171 lines
20 KiB
XML
171 lines
20 KiB
XML
<document id="D2CF6F5C25418056569B59AEDAE193E6" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B13FFE5FF79F3B9FEEDF901" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena koje Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="119" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="119" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D45719FF172EF77189EC2F9C882F4504">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="B915EA1361038B23C9BEF609E0FD1542">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="7D853642282E4375128EF631DEF7B09B">text</mods:typeOfResource>
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<mods:title id="67D75767026348B845FE5AE32BCA271C">Zootaxa</mods:title>
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<mods:part id="C97462CE846515875952BDEBCF178BCF">
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<mods:date id="972E40A2DB9AF38BFEFD4FCCB836B3EE">2011</mods:date>
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<mods:number id="DA67FF0E75349FF093AD98C4B6F5E888">2011-07-08</mods:number>
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<mods:number id="A8C207E44AE6AA7C11FCB4804A478BC4">2944</mods:number>
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<mods:url id="937F56BB98B5D6831676451F81DFA1D0">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
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<mods:classification id="87CBF1A67B9389BC41BEE0543839566C">journal article</mods:classification>
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<mods:identifier id="A5D13BA9515CE7E67A94684B030FAA3E" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
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<mods:identifier id="2534394512CED783739F80A5638F40D8" type="ISSN">11755334</mods:identifier>
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<treatment id="03B087E55B13FFE5FF79F3B9FEEDF901" ID-DOI="http://doi.org/10.5281/zenodo.5291757" ID-Zenodo-Dep="5291757" LSID="urn:lsid:plazi:treatment:03B087E55B13FFE5FF79F3B9FEEDF901" httpUri="http://treatment.plazi.org/id/03B087E55B13FFE5FF79F3B9FEEDF901" lastPageNumber="119" pageId="118" pageNumber="119">
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<subSubSection id="C30365785B13FFE5FF79F3B9FE51FE3B" box="[151,487,403,430]" pageId="118" pageNumber="119" type="nomenclature">
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<paragraph id="8BA636F35B13FFE5FF79F3B9FE51FE3B" blockId="118.[151,487,403,462]" box="[151,487,403,430]" pageId="118" pageNumber="119">
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<heading id="D0EE819F5B13FFE5FF79F3B9FE51FE3B" bold="true" box="[151,487,403,430]" fontSize="11" level="1" pageId="118" pageNumber="119" reason="1">
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<emphasis id="B96DEAE15B13FFE5FF79F3B9FE51FE3B" bold="true" box="[151,487,403,430]" pageId="118" pageNumber="119">
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<taxonomicName id="4C194D705B13FFE5FF79F3B9FEFEFE38" ID-CoL="6MJB9" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,328,404,430]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="koje" status="sp. nov.">
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<emphasis id="B96DEAE15B13FFE5FF79F3B9FEFEFE38" bold="true" box="[151,328,404,430]" italics="true" pageId="118" pageNumber="119">Hydraena koje</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25E579A5B13FFE5FEB6F3BEFE51FE3B" box="[344,487,403,429]" pageId="118" pageNumber="119" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B13FFE5FF79F39BFE34FE58" box="[151,386,438,462]" pageId="118" pageNumber="119" type="description">
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<paragraph id="8BA636F35B13FFE5FF79F39BFE34FE58" blockId="118.[151,487,403,462]" box="[151,386,438,462]" pageId="118" pageNumber="119">
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(
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<figureCitation id="13222A765B13FFE5FF71F39BFEBDFE58" box="[159,267,438,462]" captionStart="FIGURE 215" captionStartId="306.[151,250,1753,1776]" captionTargetBox="[213,1377,220,1718]" captionTargetId="figure-29@306.[192,1395,193,1732]" captionTargetPageId="306" captionText="FIGURE 215. Hydraena koje, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283836" httpUri="https://zenodo.org/record/5283836/files/figure.png" pageId="118" pageNumber="119">Figs. 215</figureCitation>
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,
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<figureCitation id="13222A765B13FFE5FEF8F39BFEF4FE58" box="[278,322,438,462]" captionStart="FIGURES 216–217" captionStartId="307.[151,264,1650,1673]" captionText="FIGURES 216–217. Aedeagi of holotypes. 216. H. torosopala. 217. H. koje." pageId="118" pageNumber="119">217</figureCitation>
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,
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<figureCitation id="13222A765B13FFE5FEA0F39BFECFFE58" box="[334,377,438,462]" captionStart="FIGURES 527–530" captionStartId="407.[151,264,1650,1673]" captionTargetBox="[165,1421,224,1616]" captionTargetId="figure-20@407.[165,1421,193,1629]" captionTargetPageId="407" captionText="FIGURES 527–530. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284135" httpUri="https://zenodo.org/record/5284135/files/figure.png" pageId="118" pageNumber="119">528</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B13FFE5FF79F3D2FDA8FD14" pageId="118" pageNumber="119" type="materials_examined">
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<paragraph id="8BA636F35B13FFE5FF79F3D2FDA8FD14" blockId="118.[151,1437,510,1687]" pageId="118" pageNumber="119">
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Type Material.
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<materialsCitation id="3B713CAE5B13FFE5FEBEF3D2FAA4FDC8" collectingDate="1988-07-01" collectingDateMax="1988-08-30" collectingDateMin="1988-07-01" collectionCode="NMW" collectorName="D. Dudgeon" elevation="160" latitude="-5.55" location="Koje Creek" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" pageId="118" pageNumber="119" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
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<typeStatus id="54A288515B13FFE5FEBEF3D2FE0BFD81" box="[336,445,511,535]" pageId="118" pageNumber="119" type="holotype">
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<emphasis id="B96DEAE15B13FFE5FEBEF3D2FE0BFD81" bold="true" box="[336,445,511,535]" pageId="118" pageNumber="119">Holotype</emphasis>
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</typeStatus>
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(male):
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<emphasis id="B96DEAE15B13FFE5FDCBF3D2FCB5FD81" bold="true" box="[549,771,510,535]" pageId="118" pageNumber="119">
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<collectingRegion id="49DDF8115B13FFE5FDCBF3D2FD48FD81" box="[549,766,510,535]" country="Papua New Guinea" name="Madang" pageId="118" pageNumber="119">Madang Province</collectingRegion>
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:
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</emphasis>
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<collectingRegion id="49DDF8115B13FFE5FCFAF3D3FCE5FD80" box="[788,851,510,534]" country="Papua New Guinea" name="West Sepik" pageId="118" pageNumber="119">Sepik</collectingRegion>
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<collectingMunicipality id="6BC2AC895B13FFE5FCB5F3D2FC5FFD80" box="[859,1001,510,534]" pageId="118" pageNumber="119">Ramu Basin</collectingMunicipality>
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,
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<location id="8EC660285B13FFE5FC1BF3D3FBCFFD80" LSID="urn:lsid:plazi:treatment:03B087E55B13FFE5FF79F3B9FEEDF901:8EC660285B13FFE5FC1BF3D3FBCFFD80" box="[1013,1145,510,534]" latitude="-5.55" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" name="Koje Creek" pageId="118" pageNumber="119" stateProvince="Madang">Kojé Creek</location>
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, unshaded creek, small to medium cobbles, gravel and fine sand (the latter in areas of slack water), filamentous algae on stony substrates, small accumulations of leaf litter,
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<quantity id="4CE19B165B13FFE5FDFDF06BFDEAFDC8" box="[531,604,582,606]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="118" pageNumber="119" unit="m" value="160.0">
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<elevation id="0034D1C05B13FFE5FDFDF06BFDEAFDC8" box="[531,604,582,606]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="118" pageNumber="119" unit="m" value="160.0">160 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE2D50345B13FFE5FD86F06BFD77FDC8" box="[616,705,582,606]" degrees="5" direction="south" minutes="33" orientation="latitude" pageId="118" pageNumber="119" precision="925" value="-5.55">5° 33' S</geoCoordinate>
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,
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<geoCoordinate id="EE2D50345B13FFE5FD23F06BFCF2FDC8" box="[717,836,582,606]" degrees="145" direction="east" minutes="23" orientation="longitude" pageId="118" pageNumber="119" precision="925" value="145.38333">145° 23' E</geoCoordinate>
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,
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<date id="FFA710335B13FFE5FCBEF06BFBAFFDC8" box="[848,1049,582,606]" pageId="118" pageNumber="119" value="1988-07-01" valueMax="1988-08-30" valueMin="1988-07-01">
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<collectingDate id="EFE3E9DB5B13FFE5FCBEF06BFBAFFDC8" box="[848,1049,582,606]" pageId="118" pageNumber="119" value="1988-07-01" valueMax="1988-08-30" valueMin="1988-07-01">1 vii–30 viii 1988</collectingDate>
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</date>
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,
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<collectorName id="26EC53255B13FFE5FBCAF06AFB1BFDC8" box="[1060,1197,582,606]" pageId="118" pageNumber="119">D. Dudgeon</collectorName>
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(
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<collectionCode id="ED08AE365B13FFE5FB52F06BFABCFDC8" box="[1212,1290,582,606]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="118" pageNumber="119">NMW</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B713CAE5B13FFE5FAF2F06AFDACFD14" collectingDate="1988-07-01" collectingDateMax="1988-08-30" collectingDateMin="1988-07-01" collectionCode="NMW" collectorName="D. Dudgeon" elevation="160" latitude="-5.55" location="Koje Creek" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" pageId="118" pageNumber="119" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
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<emphasis id="B96DEAE15B13FFE5FAF2F06AFA22FDC9" bold="true" box="[1308,1428,583,607]" pageId="118" pageNumber="119">
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<typeStatus id="54A288515B13FFE5FAF2F06AFA22FDC9" box="[1308,1428,583,607]" pageId="118" pageNumber="119" type="paratype">Paratypes</typeStatus>
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</emphasis>
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: Same data as holotype (98
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<collectionCode id="ED08AE365B13FFE5FE2BF047FDA4FD14" box="[453,530,618,642]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="118" pageNumber="119">NMW</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B13FFE5FF29F0A3FEE7FCA0" pageId="118" pageNumber="119" type="diagnosis">
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<paragraph id="8BA636F35B13FFE5FF29F0A3FEE7FCA0" blockId="118.[151,1437,510,1687]" pageId="118" pageNumber="119">
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<emphasis id="B96DEAE15B13FFE5FF29F0A3FE64FD31" bold="true" box="[199,466,654,679]" pageId="118" pageNumber="119">Differential Diagnosis.</emphasis>
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Similar in small size (ca.
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<quantity id="4CE19B165B13FFE5FD18F0A3FCECFD30" box="[758,858,654,678]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="118" pageNumber="119" unit="mm" value="1.29">1.29 mm</quantity>
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) and color to
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<taxonomicName id="4C194D705B13FFE5FC18F0A2FBC5FD30" authorityName="Perkins" authorityYear="2011" box="[1014,1139,655,678]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="tritropis">
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<emphasis id="B96DEAE15B13FFE5FC18F0A2FBBCFD30" box="[1014,1034,655,678]" italics="true" pageId="118" pageNumber="119">H</emphasis>
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.
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<emphasis id="B96DEAE15B13FFE5FBF6F0A2FBC5FD30" box="[1048,1139,655,678]" italics="true" pageId="118" pageNumber="119">tritropis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13222A765B13FFE5FB6CF0A3FB5AFD30" box="[1154,1260,654,678]" captionStart="FIGURE 211" captionStartId="303.[151,250,1753,1776]" captionTargetBox="[261,1342,226,1718]" captionTargetId="figure-26@303.[228,1360,193,1732]" captionTargetPageId="303" captionText="FIGURE 211. Hydraena tritropis, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283828" httpUri="https://zenodo.org/record/5283828/files/figure.png" pageId="118" pageNumber="119">Figs. 211</figureCitation>
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,
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<figureCitation id="13222A765B13FFE5FB16F0A3FA90FD30" box="[1272,1318,654,678]" captionStart="FIGURE 215" captionStartId="306.[151,250,1753,1776]" captionTargetBox="[213,1377,220,1718]" captionTargetId="figure-29@306.[192,1395,193,1732]" captionTargetPageId="306" captionText="FIGURE 215. Hydraena koje, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283836" httpUri="https://zenodo.org/record/5283836/files/figure.png" pageId="118" pageNumber="119">215</figureCitation>
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); differing therefrom by the narrower body, the proportionally smaller pronotum (PW/EL ca. 0.59 vs. 0.66), the concave prosternal intercoxal process, and the narrower mesoventral intercoxal process (plaque ratios ca. 6/1/5/5 vs. 7/1/5/5). The male genitalia of the two species show some similarity in basic plan, but distinctively differ in many details (
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<figureCitation id="13222A765B13FFE5FF71F133FEBDFCA0" box="[159,267,798,822]" captionStart="FIGURES 212–213" captionStartId="304.[151,264,1650,1673]" captionTargetBox="[250,1326,204,1607]" captionTargetId="figure-32@304.[226,1360,193,1629]" captionTargetPageId="304" captionText="FIGURES 212–213. Aedeagi of holotypes. 212. H. formosopala. 213. H. tritropis." figureDoi="http://doi.org/10.5281/zenodo.5283830" httpUri="https://zenodo.org/record/5283830/files/figure.png" pageId="118" pageNumber="119">Figs. 213</figureCitation>
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,
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<figureCitation id="13222A765B13FFE5FEF8F133FEF2FCA0" box="[278,324,798,822]" captionStart="FIGURES 216–217" captionStartId="307.[151,264,1650,1673]" captionText="FIGURES 216–217. Aedeagi of holotypes. 216. H. torosopala. 217. H. koje." pageId="118" pageNumber="119">217</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B13FFE5FF29F16FFAEFF9BC" pageId="118" pageNumber="119" type="description">
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<paragraph id="8BA636F35B13FFE5FF29F16FFE03FC50" blockId="118.[151,1437,510,1687]" pageId="118" pageNumber="119">
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||
<emphasis id="B96DEAE15B13FFE5FF29F16FFEEEFCCD" bold="true" box="[199,344,834,859]" inLineHeading="true" pageId="118" pageNumber="119" reason="1">Description.</emphasis>
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Size:
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<typeStatus id="54A288515B13FFE5FE42F16FFDB8FCCC" box="[428,526,834,858]" pageId="118" pageNumber="119" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.29/0.57; head 0.21/0.32; pronotum 0.30/0.46, PA 0.35, PB 0.37; elytra 0.78/0.57. Dorsum generally reddish brown, frons and elytra distinctly more darkly colored than pronotum and clypeus; legs light brown to light reddish brown; maxillary palpi testaceous, tip not darker.
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</paragraph>
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<paragraph id="8BA636F35B13FFE5FF29F1FEFED7FB08" blockId="118.[151,1437,510,1687]" pageId="118" pageNumber="119">Frons punctures ca. 1–2xef, larger near eyes than medially; interstices shining, 1–4xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum and postmentum very sparsely very finely punctulate, shining. Genae raised, shining, without posterior ridge. Pronotum subcordiform, ca. median 3/4 of anterior margin arcuate to posterior; punctures on disc ca. 1xpd frons punctures, interstices shining, 1–3xpd, punctures slightly larger and denser at anterior and posterior; PF1 very shallow, obsolete; PF2 very shallow; PF3 moderately deep; PF4 very shallow.</paragraph>
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<paragraph id="8BA636F35B13FFE5FF29F687FE78FAC4" blockId="118.[151,1437,510,1687]" pageId="118" pageNumber="119">Elytra with summit of posterior declivity slightly anterior to midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 1–3xpd, as are interstices between punctures of a row. Apices in dorsal aspect slightly separately rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
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<paragraph id="8BA636F35B13FFE5FF29F773FAEFF9BC" blockId="118.[151,1437,510,1687]" pageId="118" pageNumber="119">
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Ratios of P2 width and plaque shape (P2/w/l/s) ca. 6/1/5/5. P1 ca. 2/3 P2; median carina straight in profile; postcoxal process medially concave. P2 transversely concave, l/w ca. 5/6, sides slightly diverging toward blunt apex. Plaques very narrow, carinate, converging slightly anteriorly, at sides of deep median depression. Metaventrite concave between P2 and plaques. AIS width at straight posterior margin ca. equal P2. All legs moderately long and slender. Profemur (male) without tubercle next to trochanter; protibia slightly arcuate. Meso- and metatibia straight. Abdominal apex symmetrical; last tergite (male) deeply notched. Aedeagus as illustrated (
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<figureCitation id="13222A765B13FFE5FB04F43FFAFAF9BC" box="[1258,1356,1554,1578]" captionStart="FIGURES 216–217" captionStartId="307.[151,264,1650,1673]" captionText="FIGURES 216–217. Aedeagi of holotypes. 216. H. torosopala. 217. H. koje." pageId="118" pageNumber="119">Fig. 217</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B13FFE5FF29F41AFCB9F9D8" box="[199,783,1590,1615]" pageId="118" pageNumber="119" type="etymology">
|
||
<paragraph id="8BA636F35B13FFE5FF29F41AFCB9F9D8" blockId="118.[151,1437,510,1687]" box="[199,783,1590,1615]" pageId="118" pageNumber="119">
|
||
<emphasis id="B96DEAE15B13FFE5FF29F41AFEFBF9D9" bold="true" box="[199,333,1591,1615]" inLineHeading="true" pageId="118" pageNumber="119" reason="1">Etymology.</emphasis>
|
||
Named in reference to the
|
||
<typeStatus id="54A288515B13FFE5FD90F41AFD18F9D9" box="[638,686,1591,1615]" pageId="118" pageNumber="119">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B13FFE5FF29F477FEEDF901" pageId="118" pageNumber="119" type="distribution">
|
||
<paragraph id="8BA636F35B13FFE5FF29F477FEEDF901" blockId="118.[151,1437,510,1687]" pageId="118" pageNumber="119">
|
||
<emphasis id="B96DEAE15B13FFE5FF29F477FED6F9E5" bold="true" box="[199,352,1626,1651]" inLineHeading="true" pageId="118" pageNumber="119" reason="1">Distribution.</emphasis>
|
||
Currently known only from the
|
||
<typeStatus id="54A288515B13FFE5FD25F476FD4DF9E5" box="[715,763,1627,1651]" pageId="118" pageNumber="119">type</typeStatus>
|
||
locality, Kojé Creek, in the eastern arm of Area 3; elevation
|
||
<quantity id="4CE19B165B13FFE5FF79F453FF68F900" box="[151,222,1662,1686]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="118" pageNumber="119" unit="m" value="160.0">160 m</quantity>
|
||
(
|
||
<figureCitation id="13222A765B13FFE5FF03F453FEFAF900" box="[237,332,1662,1686]" captionStart="FIGURES 527–530" captionStartId="407.[151,264,1650,1673]" captionTargetBox="[165,1421,224,1616]" captionTargetId="figure-20@407.[165,1421,193,1629]" captionTargetPageId="407" captionText="FIGURES 527–530. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284135" httpUri="https://zenodo.org/record/5284135/files/figure.png" pageId="118" pageNumber="119">Fig. 528</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |