[151,250,1770,1792]" captionTargetBox="[152,1434,193,1747]" captionTargetId="figure@3.[152,1435,193,1749]" captionTargetPageId="3" captionText="FIGURE 2. Propodeum and tergite I of Labena spp., female: A— L. littoralis, propodeum, dorsal view (holotype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (holotype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (holotype); F— L. marginata, tergite I, dorsal view." httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="1" pageNumber="574">Fig. 2</figureCitation>
C). Tergite I of female about 2.2 × as long as posteriorly broad (
<figureCitation id="13222A74FFB96869CF428D5DA2C2F9DD" box="[364,434,1543,1568]" captionStart="FIGURE 2" captionStartId="3.[151,250,1770,1792]" captionTargetBox="[152,1434,193,1747]" captionTargetId="figure@3.[152,1435,193,1749]" captionTargetPageId="3" captionText="FIGURE 2. Propodeum and tergite I of Labena spp., female: A— L. littoralis, propodeum, dorsal view (holotype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (holotype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (holotype); F— L. marginata, tergite I, dorsal view." httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="1" pageNumber="574">Fig. 2</figureCitation>
E). Tergite I of male sub-parallel sided (
<figureCitation id="13222A74FFB96869CDAA8D5DA0B9F9DD" box="[900,969,1543,1568]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Propodeum and tergites of Labena spp., male: A— L. littoralis, propodeum, dorsal view (paratype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (paratype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (paratype); F— L. marginata, tergite I, dorsal view G— L. littoralis, tergites III—IV, dorsal view (paratype); H— L. marginata, tergites III—IV, dorsal view." httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="1" pageNumber="574">Fig. 3</figureCitation>
E). Tergites III-IV with posterior margin deeply concave and membranous (
<figureCitation id="13222A74FFB96869CC358D76A12DF9B9" box="[539,605,1580,1605]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Propodeum and tergites of Labena spp., male: A— L. littoralis, propodeum, dorsal view (paratype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (paratype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (paratype); F— L. marginata, tergite I, dorsal view G— L. littoralis, tergites III—IV, dorsal view (paratype); H— L. marginata, tergites III—IV, dorsal view." httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="1" pageNumber="574">Fig. 3</figureCitation>
G).
</paragraph>
</subSubSection>
<subSubSection id="C303657AFFB9686ECEE98D15A055F988" lastPageId="6" lastPageNumber="579" pageId="1" pageNumber="574" type="description">
<paragraph id="8BA636F1FFB96869CEE98D15A7BCF99A" blockId="1.[151,1437,1220,2037]" box="[199,1228,1615,1640]" pageId="1" pageNumber="574">
<emphasis id="B96DEAE3FFB96869CEE98D15A229F995" bold="true" box="[199,345,1615,1640]" pageId="1" pageNumber="574">Description.</emphasis>
<emphasis id="B96DEAE3FFB96869CF718D0AA2CBF995" box="[351,443,1616,1640]" italics="true" pageId="1" pageNumber="574">Female.</emphasis>
Body length (excluding ovipositor) 
<quantity id="4CE19B14FFB96869CD7D8D15A0D5F99A" box="[851,933,1615,1640]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="574" unit="mm" value="15.0">15 mm</quantity>
. Fore wing length 
<quantity id="4CE19B14FFB96869CA5B8D0AA7B8F99A" box="[1141,1224,1616,1640]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="574" unit="mm" value="12.0">12 mm</quantity>
.
</paragraph>
<paragraph id="8BA636F1FFB96869CEE98D2EA188F808" blockId="1.[151,1437,1220,2037]" pageId="1" pageNumber="574">Head. In dorsal view, about 2.1 × as wide as long; clypeus in anterior view, basally flat, with a broad shallow impression either side of slightly convex central apical region, with apical margin centrally strongly convex, about 1.8 as broad as high; clypeo-facial suture with scattered and long setae; malar space 0.3 × as long as basal width of mandible; frons flat, shiny, with very fine setiferous punctures; vertex and gena smooth and shiny with very fine setiferous punctures; gena, in dorsal view, short, abruptly constricted behind eyes, about 0.3 × as long as eye; lower part of gena very broad; interocellar area elevated; posterior ocellus separated from eye by 1.6 × its own maximum diameter; distance between posterior ocelli about 1.6 × maximum diameter of posterior ocellus; occipital carina complete, reaching hypostomal carina far from base of mandible, at a distance of 0.8 × the width of mandible base; hypostomal carina from base of mandible to the union with occipital carina strongly raised; antenna with 43 flagellomeres; subbasal flagellomeres more or less elongate, subapical ones subquadrate, apical flagellomere subtriangular with apex bearing a small flat sensillum.</paragraph>
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<paragraph id="8BA636F1FFBA686ACEB98C00A701F872" blockId="2.[151,1437,1882,1935]" pageId="2" pageNumber="575">
<emphasis id="B96DEAE3FFBA686ACEB98C00A263F892" bold="true" box="[151,275,1882,1904]" pageId="2" pageNumber="575">FIGURE 1.</emphasis>
Habitus of 
<taxonomicName id="4C194D72FFBA686ACFA78C00A2A5F88D" box="[393,469,1882,1904]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE3FFBA686ACFA78C00A2A5F88D" box="[393,469,1882,1904]" italics="true" pageId="2" pageNumber="575">Labena</emphasis>
</taxonomicName>
spp.: A— 
<taxonomicName id="4C194D72FFBA686ACC108C01A1C0F88D" box="[574,688,1882,1904]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="species" species="littoralis">
<emphasis id="B96DEAE3FFBA686ACC108C01A1C0F88D" box="[574,688,1882,1904]" italics="true" pageId="2" pageNumber="575">L. littoralis</emphasis>
</taxonomicName>
(♀ holotype); B— 
<taxonomicName id="4C194D72FFBA686ACD5E8C01A084F88D" box="[880,1012,1883,1904]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="B96DEAE3FFBA686ACD5E8C01A084F88D" box="[880,1012,1883,1904]" italics="true" pageId="2" pageNumber="575">L. marginata</emphasis>
</taxonomicName>
(♀); C— 
<taxonomicName id="4C194D72FFBA686ACA7B8C01A7B7F88D" box="[1109,1223,1882,1904]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="species" species="littoralis">
<emphasis id="B96DEAE3FFBA686ACA7B8C01A7B7F88D" box="[1109,1223,1882,1904]" italics="true" pageId="2" pageNumber="575">L. littoralis</emphasis>
</taxonomicName>
(♂ paratype); D— 
<taxonomicName id="4C194D72FFBA686ACBA98C01A270F872" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="B96DEAE3FFBA686ACBA98C01A270F872" italics="true" pageId="2" pageNumber="575">L. marginata</emphasis>
</taxonomicName>
(♂); E— 
<taxonomicName id="4C194D72FFBA686ACF4E8C20A2BCF872" box="[352,460,1913,1935]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="species" species="tekalina">
<emphasis id="B96DEAE3FFBA686ACF4E8C20A2BCF872" box="[352,460,1913,1935]" italics="true" pageId="2" pageNumber="575">L. tekalina</emphasis>
</taxonomicName>
(♂ paratype); F— 
<taxonomicName id="4C194D72FFBA686ACCA68C20A068F872" box="[648,792,1913,1935]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="species" species="madoricola">
<emphasis id="B96DEAE3FFBA686ACCA68C20A068F872" box="[648,792,1913,1935]" italics="true" pageId="2" pageNumber="575">L. madoricola</emphasis>
</taxonomicName>
(♂ holotype); G— 
<taxonomicName id="4C194D72FFBA686ACDF48C20A730F872" box="[986,1088,1913,1935]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="species" species="schausi">
<emphasis id="B96DEAE3FFBA686ACDF48C20A730F872" box="[986,1088,1913,1935]" italics="true" pageId="2" pageNumber="575">L. schausi</emphasis>
</taxonomicName>
(♂).
</paragraph>
</caption>
<caption id="DF666679FFBB686BCEB98DB0A7DBF8C0" httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="3" pageNumber="576" targetBox="[152,1434,193,1747]" targetPageId="3">
<paragraph id="8BA636F1FFBB686BCEB98DB0A7DBF8C0" blockId="3.[151,1437,1770,1854]" pageId="3" pageNumber="576">
<emphasis id="B96DEAE3FFBB686BCEB98DB0A266F902" bold="true" box="[151,278,1770,1792]" pageId="3" pageNumber="576">FIGURE 2.</emphasis>
Propodeum and tergite I of 
<taxonomicName id="4C194D72FFBB686BCC698DB0A1E3F8FD" box="[583,659,1770,1792]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE3FFBB686BCC698DB0A1E3F8FD" box="[583,659,1770,1792]" italics="true" pageId="3" pageNumber="576">Labena</emphasis>
</taxonomicName>
spp., female: A— 
<emphasis id="B96DEAE3FFBB686BCD768DB1A0A4F8FD" box="[856,980,1770,1792]" italics="true" pageId="3" pageNumber="576">
<taxonomicName id="4C194D72FFBB686BCD768DB1A0A0F8FD" box="[856,976,1770,1792]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="littoralis">L. littoralis</taxonomicName>
,
</emphasis>
propodeum, dorsal view (holotype); B— 
<emphasis id="B96DEAE3FFBB686BCBA98DB1A276F8E2" italics="true" pageId="3" pageNumber="576">
<taxonomicName id="4C194D72FFBB686BCBA98DB1A272F8E2" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="marginata">L. marginata</taxonomicName>
,
</emphasis>
propodeum, dorsal view; C— 
<emphasis id="B96DEAE3FFBB686BCC628C50A1BAF8E2" box="[588,714,1801,1823]" italics="true" pageId="3" pageNumber="576">
<taxonomicName id="4C194D72FFBB686BCC628C50A1B6F8E2" box="[588,710,1801,1823]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="littoralis">L. littoralis</taxonomicName>
,
</emphasis>
propodeum, lateral view (holotype); D— 
<emphasis id="B96DEAE3FFBB686BCAA68C50A667F8E2" box="[1160,1303,1802,1823]" italics="true" pageId="3" pageNumber="576">
<taxonomicName id="4C194D72FFBB686BCAA68C50A663F8E2" box="[1160,1299,1802,1823]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="marginata">L. marginata</taxonomicName>
,
</emphasis>
propodeum, lateral view; E— 
<emphasis id="B96DEAE3FFBB686BCF648C73A2B2F8C0" box="[330,450,1831,1854]" italics="true" pageId="3" pageNumber="576">
<taxonomicName id="4C194D72FFBB686BCF648C73A2CFF8C0" box="[330,447,1831,1854]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="littoralis">L. littoralis</taxonomicName>
,
</emphasis>
tergite I, dorsal view (holotype); F— 
<emphasis id="B96DEAE3FFBB686BCD6A8C73A0BEF8C3" box="[836,974,1833,1854]" italics="true" pageId="3" pageNumber="576">
<taxonomicName id="4C194D72FFBB686BCD6A8C73A0BAF8C3" box="[836,970,1833,1854]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="marginata">L. marginata</taxonomicName>
,
</emphasis>
tergite I, dorsal view.
</paragraph>
</caption>
<caption id="DF666679FFBC686CCEB98C0FA0B2F83A" httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="4" pageNumber="577" targetBox="[151,1436,193,1855]" targetPageId="4">
<paragraph id="8BA636F1FFBC686CCEB98C0FA0B2F83A" blockId="4.[151,1437,1877,1991]" pageId="4" pageNumber="577">
<emphasis id="B96DEAE3FFBC686CCEB98C0FA268F897" bold="true" box="[151,280,1877,1899]" pageId="4" pageNumber="577">FIGURE 3.</emphasis>
Propodeum and tergites of 
<taxonomicName id="4C194D72FFBC686CCC658C0FA1E7F896" box="[587,663,1877,1899]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE3FFBC686CCC658C0FA1E7F896" box="[587,663,1877,1899]" italics="true" pageId="4" pageNumber="577">Labena</emphasis>
</taxonomicName>
spp., male: A— 
<emphasis id="B96DEAE3FFBC686CCD638C0CA0BBF896" box="[845,971,1877,1899]" italics="true" pageId="4" pageNumber="577">
<taxonomicName id="4C194D72FFBC686CCD638C0CA0B7F896" box="[845,967,1877,1899]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="littoralis">L. littoralis</taxonomicName>
,
</emphasis>
propodeum, dorsal view (paratype); B— 
<emphasis id="B96DEAE3FFBC686CCBA98C0CA276F877" italics="true" pageId="4" pageNumber="577">
<taxonomicName id="4C194D72FFBC686CCBA98C0CA272F877" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="marginata">L. marginata</taxonomicName>
,
</emphasis>
propodeum, dorsal view; C— 
<emphasis id="B96DEAE3FFBC686CCC638C2FA1BBF877" box="[589,715,1908,1930]" italics="true" pageId="4" pageNumber="577">
<taxonomicName id="4C194D72FFBC686CCC638C2FA1B7F877" box="[589,711,1908,1930]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="littoralis">L. littoralis</taxonomicName>
,
</emphasis>
propodeum, lateral view (paratype); D— 
<emphasis id="B96DEAE3FFBC686CCAA68C2FA667F877" box="[1160,1303,1909,1930]" italics="true" pageId="4" pageNumber="577">
<taxonomicName id="4C194D72FFBC686CCAA68C2FA663F877" box="[1160,1299,1909,1930]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="marginata">L. marginata</taxonomicName>
,
</emphasis>
propodeum, lateral view; E— 
<emphasis id="B96DEAE3FFBC686CCF648CC9A2B2F855" box="[330,450,1938,1960]" italics="true" pageId="4" pageNumber="577">
<taxonomicName id="4C194D72FFBC686CCF648CC9A2CEF855" box="[330,446,1938,1960]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="littoralis">L. littoralis</taxonomicName>
,
</emphasis>
tergite I, dorsal view (paratype); F— 
<emphasis id="B96DEAE3FFBC686CCD6C8CC9A0BBF855" box="[834,971,1939,1960]" italics="true" pageId="4" pageNumber="577">
<taxonomicName id="4C194D72FFBC686CCD6C8CC9A0B7F855" box="[834,967,1939,1960]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="marginata">L. marginata</taxonomicName>
,
</emphasis>
tergite I, dorsal view G— 
<emphasis id="B96DEAE3FFBC686CCAFA8CC9A63CF855" box="[1236,1356,1938,1960]" italics="true" pageId="4" pageNumber="577">
<taxonomicName id="4C194D72FFBC686CCAFA8CC9A638F855" box="[1236,1352,1938,1960]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="littoralis">L. littoralis</taxonomicName>
,
</emphasis>
tergites III—IV, dorsal view (paratype); H— 
<emphasis id="B96DEAE3FFBC686CCC3E8CE8A1EAF83A" box="[528,666,1970,1991]" italics="true" pageId="4" pageNumber="577">
<taxonomicName id="4C194D72FFBC686CCC3E8CE8A1E6F83A" box="[528,662,1970,1991]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="marginata">L. marginata</taxonomicName>
,
</emphasis>
tergites III—IV, dorsal view.
</paragraph>
</caption>
<caption id="DF666679FFBD686DCEB98EDBA2ADFA4B" httpUri="https://zenodo.org/record/239210/files/figure.png" pageId="5" pageNumber="578" targetBox="[151,1436,193,1387]" targetPageId="5">
<paragraph id="8BA636F1FFBD686DCEB98EDBA2ADFA4B" blockId="5.[151,1437,1409,1462]" pageId="5" pageNumber="578">
<emphasis id="B96DEAE3FFBD686DCEB98EDBA265FA6B" bold="true" box="[151,277,1409,1431]" pageId="5" pageNumber="578">FIGURE 4.</emphasis>
Propodeum of 
<taxonomicName id="4C194D72FFBD686DCF978EDBA175FA6A" box="[441,517,1409,1431]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE3FFBD686DCF978EDBA175FA6A" box="[441,517,1409,1431]" italics="true" pageId="5" pageNumber="578">Labena</emphasis>
</taxonomicName>
spp., dorsal view: A— 
<taxonomicName id="4C194D72FFBD686DCCD78ED8A018FA6A" box="[761,872,1409,1431]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="tekalina">
<emphasis id="B96DEAE3FFBD686DCCD78ED8A018FA6A" box="[761,872,1409,1431]" italics="true" pageId="5" pageNumber="578">L. tekalina</emphasis>
</taxonomicName>
(♂ holotype); B— 
<taxonomicName id="4C194D72FFBD686DCA018ED8A7B1FA6A" box="[1071,1217,1409,1431]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="madoricola">
<emphasis id="B96DEAE3FFBD686DCA018ED8A7B1FA6A" box="[1071,1217,1409,1431]" italics="true" pageId="5" pageNumber="578">L. madoricola</emphasis>
</taxonomicName>
(♂ paratype); C— 
<taxonomicName id="4C194D72FFBD686DCBA98ED8A393FA4B" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="schausi">
<emphasis id="B96DEAE3FFBD686DCBA98ED8A393FA4B" italics="true" pageId="5" pageNumber="578">L. schausi</emphasis>
</taxonomicName>
(♀); D— 
<taxonomicName id="4C194D72FFBD686DCF688EFBA2DDFA4B" box="[326,429,1440,1462]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="schausi">
<emphasis id="B96DEAE3FFBD686DCF688EFBA2DDFA4B" box="[326,429,1440,1462]" italics="true" pageId="5" pageNumber="578">L. schausi</emphasis>
</taxonomicName>
(♂).
</paragraph>
</caption>
<paragraph id="8BA636F1FFBD686ECEE98E84A1A7FF05" blockId="5.[151,1437,1501,2030]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="579" pageId="5" pageNumber="578">
Mesosoma. Pronotum smooth and shiny with upper lateral corner with dense fine setiferous punctures and lower part, glabrous; in dorsal view, with two lateromedian pits centrally; epomia absent; mesoscutum finely and densely punctate, with notauli absent, virtually without transverse crest at anterior end; scutellum densely punctate, slightly convex, without lateral carinae; mesopleuron smooth and shiny, with upper end of epicnemial carina curved towards anterior margin slightly below the centre of hind margin of pronotum; metapleuron smooth and shiny, posteriorly with very fine and sparse setiferous punctures, setae long (
<figureCitation id="13222A74FFBD686DCA2A8DCBA739F957" box="[1028,1097,1681,1706]" captionStart="FIGURE 2" captionStartId="3.[151,250,1770,1792]" captionTargetBox="[152,1434,193,1747]" captionTargetId="figure@3.[152,1435,193,1749]" captionTargetPageId="3" captionText="FIGURE 2. Propodeum and tergite I of Labena spp., female: A— L. littoralis, propodeum, dorsal view (holotype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (holotype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (holotype); F— L. marginata, tergite I, dorsal view." httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="5" pageNumber="578">Fig. 2</figureCitation>
C); propodeum (
<figureCitation id="13222A74FFBD686DCB2B8DCBA63EF957" box="[1285,1358,1681,1706]" captionStart="FIGURE 2" captionStartId="3.[151,250,1770,1792]" captionTargetBox="[152,1434,193,1747]" captionTargetId="figure@3.[152,1435,193,1749]" captionTargetPageId="3" captionText="FIGURE 2. Propodeum and tergite I of Labena spp., female: A— L. littoralis, propodeum, dorsal view (holotype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (holotype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (holotype); F— L. marginata, tergite I, dorsal view." httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="5" pageNumber="578">Figs 2</figureCitation>
A, 2C) smooth and shiny with very dense setiferous punctures anteriorly to posterior transverse carina, except in area superomedia centrally; pleural carina absent anteriorly, so that area spiracularis is open externally, present posteriorly and enclosing area lateralis; in dorsal view, with area superomedia hexagonal clearly delineated anteriorly, posteriorly less strongly defined, but always separated from area petiolaris, about 1.3 × as broad as long; area basalis transverse, 2.7 × as broad as long; area dentipara confluent with area posteroexterna and area petiolaris; lateral longitudinal carina complete, area externa slightly elongate, fully enclosed, about 0.9 × as broad as long; area lateralis transverse, ventrally quite shorter than dorsally. Mid leg with tibia inflated in distal 0.7, with scattered bristles on outer surface, second tarsomere about 0.9 × as long as broad and as same length of the following two tarsomeres; hind coxa about 1.8 × as long as deep; hind tibia with scattered bristles. Fore wing with areolet transverse, 0.44 × as broad as length of distal abscissa of M, with 3rs-m subequal to 2rs-m; vein 
<emphasis id="B96DEAE3FFBE686ECB4A8BC2A6ECFF52" box="[1380,1436,152,175]" italics="true" pageId="6" pageNumber="579">Cu-a</emphasis>
opposite to vein 
<emphasis id="B96DEAE3FFBE686ECF778BE7A205FF29" box="[345,373,189,212]" italics="true" pageId="6" pageNumber="579">Rs</emphasis>
&amp; 
<emphasis id="B96DEAE3FFBE686ECFA48BE7A2D1FF29" box="[394,417,189,212]" italics="true" pageId="6" pageNumber="579">M</emphasis>
; abscissa of 
<emphasis id="B96DEAE3FFBE686ECC148BE6A12AFF2E" box="[570,602,188,211]" italics="true" pageId="6" pageNumber="579">Cu</emphasis>
1 between 1 
<emphasis id="B96DEAE3FFBE686ECCC98BE7A06FFF29" box="[743,799,189,212]" italics="true" pageId="6" pageNumber="579">m-cu</emphasis>
and 
<emphasis id="B96DEAE3FFBE686ECD748BE6A00BFF2E" box="[858,891,188,211]" italics="true" pageId="6" pageNumber="579">Cu</emphasis>
1 
<emphasis id="B96DEAE3FFBE686ECDA78BE7A0E7FF29" box="[905,919,189,212]" italics="true" pageId="6" pageNumber="579">a</emphasis>
as long as 
<emphasis id="B96DEAE3FFBE686ECA308BE6A74FFF2E" box="[1054,1087,188,211]" italics="true" pageId="6" pageNumber="579">Cu</emphasis>
1 
<emphasis id="B96DEAE3FFBE686ECA638BE6A72BFF29" box="[1101,1115,188,212]" italics="true" pageId="6" pageNumber="579">b</emphasis>
. 
<collectingCountry id="F30E7661FFBE686ECA458BE6A7D3FF28" box="[1131,1187,188,213]" name="India" pageId="6" pageNumber="579">Hind</collectingCountry>
wing with vein 
<emphasis id="B96DEAE3FFBE686ECB4A8BE6A6ECFF2E" box="[1380,1436,188,211]" italics="true" pageId="6" pageNumber="579">Cu-a</emphasis>
longer than abscissa of 
<emphasis id="B96DEAE3FFBE686ECFB38BBAA299FF0A" box="[413,489,224,247]" italics="true" pageId="6" pageNumber="579">M+Cu</emphasis>
between 
<emphasis id="B96DEAE3FFBE686ECC7A8BBBA11BFF05" box="[596,619,225,248]" italics="true" pageId="6" pageNumber="579">M</emphasis>
and 
<emphasis id="B96DEAE3FFBE686ECC8C8BBAA1B3FF0A" box="[674,707,224,247]" italics="true" pageId="6" pageNumber="579">Cu</emphasis>
1.
</paragraph>
<paragraph id="8BA636F1FFBE686ECEE98A5FA203FE50" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="579">
Metasoma. Tergite I, about 2.2 × as long as posteriorly broad, without lateromedian and lateral carinae (
<figureCitation id="13222A74FFBE686ECB418A5EA3D6FEBD" captionStart="FIGURE 2" captionStartId="3.[151,250,1770,1792]" captionTargetBox="[152,1434,193,1747]" captionTargetId="figure@3.[152,1435,193,1749]" captionTargetPageId="3" captionText="FIGURE 2. Propodeum and tergite I of Labena spp., female: A— L. littoralis, propodeum, dorsal view (holotype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (holotype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (holotype); F— L. marginata, tergite I, dorsal view." httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="6" pageNumber="579">Fig. 2</figureCitation>
E); sternite I short, reaching 0.4 of length of tergite, without a central transverse ridge; tergite II, 0.8 × as long as posteriorly broad, with fine and very dense pubescence; ovipositor sheath 1.8 × as long as hind tibia (
<figureCitation id="13222A74FFBE686ECB1C8A16A603FE99" box="[1330,1395,332,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1882,1904]" captionTargetBox="[151,1436,195,1857]" captionTargetId="figure@2.[151,1436,195,1858]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Labena spp.: A— L. littoralis (♀ holotype); B— L. marginata (♀); C— L. littoralis (♂ paratype); D— L. marginata (♂); E— L. tekalina (♂ paratype); F— L. madoricola (♂ holotype); G— L. schausi (♂)." httpUri="https://zenodo.org/record/239207/files/figure.png" pageId="6" pageNumber="579">Fig. 1</figureCitation>
A), strongly laterally compressed, the apex of the lower valve with fine file-like teeth, and with a small scabrous area basal to these teeth.
</paragraph>
<paragraph id="8BA636F1FFBE686ECEE98AEDA7BEFCCA" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="579">
Coloration (
<figureCitation id="13222A74FFBE686ECF608AEDA2FEFE2D" box="[334,398,439,464]" captionStart="FIGURE 1" captionStartId="2.[151,250,1882,1904]" captionTargetBox="[151,1436,195,1857]" captionTargetId="figure@2.[151,1436,195,1858]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Labena spp.: A— L. littoralis (♀ holotype); B— L. marginata (♀); C— L. littoralis (♂ paratype); D— L. marginata (♂); E— L. tekalina (♂ paratype); F— L. madoricola (♂ holotype); G— L. schausi (♂)." httpUri="https://zenodo.org/record/239207/files/figure.png" pageId="6" pageNumber="579">Fig. 1</figureCitation>
A). A predominantly yellow species. Head with black marks on gena along occipital carina, a transverse band through the ocelli, and longitudinal stripe centrally at occiput; scape and pedicel dorsally dark brown; flagellum basally dark brown, dorsally progressively orange until flagellomere 22, from flagellomere 23 to apex dark brown, ventrally more widely orange; mesosoma with three longitudinal black marks on mesoscutum confluent with prescutellar groove; black marks on peripheries of mesoscutum, part of tegula, posterior band on scutellum, transverse furrow at base of propodeum, and all anterior carinae of propodeum from posterior transverse carina, except abscissae closing area superomedia posteriorly; dorsal and anterior periphery on mesopleuron, upper division on metapleuron; tergite I of metasoma infuscate subapically; tergite II basally with bilobed infuscate mark, following tergites with anterior 0.6 or more, black. Legs with a light brown stripe on posterior side of front and mid femora and base of tibiae; hind coxae, trochanters, femora and tibiae; hind tarsi infuscate. Ovipositor sheath light brown, dark brown at apex. Fore wing hyaline, yellowish tinged, without a subapical black spot.
</paragraph>
<paragraph id="8BA636F1FFBE686ECEE9881EA02DFCA0" blockId="6.[151,1437,151,2013]" box="[199,861,836,861]" pageId="6" pageNumber="579">
<emphasis id="B96DEAE3FFBE686ECEE9881EA271FCA1" box="[199,257,836,860]" italics="true" pageId="6" pageNumber="579">Male</emphasis>
. Similar as female except in the following characters:
</paragraph>
<paragraph id="8BA636F1FFBE686ECEE9883DA18AFC82" blockId="6.[151,1437,151,2013]" box="[199,762,871,896]" pageId="6" pageNumber="579">
Body length 
<quantity id="4CE19B14FFBE686ECF798832A2B4FC82" box="[343,452,872,896]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="6" pageNumber="579" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
. Fore wing length 
<quantity id="4CE19B14FFBE686ECCBB883DA186FC82" box="[661,758,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="6" pageNumber="579" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
.
</paragraph>
<paragraph id="8BA636F1FFBE686ECEE988D6A0EFFAA9" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="579">
Head 1.8–2.0 × as broad as long; clypeus 1.7–1.8 × as broad as high; malar space 0.19–0.23 × as long as basal width of mandible; gena slightly rounded and constricted behind eyes, 0.27–0.35 × as long as eye, lower part of gena moderately broad; posterior ocellus separated from eye by 1.4–1.7 × its own maximum diameter; distance between posterior ocelli 1.5–2.0 × maximum diameter of posterior ocellus; occipital carina reaching hypostomal carina far from base of mandible at a distance of 0.75–0.77 × the width of mandible base; antenna with 40–45 elongate flagellomeres. Area superomedia pentagonal clearly delineated, 1.0–1.1 × as broad as long; area petiolaris obliterated by median longitudinal carina; area basalis cuadrangular, 1.25 × as broad as long (
<figureCitation id="13222A74FFBE686ECADB8F3EA64BFB81" box="[1269,1339,1124,1149]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Propodeum and tergites of Labena spp., male: A— L. littoralis, propodeum, dorsal view (paratype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (paratype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (paratype); F— L. marginata, tergite I, dorsal view G— L. littoralis, tergites III—IV, dorsal view (paratype); H— L. marginata, tergites III—IV, dorsal view." httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="6" pageNumber="579">Fig. 3</figureCitation>
A); area dentipara separated from area posteroexterna; area externa 0.86–0.89 × as broad as long (
<figureCitation id="13222A74FFBE686ECAB88FDDA7AAFB5D" box="[1174,1242,1159,1184]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Propodeum and tergites of Labena spp., male: A— L. littoralis, propodeum, dorsal view (paratype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (paratype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (paratype); F— L. marginata, tergite I, dorsal view G— L. littoralis, tergites III—IV, dorsal view (paratype); H— L. marginata, tergites III—IV, dorsal view." httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="6" pageNumber="579">Fig. 3</figureCitation>
A). Mid leg with tibia not inflated, second tarsomere elongate, 6.0 × as long as broad, 1.0–1.1 × as long as following two tarsomeres; hind coxa about 3.8–4.5 × as long as deep. Fore wing with areolet, 0.30–0.38 × as broad as length of distal abscissa of M. Tergite I, 3.4–3.9 × as long as posteriorly broad, sub-parallel sided (
<figureCitation id="13222A74FFBE686ECDE38FAEA77FFAF1" box="[973,1039,1268,1293]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Propodeum and tergites of Labena spp., male: A— L. littoralis, propodeum, dorsal view (paratype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (paratype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (paratype); F— L. marginata, tergite I, dorsal view G— L. littoralis, tergites III—IV, dorsal view (paratype); H— L. marginata, tergites III—IV, dorsal view." httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="6" pageNumber="579">Fig. 3</figureCitation>
E); sternite I, reaching 0.5 of length of tergite; tergite II, 1.9–2.0 × as long as posteriorly broad, with fine and moderately dense pubescence; tergites III– IV with posterior margin deeply concave and membranous (
<figureCitation id="13222A74FFBE686ECD168E66A00BFAA9" box="[824,891,1340,1365]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Propodeum and tergites of Labena spp., male: A— L. littoralis, propodeum, dorsal view (paratype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (paratype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (paratype); F— L. marginata, tergite I, dorsal view G— L. littoralis, tergites III—IV, dorsal view (paratype); H— L. marginata, tergites III—IV, dorsal view." httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="6" pageNumber="579">Fig. 3</figureCitation>
G).
</paragraph>
<paragraph id="8BA636F1FFBE686ECEE98E05A61FF9F5" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="579">
Coloration (
<figureCitation id="13222A74FFBE686ECF638E05A2FCFA85" box="[333,396,1375,1400]" captionStart="FIGURE 1" captionStartId="2.[151,250,1882,1904]" captionTargetBox="[151,1436,195,1857]" captionTargetId="figure@2.[151,1436,195,1858]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Labena spp.: A— L. littoralis (♀ holotype); B— L. marginata (♀); C— L. littoralis (♂ paratype); D— L. marginata (♂); E— L. tekalina (♂ paratype); F— L. madoricola (♂ holotype); G— L. schausi (♂)." httpUri="https://zenodo.org/record/239207/files/figure.png" pageId="6" pageNumber="579">Fig. 1</figureCitation>
C). As female but pedicel orange; flagellum orange with former and apical 4–6 flagellomeres dark brown to black; all carinae of propodeum (
<figureCitation id="13222A74FFBE686ECCED8EDEA07CFA61" box="[707,780,1412,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Propodeum and tergites of Labena spp., male: A— L. littoralis, propodeum, dorsal view (paratype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (paratype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (paratype); F— L. marginata, tergite I, dorsal view G— L. littoralis, tergites III—IV, dorsal view (paratype); H— L. marginata, tergites III—IV, dorsal view." httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="6" pageNumber="579">Figs 3</figureCitation>
A, 3C) dark brown to black, except posterior abscissa of lateral longitudinal carina separating area posteroexterna from area coxalis, posterior carina enclosing area superomedia and longitudinal carina obliterating area petiolaris; tergite II dark brown to black in anterior half; front and mid legs entirely yellow; hind leg mostly light brown, tibia infuscate at distal part, tarsomeres dark brown.
</paragraph>
<paragraph id="8BA636F1FFBE686ECEE98D4EA055F988" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="579">
<emphasis id="B96DEAE3FFBE686ECEE98D4EA24BF9D0" bold="true" box="[199,315,1556,1581]" pageId="6" pageNumber="579">Variation.</emphasis>
Some males have a black spot at central part of mesopleuron and first tergite not infuscate and/or black marks of tergites tending to be brown, posteriorly. Specimen from French 
<collectingCountry id="F30E7661FFBE686ECA038D62A7F4F9AD" box="[1069,1156,1592,1616]" name="Guyana" pageId="6" pageNumber="579">Guyana</collectingCountry>
has antenna dark brown ventrally, and tergites from V to apex entirely dark brown.
</paragraph>
</subSubSection>
<subSubSection id="C303657AFFBE686FCEE98D25A005F92D" lastPageId="7" lastPageNumber="580" pageId="6" pageNumber="579" type="etymology">
<paragraph id="8BA636F1FFBE686ECEE98D25A792F965" blockId="6.[151,1437,151,2013]" box="[199,1250,1663,1688]" pageId="6" pageNumber="579">
<emphasis id="B96DEAE3FFBE686ECEE98D25A23EF965" bold="true" box="[199,334,1663,1688]" pageId="6" pageNumber="579">Etymology.</emphasis>
The name refers to the littoral area of Yucatán, where the 
<typeStatus id="54A28853FFBE686ECDF78D25A74BF965" box="[985,1083,1663,1688]" pageId="6" pageNumber="579" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
<paragraph id="8BA636F1FFBE686ECEE98DFEA2C5F940" blockId="6.[151,1437,151,2013]" box="[199,437,1700,1725]" pageId="6" pageNumber="579">
<emphasis id="B96DEAE3FFBE686ECEE98DFEA239F940" bold="true" box="[199,329,1700,1725]" pageId="6" pageNumber="579">Phenology.</emphasis>
<figureCitation id="13222A74FFBE686ECF7E8DFEA2C1F940" box="[336,433,1700,1725]" captionStart="FIGURE 6" captionStartId="8.[151,250,1409,1431]" captionTargetBox="[150,1425,667,1378]" captionTargetId="figure@8.[150,1435,663,1381]" captionTargetPageId="8" captionText="FIGURE 6. Phenology of L. littoralis." httpUri="https://zenodo.org/record/239212/files/figure.png" pageId="6" pageNumber="579">Figure 6</figureCitation>
.
</paragraph>
<paragraph id="8BA636F1FFBE686ECEE98D9DA09BF8F8" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="579">
<emphasis id="B96DEAE3FFBE686ECEE98D9DA2CDF91D" bold="true" box="[199,445,1735,1760]" pageId="6" pageNumber="579">Taxonomic remarks.</emphasis>
In comparison with the other Mexican species, 
<taxonomicName id="4C194D72FFBE686ECDF48D93A728F91D" box="[986,1112,1736,1760]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="littoralis">
<emphasis id="B96DEAE3FFBE686ECDF48D93A728F91D" box="[986,1112,1736,1760]" italics="true" pageId="6" pageNumber="579">L. littoralis</emphasis>
</taxonomicName>
is very close to 
<taxonomicName id="4C194D72FFBE686ECB3F8D93A6ECF922" box="[1297,1436,1736,1760]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="B96DEAE3FFBE686ECB3F8D93A6ECF922" box="[1297,1436,1736,1760]" italics="true" pageId="6" pageNumber="579">L marginata</emphasis>
</taxonomicName>
but can be distinguished from it by the following characters in combination:
</paragraph>
<paragraph id="8BA636F1FFBE686ECEE98C4BA6F6F821" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="579">
<emphasis id="B96DEAE3FFBE686ECEE98C4BA220F8D5" box="[199,336,1808,1832]" italics="true" pageId="6" pageNumber="579">
<taxonomicName id="4C194D72FFBE686ECEE98C4BA238F8D5" box="[199,328,1808,1832]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="littoralis">L. littoralis</taxonomicName>
:
</emphasis>
Propodeum (
<figureCitation id="13222A74FFBE686ECFC78C55A141F8D5" box="[489,561,1807,1832]" captionStart="FIGURE 2" captionStartId="3.[151,250,1770,1792]" captionTargetBox="[152,1434,193,1747]" captionTargetId="figure@3.[152,1435,193,1749]" captionTargetPageId="3" captionText="FIGURE 2. Propodeum and tergite I of Labena spp., female: A— L. littoralis, propodeum, dorsal view (holotype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (holotype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (holotype); F— L. marginata, tergite I, dorsal view." httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="6" pageNumber="579">Figs 2</figureCitation>
A, 2C, 3A, 3C) with area basalis very transverse, in female 2.7 × as broad as long, in male, 1.25 × as broad as long; area petiolaris not defined by median lateral carinae, confluent with area posteroexterna and dentipara; area externa longer than broad; area lateralis conspicuously shorter than broad, very transverse. Tergite I of metasoma about 2.2 × as long as posteriorly broad in female (
<figureCitation id="13222A74FFBE686ECA678C26A7FBF869" box="[1097,1163,1916,1941]" captionStart="FIGURE 2" captionStartId="3.[151,250,1770,1792]" captionTargetBox="[152,1434,193,1747]" captionTargetId="figure@3.[152,1435,193,1749]" captionTargetPageId="3" captionText="FIGURE 2. Propodeum and tergite I of Labena spp., female: A— L. littoralis, propodeum, dorsal view (holotype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (holotype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (holotype); F— L. marginata, tergite I, dorsal view." httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="6" pageNumber="579">Fig. 2</figureCitation>
E); in male, 3.4–3.9 × as long as posteriorly broad, sub-parallel sided (
<figureCitation id="13222A74FFBE686ECCB28CFAA190F845" box="[668,736,1952,1977]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Propodeum and tergites of Labena spp., male: A— L. littoralis, propodeum, dorsal view (paratype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (paratype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (paratype); F— L. marginata, tergite I, dorsal view G— L. littoralis, tergites III—IV, dorsal view (paratype); H— L. marginata, tergites III—IV, dorsal view." httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="6" pageNumber="579">Fig. 3</figureCitation>
E). Male with flagellum orange, dorsally at base and apically dark brown to black (
<figureCitation id="13222A74FFBE686ECFA98C9EA2B6F821" box="[391,454,1988,2013]" captionStart="FIGURE 1" captionStartId="2.[151,250,1882,1904]" captionTargetBox="[151,1436,195,1857]" captionTargetId="figure@2.[151,1436,195,1858]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Labena spp.: A— L. littoralis (♀ holotype); B— L. marginata (♀); C— L. littoralis (♂ paratype); D— L. marginata (♂); E— L. tekalina (♂ paratype); F— L. madoricola (♂ holotype); G— L. schausi (♂)." httpUri="https://zenodo.org/record/239207/files/figure.png" pageId="6" pageNumber="579">Fig. 1</figureCitation>
C). Tergites III–IV with posterior margin deeply concave and membranous (
<figureCitation id="13222A74FFBE686ECB318C9EA611F821" box="[1311,1377,1988,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Propodeum and tergites of Labena spp., male: A— L. littoralis, propodeum, dorsal view (paratype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (paratype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (paratype); F— L. marginata, tergite I, dorsal view G— L. littoralis, tergites III—IV, dorsal view (paratype); H— L. marginata, tergites III—IV, dorsal view." httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="6" pageNumber="579">Fig. 3</figureCitation>
G).
</paragraph>
<caption id="DF666679FFBF686FCEB98EDEA7C5FA2A" httpUri="https://zenodo.org/record/239211/files/figure.png" pageId="7" pageNumber="580" targetBox="[151,1436,193,1390]" targetPageId="7">
<paragraph id="8BA636F1FFBF686FCEB98EDEA7C5FA2A" blockId="7.[151,1436,1412,1495]" pageId="7" pageNumber="580">
<emphasis id="B96DEAE3FFBF686FCEB98EDEA262FA64" bold="true" box="[151,274,1412,1434]" pageId="7" pageNumber="580">FIGURE 5.</emphasis>
Propodeum and tergite I of 
<taxonomicName id="4C194D72FFBF686FCC038EDEA109FA67" box="[557,633,1412,1434]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE3FFBF686FCC038EDEA109FA67" box="[557,633,1412,1434]" italics="true" pageId="7" pageNumber="580">Labena</emphasis>
</taxonomicName>
spp.: A— 
<taxonomicName id="4C194D72FFBF686FCCCF8EDFA03DFA67" box="[737,845,1412,1434]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="tekalina">
<emphasis id="B96DEAE3FFBF686FCCCF8EDFA03DFA67" box="[737,845,1412,1434]" italics="true" pageId="7" pageNumber="580">L. tekalina</emphasis>
</taxonomicName>
(♂ paratype), propodeum, lateral view; B— 
<taxonomicName id="4C194D72FFBF686FCB238EDFA6ECFA67" box="[1293,1436,1412,1434]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="madoricola">
<emphasis id="B96DEAE3FFBF686FCB238EDFA6ECFA67" box="[1293,1436,1412,1434]" italics="true" pageId="7" pageNumber="580">L. madoricola</emphasis>
</taxonomicName>
(♂ holotype), propodeum, lateral view; C— 
<taxonomicName id="4C194D72FFBF686FCC4D8EFEA1A1FA44" box="[611,721,1443,1465]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="tekalina">
<emphasis id="B96DEAE3FFBF686FCC4D8EFEA1A1FA44" box="[611,721,1443,1465]" italics="true" pageId="7" pageNumber="580">L. tekalina</emphasis>
</taxonomicName>
(♂ holotype), tergite I, dorsal view; D— 
<taxonomicName id="4C194D72FFBF686FCA548EFEA67BFA44" box="[1146,1291,1443,1465]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="madoricola">
<emphasis id="B96DEAE3FFBF686FCA548EFEA67BFA44" box="[1146,1291,1443,1465]" italics="true" pageId="7" pageNumber="580">L. madoricola</emphasis>
</taxonomicName>
(♂ paratype), tergite I, dorsal view; E— 
<taxonomicName id="4C194D72FFBF686FCFB08E98A174FA2A" box="[414,516,1473,1495]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="schausi">
<emphasis id="B96DEAE3FFBF686FCFB08E98A174FA2A" box="[414,516,1473,1495]" italics="true" pageId="7" pageNumber="580">L. schausi</emphasis>
</taxonomicName>
(♀), tergite I, dorsal view; F— 
<taxonomicName id="4C194D72FFBF686FCD6F8E98A0D8FA2A" box="[833,936,1473,1495]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="schausi">
<emphasis id="B96DEAE3FFBF686FCD6F8E98A0D8FA2A" box="[833,936,1473,1495]" italics="true" pageId="7" pageNumber="580">L. schausi</emphasis>
</taxonomicName>
(♂), tergite I, dorsal view.
</paragraph>
</caption>
<paragraph id="8BA636F1FFBF686FCEE98D5FA005F92D" blockId="7.[151,1436,1540,2033]" pageId="7" pageNumber="580">
<taxonomicName id="4C194D72FFBF686FCEE98D5FA228F9E1" box="[199,344,1541,1564]" class="Insecta" family="Ichneumonidae" genus="Labena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="580" phylum="Arthropoda" rank="species" species="marginata">
<emphasis id="B96DEAE3FFBF686FCEE98D5FA228F9E1" box="[199,344,1541,1564]" italics="true" pageId="7" pageNumber="580">L. marginata</emphasis>
</taxonomicName>
(
<figureCitation id="13222A74FFBF686FCF498D5EA2DBF9E1" box="[359,427,1540,1565]" captionStart="FIGURE 1" captionStartId="2.[151,250,1882,1904]" captionTargetBox="[151,1436,195,1857]" captionTargetId="figure@2.[151,1436,195,1858]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Labena spp.: A— L. littoralis (♀ holotype); B— L. marginata (♀); C— L. littoralis (♂ paratype); D— L. marginata (♂); E— L. tekalina (♂ paratype); F— L. madoricola (♂ holotype); G— L. schausi (♂)." httpUri="https://zenodo.org/record/239207/files/figure.png" pageId="7" pageNumber="580">Figs 1</figureCitation>
B, 1D): Propodeum (
<figureCitation id="13222A74FFBF686FCCB48D5EA192F9E1" box="[666,738,1540,1565]" captionStart="FIGURE 2" captionStartId="3.[151,250,1770,1792]" captionTargetBox="[152,1434,193,1747]" captionTargetId="figure@3.[152,1435,193,1749]" captionTargetPageId="3" captionText="FIGURE 2. Propodeum and tergite I of Labena spp., female: A— L. littoralis, propodeum, dorsal view (holotype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (holotype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (holotype); F— L. marginata, tergite I, dorsal view." httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="7" pageNumber="580">Figs 2</figureCitation>
B, 2D, 3B, 3D) with area basalis not more than twice as broad as long, usually longer than broad in males; area petiolaris defined by median lateral carinae or obliterated; area externa conspicuously broader than long; area lateralis almost as long as wide, slightly transverse. Tergite I of metasoma 1.7–1.8 × as long as posteriorly broad in female (
<figureCitation id="13222A74FFBF686FCD798D2AA0EEF975" box="[855,926,1648,1673]" captionStart="FIGURE 2" captionStartId="3.[151,250,1770,1792]" captionTargetBox="[152,1434,193,1747]" captionTargetId="figure@3.[152,1435,193,1749]" captionTargetPageId="3" captionText="FIGURE 2. Propodeum and tergite I of Labena spp., female: A— L. littoralis, propodeum, dorsal view (holotype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (holotype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (holotype); F— L. marginata, tergite I, dorsal view." httpUri="https://zenodo.org/record/239208/files/figure.png" pageId="7" pageNumber="580">Fig. 2</figureCitation>
F); in male, 2.9-3.1 × as long as posteriorly broad, with divergent sides from base to apex (
<figureCitation id="13222A74FFBF686FCC878DCEA19DF951" box="[681,749,1684,1709]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Propodeum and tergites of Labena spp., male: A— L. littoralis, propodeum, dorsal view (paratype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (paratype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (paratype); F— L. marginata, tergite I, dorsal view G— L. littoralis, tergites III—IV, dorsal view (paratype); H— L. marginata, tergites III—IV, dorsal view." httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="7" pageNumber="580">Fig. 3</figureCitation>
F). Male with flagellum entirely orange. Tergites III–IV with posterior margin straight, not with exposing membrane (
<figureCitation id="13222A74FFBF686FCD218DE2A021F92D" box="[783,849,1720,1745]" captionStart="FIGURE 3" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. Propodeum and tergites of Labena spp., male: A— L. littoralis, propodeum, dorsal view (paratype); B— L. marginata, propodeum, dorsal view; C— L. littoralis, propodeum, lateral view (paratype); D— L. marginata, propodeum, lateral view; E— L. littoralis, tergite I, dorsal view (paratype); F— L. marginata, tergite I, dorsal view G— L. littoralis, tergites III—IV, dorsal view (paratype); H— L. marginata, tergites III—IV, dorsal view." httpUri="https://zenodo.org/record/239209/files/figure.png" pageId="7" pageNumber="580">Fig. 3</figureCitation>
H).
</paragraph>
</subSubSection>
<subSubSection id="C303657AFFBF6860CEE98D86A0A7FD9D" lastPageId="8" lastPageNumber="581" pageId="7" pageNumber="580" type="materials_examined">
<paragraph id="8BA636F1FFBF6860CEE98D86A0A7FD9D" blockId="7.[151,1436,1540,2033]" lastBlockId="8.[151,1437,151,608]" lastPageId="8" lastPageNumber="581" pageId="7" pageNumber="580">
<emphasis id="B96DEAE3FFBF686FCEE98D86A141F908" bold="true" box="[199,561,1756,1781]" pageId="7" pageNumber="580">
Material examined. 
<typeStatus id="54A28853FFBF686FCFEB8D86A141F908" box="[453,561,1756,1781]" pageId="7" pageNumber="580" type="holotype">Holotype</typeStatus>
</emphasis>
female (CER-UADY currently loaned in 
<collectionCode id="ED08AE34FFBF686FCA018D86A7F0F909" box="[1071,1152,1756,1780]" pageId="7" pageNumber="580">CEUA</collectionCode>
), 
<collectingCountry id="F30E7661FFBF686FCAB78D86A780F908" box="[1177,1264,1756,1781]" name="Mexico" pageId="7" pageNumber="580">
<emphasis id="B96DEAE3FFBF686FCAB78D86A780F908" bold="true" box="[1177,1264,1756,1781]" pageId="7" pageNumber="580">Mexico</emphasis>
</collectingCountry>
, Yucatán, Rio Lagartos, Nucleo, 
<date id="FFA71031FFBF686FCF468C5AA283F8E5" box="[360,499,1792,1816]" pageId="7" pageNumber="580" value="2008-12-23">23.XII.2008</date>
– 
<date id="FFA71031FFBF686FCC2D8C5AA11DF8E5" box="[515,621,1792,1816]" pageId="7" pageNumber="580" value="2009-01-07">07.I.2009</date>
, Sabana, TM 7, leg. González. 
<emphasis id="B96DEAE3FFBF686FCDF38C5AA7CDF8E4" bold="true" box="[989,1213,1792,1817]" pageId="7" pageNumber="580">
<typeStatus id="54A28853FFBF686FCDF38C5AA729F8E4" box="[989,1113,1792,1817]" pageId="7" pageNumber="580" type="paratype">Paratypes</typeStatus>
: 
<collectingCountry id="F30E7661FFBF686FCA488C5AA7CDF8E4" box="[1126,1213,1792,1817]" name="Mexico" pageId="7" pageNumber="580">Mexico</collectingCountry>
</emphasis>
, Yucatán, El Cuyo, Cementerio, 
<date id="FFA71031FFBF686FCF1C8C7EA28EF8C1" box="[306,510,1828,1852]" pageId="7" pageNumber="580" value="2009-03-18" valueMax="2009-04-01" valueMin="2009-03-18">18.III–01.IV.2009</date>
, Matorral, TM6, leg. González, 
<specimenCount id="9D1FFD78FFBF686FCDB38C7EA0A2F8C1" box="[925,978,1828,1852]" pageId="7" pageNumber="580" type="generic">1 ♂</specimenCount>
(CER-UADY); same locality, 
<date id="FFA71031FFBF686FCB798C7EA269F89D" pageId="7" pageNumber="580" value="2008-10-22" valueMax="2008-11-10" valueMin="2008-10-22">22.X– 10.XI.2008</date>
, Matorral, TM6, leg. González, 
<specimenCount id="9D1FFD78FFBF686FCC878C12A1ABF89D" box="[681,731,1864,1888]" pageId="7" pageNumber="580" type="generic">2 ♂</specimenCount>
(CER-UADY); El Cuyo, Punta Mecoh, 
<date id="FFA71031FFBF686FCAFE8C12A6E8F89D" box="[1232,1432,1864,1888]" pageId="7" pageNumber="580" value="2008-10-22" valueMax="2008-11-10" valueMin="2008-10-22">22.X–10.XI.2008</date>
, Matorral, TM3, leg. González, 
<specimenCount id="9D1FFD78FFBF686FCFDD8C36A16CF879" box="[499,540,1900,1924]" pageId="7" pageNumber="580" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFBF686FCC048C36A10CF879" box="[554,636,1900,1924]" pageId="7" pageNumber="580">CEUA</collectionCode>
); same locality and date, TM4, leg. González, 
<specimenCount id="9D1FFD78FFBF686FCAAF8C36A7DBF879" box="[1153,1195,1900,1924]" pageId="7" pageNumber="580" type="generic">4 ♂</specimenCount>
(CER-UADY); same locality, 
<date id="FFA71031FFBF686FCED48CCAA2A3F854" box="[250,467,1936,1961]" pageId="7" pageNumber="580" value="2008-11-26" valueMax="2008-12-10" valueMin="2008-11-26">26.XI–10.XII.2008</date>
, Matorral, TM 4, leg. González, 
<specimenCount id="9D1FFD78FFBF686FCD7A8CCAA0F1F855" box="[852,897,1936,1960]" pageId="7" pageNumber="580" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFBF686FCDBC8CCAA095F855" box="[914,997,1936,1960]" pageId="7" pageNumber="580">CEUA</collectionCode>
); same locality, 
<date id="FFA71031FFBF686FCA8F8CCAA65CF855" box="[1185,1324,1936,1960]" pageId="7" pageNumber="580" value="2008-12-23">23.XII.2008</date>
– 
<date id="FFA71031FFBF686FCB128CCAA6E7F855" box="[1340,1431,1936,1960]" pageId="7" pageNumber="580" value="2009-01-07">7.I.2009</date>
, Matorral, TM 4, leg. González, 
<specimenCount id="9D1FFD78FFBF686FCC2B8CEEA141F831" box="[517,561,1972,1996]" pageId="7" pageNumber="580" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFBF686FCC6C8CEEA1E8F831" box="[578,664,1972,1996]" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="7" pageNumber="580">ZMUT</collectionCode>
); same locality, 
<date id="FFA71031FFBF686FCD7F8CEEA085F831" box="[849,1013,1972,1996]" pageId="7" pageNumber="580" value="2009-01-21" valueMax="2009-02-04" valueMin="2009-01-21">21.I–4.II.2009</date>
, Matorral, TM 4, leg. González, 
<specimenCount id="9D1FFD78FFBF686FCB5E8CEEA6ECF831" box="[1392,1436,1972,1996]" pageId="7" pageNumber="580" type="generic">1 ♂</specimenCount>
(CER-UADY); El Cuyo, Sacboh, 
<date id="FFA71031FFBF686FCC308C82A1CAF80D" box="[542,698,2008,2032]" pageId="7" pageNumber="580" value="2008-03-04" valueMax="2008-03-18" valueMin="2008-03-04">4–18.III.2008</date>
, Selva, TM 2, leg. González, 
<specimenCount id="9D1FFD78FFBF686FCA388C82A732F80D" box="[1046,1090,2008,2032]" pageId="7" pageNumber="580" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFBF686FCA7A8C82A7D6F80D" box="[1108,1190,2008,2032]" pageId="7" pageNumber="580">CEUA</collectionCode>
); same locality, 
<date id="FFA71031FFBF6860CB4C8C82A270FF4D" lastPageId="8" lastPageNumber="581" pageId="7" pageNumber="580" value="2009-01-21" valueMax="2009-02-04" valueMin="2009-01-21">21.I– 4.II.2009</date>
, Selva, TM 2, leg. González, 
<specimenCount id="9D1FFD78FFB06860CC798BC2A1F3FF52" box="[599,643,152,176]" pageId="8" pageNumber="581" type="generic">1 ♂</specimenCount>
(CER-UADY); El Cuyo, Tekal, 
<date id="FFA71031FFB06860CDD58BC2A7C9FF4D" box="[1019,1209,152,176]" pageId="8" pageNumber="581" value="2009-03-18" valueMax="2009-04-01" valueMin="2009-03-18">18.III–1.IV.2009</date>
, Selva, TM 12, leg. González, 
<specimenCount id="9D1FFD78FFB06860CF3C8BE6A24FFF29" box="[274,319,188,212]" pageId="8" pageNumber="581" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFB06860CF7E8BE6A2D3FF29" box="[336,419,188,212]" pageId="8" pageNumber="581">CEUA</collectionCode>
); Rio Lagartos, Nucleo, 
<date id="FFA71031FFB06860CCEE8BE6A0EAFF28" box="[704,922,188,213]" pageId="8" pageNumber="581" value="2008-11-26" valueMax="2008-12-10" valueMin="2008-11-26">26.XI–10.XII.2008</date>
, Sabana TM 8, leg González, 
<specimenCount id="9D1FFD78FFB06860CAD08BE6A65BFF29" box="[1278,1323,188,212]" pageId="8" pageNumber="581" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFB06860CB138BE6A6E0FF29" box="[1341,1424,188,212]" pageId="8" pageNumber="581">CEUA</collectionCode>
); same locality, 
<date id="FFA71031FFB06860CF108BBAA283FF05" box="[318,499,224,248]" pageId="8" pageNumber="581" value="2008-12-10" valueMax="2008-12-23" valueMin="2008-12-10">10–23.XII.2008</date>
, Sabana, TM 8, leg. González, 
<specimenCount id="9D1FFD78FFB06860CD4A8BBAA0E1FF0A" box="[868,913,224,248]" pageId="8" pageNumber="581" type="generic">2 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFB06860CD8D8BBAA086FF05" box="[931,1014,224,248]" pageId="8" pageNumber="581">CEUA</collectionCode>
); same locality and date, TM 7, leg. González, 
<specimenCount id="9D1FFD78FFB06860CF218A5EA24AFEE1" box="[271,314,260,284]" pageId="8" pageNumber="581" type="generic">2 ♂</specimenCount>
(CER-UADY); same locality, 
<date id="FFA71031FFB06860CCB68A5EA052FEE1" box="[664,802,260,284]" pageId="8" pageNumber="581" value="2008-12-23">23.XII.2008</date>
– 
<date id="FFA71031FFB06860CD1D8A5EA0EDFEE1" box="[819,925,260,284]" pageId="8" pageNumber="581" value="2009-01-07">07.I.2009</date>
, Sabana, TM 8, leg. González, 
<specimenCount id="9D1FFD78FFB06860CB2C8A5EA65EFEE1" box="[1282,1326,260,284]" pageId="8" pageNumber="581" type="generic">3 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFB06860CB138A5EA6E0FEE1" box="[1341,1424,260,284]" pageId="8" pageNumber="581">CEUA</collectionCode>
); same locality, 
<date id="FFA71031FFB06860CF178A72A2ADFEBD" box="[313,477,296,320]" pageId="8" pageNumber="581" value="2009-01-21" valueMax="2009-02-04" valueMin="2009-01-21">21.I–4.II.2009</date>
, Sabana TM 7, leg González, 
<specimenCount id="9D1FFD78FFB06860CD1C8A72A02DFEC2" box="[818,861,296,320]" pageId="8" pageNumber="581" type="generic">2 ♂</specimenCount>
(CER-UADY); same locality, 
<date id="FFA71031FFB06860CA948A72A64FFEBD" box="[1210,1343,296,320]" pageId="8" pageNumber="581" value="2009-02-04" valueMax="2009-02-08" valueMin="2009-02-04">4–8.II.2009</date>
, Sabana TM 8, leg. González, 
<specimenCount id="9D1FFD78FFB06860CFA48A16A2C4FE99" box="[394,436,332,357]" pageId="8" pageNumber="581" type="generic">5 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFB06860CFEC8A16A165FE99" box="[450,533,332,356]" pageId="8" pageNumber="581">CEUA</collectionCode>
); same locality and date, TM 7, leg. González, 
<specimenCount id="9D1FFD78FFB06860CA0F8A16A73BFE99" box="[1057,1099,332,356]" pageId="8" pageNumber="581" type="generic">1 ♂</specimenCount>
(CER-UADY); same locality, 
<date id="FFA71031FFB06860CEB98A2AA23CFE75" box="[151,332,368,392]" pageId="8" pageNumber="581" value="2009-02-18" valueMax="2009-03-04" valueMin="2009-02-18">18.II–4.III.2009</date>
, Sabana TM 7, leg. González, 
<specimenCount id="9D1FFD78FFB06860CC828A2AA1A6FE7A" box="[684,726,368,392]" pageId="8" pageNumber="581" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFB06860CCC88A2AA049FE75" box="[742,825,368,392]" pageId="8" pageNumber="581">CEUA</collectionCode>
); same locality, 
<date id="FFA71031FFB06860CDDE8A2AA7C8FE75" box="[1008,1208,368,392]" pageId="8" pageNumber="581" value="2009-05-27" valueMax="2009-06-10" valueMin="2009-05-27">27.V–10.VI.2009</date>
, Sabana, TM 8, leg. González, 
<specimenCount id="9D1FFD78FFB06860CF208ACEA248FE51" box="[270,312,404,428]" pageId="8" pageNumber="581" type="generic">1 ♂</specimenCount>
(CER-UADY); Rio Lagartos, Poste, 
<date id="FFA71031FFB06860CCF68ACEA0D9FE50" box="[728,937,404,429]" pageId="8" pageNumber="581" value="2008-11-10" valueMax="2008-11-26" valueMin="2008-11-10">10.XI–26.XI.2008</date>
, Sabana, TM 9, leg. González, 
<specimenCount id="9D1FFD78FFB06860CB2B8ACEA65FFE51" box="[1285,1327,404,428]" pageId="8" pageNumber="581" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFB06860CB138ACEA6E0FE51" box="[1341,1424,404,428]" pageId="8" pageNumber="581">CEUA</collectionCode>
); same locality, 
<date id="FFA71031FFB06860CF158AEDA164FE2D" box="[315,532,439,464]" pageId="8" pageNumber="581" value="2008-11-26" valueMax="2008-12-10" valueMin="2008-11-26">26.XI–10.XII.2008</date>
, Sabana, TM 9, leg. González, 
<specimenCount id="9D1FFD78FFB06860CD538AE2A0D8FE32" box="[893,936,440,464]" pageId="8" pageNumber="581" type="generic">1 ♂</specimenCount>
(CER-UADY); same locality and date, TM 10, leg. González, 
<specimenCount id="9D1FFD78FFB06860CF588A86A2D5FE09" box="[374,421,476,500]" pageId="8" pageNumber="581" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFB06860CF968A86A17BFE09" box="[440,523,476,500]" pageId="8" pageNumber="581">CEUA</collectionCode>
); same locality, 
<date id="FFA71031FFB06860CCE28A86A0F1FE09" box="[716,897,476,500]" pageId="8" pageNumber="581" value="2008-12-10" valueMax="2008-12-23" valueMin="2008-12-10">10–23.XII.2008</date>
, Sabana, TM 9, leg. González, 
<specimenCount id="9D1FFD78FFB06860CAD28A86A65AFE09" box="[1276,1322,476,500]" pageId="8" pageNumber="581" type="generic">2 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFB06860CB138A86A6E0FE09" box="[1341,1424,476,500]" pageId="8" pageNumber="581">CEUA</collectionCode>
); 
<date id="FFA71031FFB06860CEB9895AA252FDE5" box="[151,290,512,536]" pageId="8" pageNumber="581" value="2008-12-23">23.XII.2008</date>
– 
<date id="FFA71031FFB06860CF1F895AA2ECFDE5" box="[305,412,512,536]" pageId="8" pageNumber="581" value="2009-01-07">07.I.2009</date>
, Sabana, TM 9, leg González, 
<specimenCount id="9D1FFD78FFB06860CCDE895AA06AFDEA" box="[752,794,512,536]" pageId="8" pageNumber="581" type="generic">2 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFB06860CD06895AA00BFDE5" box="[808,891,512,536]" pageId="8" pageNumber="581">CEUA</collectionCode>
); 
<date id="FFA71031FFB06860CDA3895AA767FDE5" box="[909,1047,512,536]" pageId="8" pageNumber="581" value="2009-01-07" valueMax="2009-01-21" valueMin="2009-01-07">7–21.I.2009</date>
, Sabana, TM 8, leg. González, 
<specimenCount id="9D1FFD78FFB06860CB5D895AA6ECFDEA" box="[1395,1436,512,536]" pageId="8" pageNumber="581" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFB06860CEB1897EA385FDC1" box="[159,245,548,572]" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="8" pageNumber="581">ZMUT</collectionCode>
); same locality and date, TM 9, leg. González, 
<specimenCount id="9D1FFD78FFB06860CD28897EA040FDC1" box="[774,816,548,572]" pageId="8" pageNumber="581" type="generic">3 ♂</specimenCount>
(
<specimenCount id="9D1FFD78FFB06860CD6E897EA014FDC1" box="[832,868,548,572]" pageId="8" pageNumber="581" type="generic">2♂</specimenCount>
<collectionCode id="ED08AE34FFB06860CD45897EA0CAFDC1" box="[875,954,548,572]" pageId="8" pageNumber="581">CEUA</collectionCode>
, 
<specimenCount id="9D1FFD78FFB06860CDE8897EA080FDC1" box="[966,1008,548,572]" pageId="8" pageNumber="581" type="generic">1 ♂</specimenCount>
<collectionCode id="ED08AE34FFB06860CDD9897EA73CFDC1" box="[1015,1100,548,572]" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="8" pageNumber="581">ZMUT</collectionCode>
). 
<emphasis id="B96DEAE3FFB06860CA70897EA386FD9D" bold="true" pageId="8" pageNumber="581">
Non 
<typeStatus id="54A28853FFB06860CAB9897EA7BAFDC0" box="[1175,1226,548,573]" pageId="8" pageNumber="581">type</typeStatus>
material: French 
<collectingCountry id="F30E7661FFB06860CEB9891DA386FD9D" box="[151,246,583,608]" name="Guyana" pageId="8" pageNumber="581">Guyana</collectingCountry>
</emphasis>
, RN Trinité, 
<date id="FFA71031FFB06860CFA5891DA15BFD9D" box="[395,555,583,608]" pageId="8" pageNumber="581" value="2012-05-26" valueMax="2012-05-28" valueMin="2012-05-26">26–28.V.2012</date>
, sweeping, 
<collectionCode id="ED08AE34FFB06860CC838912A186FD9D" box="[685,758,584,608]" pageId="8" pageNumber="581">SEAG</collectionCode>
coll., 
<specimenCount id="9D1FFD78FFB06860CD128912A016FDA2" box="[828,870,584,608]" pageId="8" pageNumber="581" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED08AE34FFB06860CD5B8912A0BBFD9D" box="[885,971,584,608]" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="8" pageNumber="581">ZMUT</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>