treatments-xml/data/03/B0/87/03B087E55B1CFFE9FF79F673FD23FD3E.xml
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<document id="EA9134628AF79875F034EF130D1B7C92" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B1CFFE9FF79F673FD23FD3E" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena pectenata Perkins 2011, new species" docType="treatment" docVersion="4" lastPageNumber="123" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="122" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C43C556D299D0825D5A43151A11F292E">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="EB3EFB0EE73BE8580191BDFC1FF612DE">Perkins, Philip D.</mods:namePart>
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<mods:date id="9047CC792D3A4564DEEFE58D32AC7125">2011</mods:date>
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<mods:classification id="BAAFA41EF804A0CB083D2F886F7C0C24">journal article</mods:classification>
<mods:identifier id="6AC429A913780E80609F15772FBD1962" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
<mods:identifier id="3F5D92FEB1E4AD47547CA7C5B003A7D0" type="ISSN">1175­5334</mods:identifier>
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<treatment id="03B087E55B1CFFE9FF79F673FD23FD3E" LSID="urn:lsid:plazi:treatment:03B087E55B1CFFE9FF79F673FD23FD3E" httpUri="http://treatment.plazi.org/id/03B087E55B1CFFE9FF79F673FD23FD3E" lastPageId="122" lastPageNumber="123" pageId="121" pageNumber="122">
<subSubSection id="C30365785B1CFFEAFF79F673FD9EFBEE" box="[151,552,1118,1144]" pageId="121" pageNumber="122" type="nomenclature">
<paragraph id="8BA636F35B1CFFEAFF79F673FD9EFBEE" blockId="121.[151,552,1118,1177]" box="[151,552,1118,1144]" pageId="121" pageNumber="122">
<heading id="D0EE819F5B1CFFEAFF79F673FD9EFBEE" bold="true" box="[151,552,1118,1144]" fontSize="11" level="1" pageId="121" pageNumber="122" reason="1">
<emphasis id="B96DEAE15B1CFFEAFF79F673FD9EFBEE" bold="true" box="[151,552,1118,1144]" pageId="121" pageNumber="122">
<taxonomicName id="4C194D705B1CFFEAFF79F673FE3FFBEE" ID-CoL="3N3F7" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,393,1118,1144]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="pectenata" status="sp. nov.">
<emphasis id="B96DEAE15B1CFFEAFF79F673FE3FFBEE" bold="true" box="[151,393,1118,1144]" italics="true" pageId="121" pageNumber="122">Hydraena pectenata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B1CFFEAFE76F673FD9EFBEE" box="[408,552,1118,1144]" pageId="121" pageNumber="122" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B1CFFEAFF79F6ACFE0CFB0F" box="[151,442,1153,1177]" pageId="121" pageNumber="122" type="description">
<paragraph id="8BA636F35B1CFFEAFF79F6ACFE0CFB0F" blockId="121.[151,552,1118,1177]" box="[151,442,1153,1177]" pageId="121" pageNumber="122">
(
<figureCitation id="13222A765B1CFFEAFF71F6ACFEBDFB0F" box="[159,267,1153,1177]" captionStart="FIGURE 223" captionStartId="312.[151,250,1753,1776]" captionTargetBox="[218,1365,213,1716]" captionTargetId="figure-28@312.[202,1384,193,1732]" captionTargetPageId="312" captionText="FIGURE 223. Hydraena pectenata, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283852" httpUri="https://zenodo.org/record/5283852/files/figure.png" pageId="121" pageNumber="122">Figs. 223</figureCitation>
,
<figureCitation id="13222A765B1CFFEAFEF8F6ACFEF4FB0F" box="[278,322,1153,1177]" captionStart="FIGURES 224225" captionStartId="313.[151,264,1650,1673]" captionText="FIGURES 224225. Aedeagi of holotypes. 224. H. ulna. 225. H. pectenata." pageId="121" pageNumber="122">225</figureCitation>
,
<figureCitation id="13222A765B1CFFEAFEA0F6ACFECCFB0F" box="[334,378,1153,1177]" captionStart="FIGURES 412415" captionStartId="375.[151,264,1650,1673]" captionTargetBox="[284,1308,216,1584]" captionTargetId="figure-25@375.[263,1323,193,1629]" captionTargetPageId="375" captionText="FIGURES 412415. Female tergite X, gonocoxite, and spermatheca of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284037" httpUri="https://zenodo.org/record/5284037/files/figure.png" pageId="121" pageNumber="122">413</figureCitation>
,
<figureCitation id="13222A765B1CFFEAFE68F6ACFE07FB0F" box="[390,433,1153,1177]" captionStart="FIGURES 535538" captionStartId="409.[151,264,1648,1671]" captionTargetBox="[202,1385,213,1621]" captionTargetId="figure-20@409.[202,1385,193,1627]" captionTargetPageId="409" captionText="FIGURES 535538. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284139" httpUri="https://zenodo.org/record/5284139/files/figure.png" pageId="121" pageNumber="122">536</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B1CFFEAFF79F6E9FCA3FAB2" pageId="121" pageNumber="122" type="materials_examined">
<paragraph id="8BA636F35B1CFFEAFF79F6E9FCA3FAB2" blockId="121.[151,1437,1219,2036]" pageId="121" pageNumber="122">
Type Material.
<materialsCitation id="3B713CAE5B1CFFEAFEBBF6E9FF58FAB2" collectingDate="1964-10-08" collectionCode="NHM" collectorName="M. E. Bacchus" county="Finisterre Mts." elevation="1067" latitude="-5.9166665" location="Damanti" longLatPrecision="1305" longitude="145.96666" municipality="Naho River Valley" pageId="121" pageNumber="122" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<typeStatus id="54A288515B1CFFEAFEBBF6E9FE74FB4A" box="[341,450,1220,1244]" pageId="121" pageNumber="122" type="holotype">
<emphasis id="B96DEAE15B1CFFEAFEBBF6E9FE74FB4A" bold="true" box="[341,450,1220,1244]" pageId="121" pageNumber="122">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B1CFFEAFDDDF6E9FCA5FB4A" bold="true" box="[563,787,1219,1244]" pageId="121" pageNumber="122">
<collectingRegion id="49DDF8115B1CFFEAFDDDF6E9FCBBFB4A" box="[563,781,1219,1244]" country="Papua New Guinea" name="Madang" pageId="121" pageNumber="122">Madang Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62C74E7F5B1CFFEAFCC6F6E9FC65FB4D" box="[808,979,1220,1244]" pageId="121" pageNumber="122">Finisterre Mts.</collectingCounty>
,
<collectingMunicipality id="6BC2AC895B1CFFEAFC0CF6E9FB0CFB4A" box="[994,1210,1220,1244]" pageId="121" pageNumber="122">Naho River Valley</collectingMunicipality>
,
<location id="8EC660285B1CFFEAFB27F6E9FA9AFB4A" LSID="urn:lsid:plazi:treatment:03B087E55B1CFFE9FF79F673FD23FD3E:8EC660285B1CFFEAFB27F6E9FA9AFB4A" box="[1225,1324,1220,1244]" county="Finisterre Mts." latitude="-5.9166665" longLatPrecision="1305" longitude="145.96666" municipality="Naho River Valley" name="Damanti" pageId="121" pageNumber="122" stateProvince="Madang">Damanti</location>
, ex small pool, probably permanent, in clay in dense shade,
<quantity id="4CE19B165B1CFFEAFD28F6CAFCABFB69" box="[710,797,1255,1279]" metricMagnitude="3" metricUnit="m" metricValue="1.067" pageId="121" pageNumber="122" unit="m" value="1067.0">
<elevation id="0034D1C05B1CFFEAFD28F6CAFCABFB69" box="[710,797,1255,1279]" metricMagnitude="3" metricUnit="m" metricValue="1.067" pageId="121" pageNumber="122" unit="m" value="1067.0">1067 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B1CFFEAFCC7F6CAFC37FB69" box="[809,897,1255,1279]" degrees="5" direction="south" minutes="55" orientation="latitude" pageId="121" pageNumber="122" precision="925" value="-5.9166665">5° 55' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B1CFFEAFC60F6CAFBB2FB69" box="[910,1028,1255,1279]" degrees="145" direction="east" minutes="58" orientation="longitude" pageId="121" pageNumber="122" precision="925" value="145.96666">145° 58' E</geoCoordinate>
,
<date id="FFA710335B1CFFEAFBFEF6CAFBC2FB69" box="[1040,1140,1255,1279]" pageId="121" pageNumber="122" value="1964-10-08">
<collectingDate id="EFE3E9DB5B1CFFEAFBFEF6CAFBC2FB69" box="[1040,1140,1255,1279]" pageId="121" pageNumber="122" value="1964-10-08">8 x 1964</collectingDate>
</date>
,
<collectorName id="26EC53255B1CFFEAFB91F6C5FA95FB69" box="[1151,1315,1255,1279]" pageId="121" pageNumber="122">M. E. Bacchus</collectorName>
(MEB 39) (
<collectionCode id="ED08AE365B1CFFEAFF71F721FF50FAB2" box="[159,230,1292,1316]" pageId="121" pageNumber="122">NHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5B1CFFEAFF17F721FCA7FAB2" box="[249,785,1292,1316]" collectingDate="1964-10-08" collectionCode="NHM" collectorName="M. E. Bacchus" county="Finisterre Mts." elevation="1067" latitude="-5.9166665" location="Damanti" longLatPrecision="1305" longitude="145.96666" municipality="Naho River Valley" pageId="121" pageNumber="122" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<emphasis id="B96DEAE15B1CFFEAFF17F721FEC7FAB2" bold="true" box="[249,369,1292,1316]" pageId="121" pageNumber="122">
<typeStatus id="54A288515B1CFFEAFF17F721FEC7FAB2" box="[249,369,1292,1316]" pageId="121" pageNumber="122" type="paratype">Paratypes</typeStatus>
</emphasis>
: Same data as holotype (213
<collectionCode id="ED08AE365B1CFFEAFD2CF721FCBEFAB2" box="[706,776,1292,1316]" pageId="121" pageNumber="122">NHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B1CFFEAFF29F702FE17FA22" pageId="121" pageNumber="122" type="diagnosis">
<paragraph id="8BA636F35B1CFFEAFF29F702FE17FA22" blockId="121.[151,1437,1219,2036]" pageId="121" pageNumber="122">
<emphasis id="B96DEAE15B1CFFEAFF29F702FE64FADE" bold="true" box="[199,466,1327,1352]" pageId="121" pageNumber="122">Differential Diagnosis.</emphasis>
Similar in dorsal habitus and plaque ratios to
<taxonomicName id="4C194D705B1CFFEAFC3BF71CFB9CFADE" authorityName="Perkins" authorityYear="2011" box="[981,1066,1328,1352]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="ulna">
<emphasis id="B96DEAE15B1CFFEAFC3BF71CFC5FFADE" box="[981,1001,1329,1352]" italics="true" pageId="121" pageNumber="122">H</emphasis>
.
<emphasis id="B96DEAE15B1CFFEAFC16F71DFB9CFADE" box="[1016,1066,1328,1352]" italics="true" pageId="121" pageNumber="122">ulna</emphasis>
</taxonomicName>
(
<figureCitation id="13222A765B1CFFEAFBD6F702FB13FAD1" box="[1080,1189,1327,1351]" captionStart="FIGURE 222" captionStartId="311.[151,250,1753,1776]" captionTargetBox="[206,1388,209,1724]" captionTargetId="figure-26@311.[186,1401,193,1732]" captionTargetPageId="311" captionText="FIGURE 222. Hydraena ulna, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283850" httpUri="https://zenodo.org/record/5283850/files/figure.png" pageId="121" pageNumber="122">Figs. 222</figureCitation>
,
<figureCitation id="13222A765B1CFFEAFB5EF702FB68FAD1" box="[1200,1246,1327,1351]" captionStart="FIGURE 223" captionStartId="312.[151,250,1753,1776]" captionTargetBox="[218,1365,213,1716]" captionTargetId="figure-28@312.[202,1384,193,1732]" captionTargetPageId="312" captionText="FIGURE 223. Hydraena pectenata, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283852" httpUri="https://zenodo.org/record/5283852/files/figure.png" pageId="121" pageNumber="122">223</figureCitation>
); differing therefrom in slightly smaller size (ca. 1.46 vs.
<quantity id="4CE19B165B1CFFEAFD8BF779FD65FAFA" box="[613,723,1364,1388]" metricMagnitude="-3" metricUnit="m" metricValue="1.57" pageId="121" pageNumber="122" unit="mm" value="1.57">1. 57 mm</quantity>
), more densely punctate pronotum (discal interstices ca. 12 vs. 24 xpd). The male genitalia of the two species also show a relationship, while differing distinctively in several details (
<figureCitation id="13222A765B1CFFEAFF01F7B1FE22FA22" box="[239,404,1436,1460]" captionStart="FIGURES 224225" captionStartId="313.[151,264,1650,1673]" captionText="FIGURES 224225. Aedeagi of holotypes. 224. H. ulna. 225. H. pectenata." pageId="121" pageNumber="122">Figs. 224, 225</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B1CFFE9FF29F792FB19FD81" lastPageId="122" lastPageNumber="123" pageId="121" pageNumber="122" type="description">
<paragraph id="8BA636F35B1CFFEAFF29F792FF74F9FE" blockId="121.[151,1437,1219,2036]" pageId="121" pageNumber="122">
<emphasis id="B96DEAE15B1CFFEAFF29F792FEEEFA4E" bold="true" box="[199,344,1471,1496]" inLineHeading="true" pageId="121" pageNumber="122" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B1CFFEAFE42F792FDB8FA41" box="[428,526,1471,1495]" pageId="121" pageNumber="122" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.46/0.65; head 0.20/0.34; pronotum 0.40/0.50, PA 0.38, PB 0.46; elytra 0.87/0.65. Dorsum of head dark brown to piceous; pronotum dark brown in front of and behind piceous, very diffusely margined, fascia, ratios of color bands, as measured in midline, ca. 7/15/5; elytra dark brown; legs brown; maxillary palpi testaceous, tip not darker. Dorsum moderately shining.
</paragraph>
<paragraph id="8BA636F35B1CFFEAFF29F459FE67F83A" blockId="121.[151,1437,1219,2036]" pageId="121" pageNumber="122">Frons punctures ca. 1xef near eyes, smaller and sparser medially; interstices 14xpd, microreticulate and dull laterally, smooth and shining medially. Clypeus effacedly microreticulate laterally, very finely sparsely punctulate and shining medially. Mentum effacedly microreticulate, very sparsely very finely punctulate, weakly shining. Postmentum microreticulate in shallow median depression, otherwise finely sparsely punctate. Genae very slightly raised, shining, with shallow, obsolete median impression, without posterior ridge. Pronotum transverse, anterior margin straight, sides straight or very slightly arcuate between anterior angle and widest part, arcuate at middle, sinuate between widest part and posterior angle; punctures on disc ca. 23xpd largest frons punctures, interstices shining, ca. 12xpd, punctures anteriorly and posteriorly ca. 2xpd those on disc; PF1, PF2 and PF4 very shallow; PF3 moderately deep, wide.</paragraph>
<paragraph id="8BA636F35B1CFFE9FF29F595FD24FF61" blockId="121.[151,1437,1219,2036]" lastBlockId="122.[151,1436,151,680]" lastPageId="122" lastPageNumber="123" pageId="121" pageNumber="122">Elytra widest at about midlength; summit of posterior declivity at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures slightly smaller than largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 12xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BA636F35B1FFFE9FF29F329FB19FD81" blockId="122.[151,1436,151,680]" pageId="122" pageNumber="123">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 3/2/8/3. P1 laminate; median carina sinuate in profile, slightly raised behind coxae. P2 length/width ca. 5/3, sides very slightly diverging toward blunt, slightly raised apex. Plaques flat or slightly rounded, at sloping sides of median depression, anteriorly tapering and convergent, narrowly separated at anterior extreme. Metaventrite flat between mesocoxae. AIS width at straight posterior margin ca. 1.5x P2. All legs of moderate length. Profemur (male) with small tubercle on ventral margin near trochanter; protibia straight, medial margin slightly widened in distal 1/2. Meso- and metatibia straight, slender. Abdominal apex very slightly asymmetrical; last tergite (male) with very small apicomedian notch. Aedeagus as illustrated (
<figureCitation id="13222A765B1FFFE9FEF8F3D2FECFFD81" box="[278,377,511,535]" captionStart="FIGURES 224225" captionStartId="313.[151,264,1650,1673]" captionText="FIGURES 224225. Aedeagi of holotypes. 224. H. ulna. 225. H. pectenata." pageId="122" pageNumber="123">Fig. 225</figureCitation>
). Female tergite X, gonocoxite, and spermatheca as illustrated (
<figureCitation id="13222A765B1FFFE9FBAEF3D2FB14FD81" box="[1088,1186,511,535]" captionStart="FIGURES 412415" captionStartId="375.[151,264,1650,1673]" captionTargetBox="[284,1308,216,1584]" captionTargetId="figure-25@375.[263,1323,193,1629]" captionTargetPageId="375" captionText="FIGURES 412415. Female tergite X, gonocoxite, and spermatheca of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284037" httpUri="https://zenodo.org/record/5284037/files/figure.png" pageId="122" pageNumber="123">Fig. 413</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B1FFFE9FF29F009FBEEFDC9" pageId="122" pageNumber="123" type="etymology">
<paragraph id="8BA636F35B1FFFE9FF29F009FBEEFDC9" blockId="122.[151,1436,151,680]" pageId="122" pageNumber="123">
<emphasis id="B96DEAE15B1FFFE9FF29F009FEFBFDAA" bold="true" box="[199,333,548,572]" inLineHeading="true" pageId="122" pageNumber="123" reason="1">Etymology.</emphasis>
&quot;Comb-like qualities&quot;; named in reference to the rows of spines on the meso- and metatibiae of males, the slender processes of the aedeagus, and the dentate tenth tergum of females.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B1FFFE9FF29F046FD23FD3E" pageId="122" pageNumber="123" type="distribution">
<paragraph id="8BA636F35B1FFFE9FF29F046FD23FD3E" blockId="122.[151,1436,151,680]" pageId="122" pageNumber="123">
<emphasis id="B96DEAE15B1FFFE9FF29F046FED6FD12" bold="true" box="[199,352,619,644]" inLineHeading="true" pageId="122" pageNumber="123" reason="1">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54A288515B1FFFE9FD24F041FD4CFD12" box="[714,762,620,644]" pageId="122" pageNumber="123">type</typeStatus>
locality, Damanti, in the Finisterre Mts. (Area 5), at the bor- der with Area 3; elevation
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(
<figureCitation id="13222A765B1FFFE9FDC8F0A2FD30FD31" box="[550,646,655,679]" captionStart="FIGURES 535538" captionStartId="409.[151,264,1648,1671]" captionTargetBox="[202,1385,213,1621]" captionTargetId="figure-20@409.[202,1385,193,1627]" captionTargetPageId="409" captionText="FIGURES 535538. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284139" httpUri="https://zenodo.org/record/5284139/files/figure.png" pageId="122" pageNumber="123">Fig. 536</figureCitation>
).
</paragraph>
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</treatment>
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