treatments-xml/data/03/B0/87/03B087CCF260FFACFF390CBAAC321BAD.xml

<document id="DE03506A9E0DF50D2AD3FFD92661C80E" ID-CLB-Dataset="32535" ID-DOI="10.11646/zootaxa.4303.1.1" ID-GBIF-Dataset="958ff34b-f5e0-4e8f-903f-2b4473bb69b3" ID-ISSN="1175-5326" ID-Zenodo-Dep="840417" ID-ZooBank="9E9C21C7-AB32-4A65-A2C0-1A0B826F5D64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1502221956248" checkinUser="plazi" docAuthor="Rabosky, Daniel L., Doughty, Paul &amp; Huang, Huateng" docDate="2017" docId="03B087CCF260FFACFF390CBAAC321BAD" docLanguage="en" docName="zootaxa.4303.1.1.pdf" docOrigin="Zootaxa 4303 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ctenotus piankai subsp. duricola , Storr 1975" docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="FF89FFB4F26FFFBDFFAE0E6CA9301C49" masterDocTitle="Lizards in pinstripes: morphological and genomic evidence for two new species of scincid lizards within Ctenotus piankai Storr and C. duricola Storr (Reptilia: Scincidae) in the Australian arid zone" masterLastPageNumber="26" masterPageNumber="1" pageNumber="16" updateTime="1698466361088" updateUser="ExternalLinkService">
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<mods:title id="C1570C54FDD95D6172810ADE1CD010E5">Lizards in pinstripes: morphological and genomic evidence for two new species of scincid lizards within Ctenotus piankai Storr and C. duricola Storr (Reptilia: Scincidae) in the Australian arid zone</mods:title>
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<mods:namePart id="9E3C6F08484E1C156849C339129F0C06">Rabosky, Daniel L.</mods:namePart>
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<mods:namePart id="2D447B5D6388E28049887C2139CD93E9">Doughty, Paul</mods:namePart>
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<mods:namePart id="470D046F93BC865A5754D695F59FA9AB">Huang, Huateng</mods:namePart>
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<paragraph id="8BA636DAF260FFB2FF390CBAABF31EB9" blockId="15.[151,707,726,822]" box="[151,707,726,752]" pageId="15" pageNumber="16">
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<emphasis id="B96DEAC8F260FFB2FF390CBAABF31EB9" bold="true" box="[151,707,726,752]" pageId="15" pageNumber="16">
<taxonomicName id="4C194D59F260FFB2FF390CBAAB541EB9" authority="Storr, 1975" authorityName=", Storr" authorityYear="1975" box="[151,612,726,752]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="subSpecies" species="piankai" subSpecies="duricola">
<emphasis id="B96DEAC8F260FFB2FF390CBAA8E11EB9" bold="true" box="[151,465,726,752]" italics="true" pageId="15" pageNumber="16">Ctenotus piankai duricola</emphasis>
<bibRefCitation id="EF884B2BF260FFB2FE760CBAAB541EB9" author="Storr" box="[472,612,726,752]" pageId="15" pageNumber="16" refString="Storr, G. M. (1975) The genus Ctenotus (Lacertilia: Scincidae) in the Kimberley and North-west Divisions of Western Australia. Records of the Western Australian Museum, 3, 209 - 243." type="journal article" year="1975">Storr, 1975</bibRefCitation>
</taxonomicName>
: p. 239.
</emphasis>
</heading>
</paragraph>
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Eastern Pilbara Lined 
<taxonomicName id="4C194D59F260FFB2FE3E0C96A8C41F5B" authorityName="Storr" authorityYear="1964" box="[400,500,762,786]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">Ctenotus</taxonomicName>
</paragraph>
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(
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, 
<figureCitation id="13222A5FF260FFB2FF540D72A8381F7F" box="[250,264,798,822]" captionStart="FIGURE 3" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[187,1402,193,1921]" captionTargetId="figure@5.[187,1402,193,1921]" captionTargetPageId="5" captionText="FIGURE 3. Lateral-dorsal perspective of the forebodies of Ctenotus. From top to bottom: A) C. piankai (WAM R 108934), B) C. rhabdotus sp. nov. (WAM R 110581), C) C. duricola (WAM R 111876), and D) C. pallasotus sp. nov. (WAM R 119931). Scale bar is 10 mm." httpUri="https://zenodo.org/record/840425/files/figure.png" pageId="15" pageNumber="16">3</figureCitation>
C, 7, 14)
</paragraph>
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<typeStatus id="54A28878F260FFB2FF390D09A8361F37" box="[151,262,869,894]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
.
</emphasis>
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R17163 (female), collected from 
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, 
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(
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, 
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) by 
<collectorName id="26EC530CF260FFB2FAA60D0AAC4E1F37" box="[1288,1406,870,894]" pageId="15" pageNumber="16">G.M. Storr</collectorName>
&amp; 
<collectorName id="26EC530CF260FFB2FF390DE6A8321FEB" box="[151,258,905,930]" pageId="15" pageNumber="16">B.T. Clay</collectorName>
on 
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<date id="FFA7101AF260FFB2FE830DE5A8C71FEB" box="[301,503,905,930]" pageId="15" pageNumber="16" value="1961-02-15">15 February 1961</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
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<paragraph id="8BA636DAF260FFB2FF690DC1AC0F1FA3" blockId="15.[151,1437,869,1866]" pageId="15" pageNumber="16">
<emphasis id="B96DEAC8F260FFB2FF690DC1AB391F8F" bold="true" box="[199,521,941,966]" pageId="15" pageNumber="16">
Measurements of 
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</emphasis>
: SVL 
<quantity id="4CE19B3FF260FFB2FDF50DC1AB801F8C" box="[603,688,941,966]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="15" pageNumber="16" unit="mm" value="54.0">54 mm</quantity>
; TL 
<quantity id="4CE19B3FF260FFB2FD420DC2AA0E1F8C" box="[748,830,942,966]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="16" unit="mm" value="18.0">18 mm</quantity>
(broken); HL 
<quantity id="4CE19B3FF260FFB2FC450DC1AD651F8C" box="[1003,1109,941,966]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="15" pageNumber="16" unit="mm" value="10.6">10.6 mm</quantity>
; HW 
<quantity id="4CE19B3FF260FFB2FB330DC2ADC91F8C" box="[1181,1273,942,966]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="15" pageNumber="16" unit="mm" value="7.2">7.2 mm</quantity>
; HD 
<quantity id="4CE19B3FF260FFB2FA950DC1ACA81F8C" box="[1339,1432,941,966]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="15" pageNumber="16" unit="mm" value="6.5">6.5 mm</quantity>
; 4ToeL 
<quantity id="4CE19B3FF260FFB2FF480DBEA80F1FA0" box="[230,319,978,1002]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="15" pageNumber="16" unit="mm" value="7.8">7.8 mm</quantity>
; ToeLam 24; MBSR 28; SupLab 8; InfLab 8; supraciliaries 8; EarLob 3 (left) and 4 (right).
</paragraph>
<paragraph id="8BA636DAF260FFB2FF690D99AC0D181F" blockId="15.[151,1437,869,1866]" pageId="15" pageNumber="16">
<materialsCitation id="3B713C87F260FFB2FF690D99ABD41847" ID-GBIF-Occurrence="1585968301" box="[199,740,1013,1038]" collectionCode="WAM" location="Woodstock" pageId="15" pageNumber="16" specimenCount="1" typeStatus="paratype">
<emphasis id="B96DEAC8F260FFB2FF690D99A8761847" bold="true" box="[199,326,1013,1038]" pageId="15" pageNumber="16">
<typeStatus id="54A28878F260FFB2FF690D99A8711847" box="[199,321,1013,1038]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED08AE1FF260FFB2FEE20D9AA8A11847" box="[332,401,1014,1038]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="15" pageNumber="16">WAM</collectionCode>
R12629 (female), 
<location id="8EC66001F260FFB2FDCD0D99ABD41847" LSID="urn:lsid:plazi:treatment:03B087CCF260FFACFF390CBAAC321BAD:8EC66001F260FFB2FDCD0D99ABD41847" box="[611,740,1013,1038]" name="Woodstock" pageId="15" pageNumber="16">Woodstock</location>
</materialsCitation>
; 
<materialsCitation id="3B713C87F260FFB2FD400D9AAC7D1847" ID-GBIF-Occurrence="1585968306" box="[750,1357,1013,1038]" collectionCode="WAM" location="Tambrey" pageId="15" pageNumber="16" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED08AE1FF260FFB2FD400D9AAA021847" box="[750,818,1014,1038]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="15" pageNumber="16">WAM</collectionCode>
R20115* (field tag—231) (damaged), 
<location id="8EC66001F260FFB2FB480D99AC7D1847" LSID="urn:lsid:plazi:treatment:03B087CCF260FFACFF390CBAAC321BAD:8EC66001F260FFB2FB480D99AC7D1847" box="[1254,1357,1013,1038]" name="Tambrey" pageId="15" pageNumber="16">Tambrey</location>
</materialsCitation>
; 
<materialsCitation id="3B713C87F260FFB2FAF60D9AABFC187B" ID-GBIF-Occurrence="1585968299" collectionCode="WAM" location="Mt Herbert" pageId="15" pageNumber="16" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED08AE1FF260FFB2FAF60D9AACAD1847" box="[1368,1437,1014,1038]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="15" pageNumber="16">WAM</collectionCode>
R20117* (field tag—439) (juvenile), 
<location id="8EC66001F260FFB2FDE60A76ABFC187B" LSID="urn:lsid:plazi:treatment:03B087CCF260FFACFF390CBAAC321BAD:8EC66001F260FFB2FDE60A76ABFC187B" box="[584,716,1049,1074]" name="Mt Herbert" pageId="15" pageNumber="16">Mt Herbert</location>
</materialsCitation>
; 
<materialsCitation id="3B713C87F260FFB2FD750A76AC0D181F" ID-GBIF-Occurrence="1585968292" collectionCode="WAM" location="Barrow Island" pageId="15" pageNumber="16" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED08AE1FF260FFB2FD750A76AA10187B" box="[731,800,1050,1074]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="15" pageNumber="16">WAM</collectionCode>
R45362* (field tag—258) (female) &amp; 
<collectionCode id="ED08AE1FF260FFB2FB490A76AC1C187B" box="[1255,1324,1050,1074]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="15" pageNumber="16">WAM</collectionCode>
R45363* (male), 
<location id="8EC66001F260FFB2FF400A52A8BD181F" LSID="urn:lsid:plazi:treatment:03B087CCF260FFACFF390CBAAC321BAD:8EC66001F260FFB2FF400A52A8BD181F" box="[238,397,1085,1110]" name="Barrow Island" pageId="15" pageNumber="16">Barrow Island</location>
(* species = 
<taxonomicName id="4C194D59F260FFB2FD8F0A52AB85181F" authority="Rabosky &amp; Doughty &amp; Huang, 2017" authorityName="Rabosky &amp; Doughty &amp; Huang" authorityYear="2017" box="[545,693,1085,1110]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="pallasotus" status="sp. nov.">
<emphasis id="B96DEAC8F260FFB2FD8F0A52AB85181F" box="[545,693,1085,1110]" italics="true" pageId="15" pageNumber="16">C. pallasotus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B3F260FFB2FD120A51AA26181F" box="[700,790,1085,1110]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B96DEAC8F260FFB2FD120A51AA26181F" bold="true" box="[700,790,1085,1110]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
, but maintains status as 
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of 
<taxonomicName id="4C194D59F260FFB2FB030A52AC1C181F" authorityName="Storr" authorityYear="1975" box="[1197,1324,1085,1110]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="duricola">
<emphasis id="B96DEAC8F260FFB2FB030A52AC1C181F" box="[1197,1324,1085,1110]" italics="true" pageId="15" pageNumber="16">C. duricola</emphasis>
</taxonomicName>
).
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</paragraph>
</subSubSection>
<subSubSection id="C3036551F260FFB2FF690A0DACB3191B" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8BA636DAF260FFB2FF690A0DACB3191B" blockId="15.[151,1437,869,1866]" pageId="15" pageNumber="16">
<emphasis id="B96DEAC8F260FFB2FF690A0DA80E1833" bold="true" box="[199,318,1121,1146]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
A small-bodied (to 
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SVL), elongate 
<taxonomicName id="4C194D59F260FFB2FCB00A0EAA4F1830" authorityName="Storr" authorityYear="1964" box="[798,895,1122,1145]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B96DEAC8F260FFB2FCB00A0EAA4F1830" box="[798,895,1122,1145]" italics="true" pageId="15" pageNumber="16">Ctenotus</emphasis>
</taxonomicName>
, nasals in contact, prefrontals in contact or not in contact, 26–29 mid-body scale rows, 20–25 compressed lamellae under toes with narrow callus, usually eight supralabials and supraciliary scales; pattern simple and full including six (occasionally eight) pale narrow longitudinal stripes on a blackish-brown ground color, dorsal stripes not continuing on head to snout, tail not red or blue, absence of upper lateral row of spots between dorsolateral and lateral stripes, dark vertebral stripe continues to anterior-most nuchal or parietal, parietals without blotches or spots, pale dorsolateral stripe continues anteriorly to eye, pale paravertebral stripes join on tail anterior to level of heel of extended hindlimb, lower labial scales variably stippled.
</paragraph>
</subSubSection>
<subSubSection id="C3036551F260FFACFF690B31AC651D54" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
<paragraph id="8BA636DAF260FFB2FF690B31AA3A1A4F" blockId="15.[151,1437,869,1866]" pageId="15" pageNumber="16">
<emphasis id="B96DEAC8F260FFB2FF690B31A869193F" bold="true" box="[199,345,1373,1398]" pageId="15" pageNumber="16">Description.</emphasis>
Head triangular, with snout narrowing to rounded tip; body long and cylindrical with flattened ventrum, neck only weakly constricted from head and body, concavity posterior to forelimbs; limbs short and welldeveloped, palmar and plantar surfaces with raised conical to triangular scales with rounded tips, fingers and toes five, finger length: 3&gt;4&gt;2&gt;5&gt;1, toe length: 4&gt;3&gt;2&gt;5&gt;1, digits laterally compressed with obtuse keel, claws long and recurved; tail long and thin, tapering to a fine point.
</paragraph>
<paragraph id="8BA636DAF260FFB2FF69087DAA3A1A3B" blockId="15.[151,1437,869,1866]" pageId="15" pageNumber="16">Dorsal scales smooth, flat, reflective and imbricate, posterior edge of scale convex; scales on midline of dorsum widest, decreasing in size to ventrolateral edge; on tail, ventral scale rows along midline much wider than other scale rows, distal portion of tail often regenerated.</paragraph>
<paragraph id="8BA636DAF260FFB2FF690811ABA11B03" blockId="15.[151,1437,869,1866]" pageId="15" pageNumber="16">
Nostril located in center of nasal scale and directed anteriorly, frontoparietals divided, supraoculars four (2nd largest), supraciliaries 7–9, loreals two, preoculars two, presuboculars one, supralabials 7–9, infralabials 7–10, upper eyelid with translucent scales, lower eyelid scaly, ear opening D-shaped with 2–6 ear lobules on anterior edge, nuchals 2–7; mental with straight posterior edge, gulars imbricate decreasing in size posteriorly towards neck then increasing again on ventrum; two greatly enlarged scales anterior to cloaca. Variation in continuous and meristic characters is summarized in 
<tableCitation id="C69B0361F260FFB2FD99095DABBB1B03" box="[567,651,1841,1866]" captionStart="TABLE 1" captionStartId="17.[151,239,1088,1111]" captionText="TABLE 1. Morphological characteristics of the Ctenotus species considered in this study. Number in parentheses after taxon name is number of individuals measured. Definitions for each character are provided in the Materials and Methods. Summaries show means ± standard deviations for each character (range in parentheses). Measurements for continuous characters in mm." pageId="15" pageNumber="16">Table 1</tableCitation>
.
</paragraph>
<caption id="DF666652F27FFFADFF3908ABAD331A94" box="[151,1027,1735,1758]" httpUri="https://zenodo.org/record/840447/files/figure.png" pageId="16" pageNumber="17" targetBox="[159,1423,195,1715]" targetPageId="16">
<paragraph id="8BA636DAF27FFFADFF3908ABAD331A94" blockId="16.[151,1027,1735,1758]" box="[151,1027,1735,1758]" pageId="16" pageNumber="17">
<emphasis id="B96DEAC8F27FFFADFF3908ABA82F1A95" bold="true" box="[151,287,1735,1757]" pageId="16" pageNumber="17">FIGURE 14.</emphasis>
<taxonomicName id="4C194D59F27FFFADFE8B08A4A8EC1A94" authorityName="Storr" authorityYear="1975" box="[293,476,1735,1757]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="duricola">
<emphasis id="B96DEAC8F27FFFADFE8B08A4A8EC1A94" box="[293,476,1735,1757]" italics="true" pageId="16" pageNumber="17">Ctenotus duricola</emphasis>
</taxonomicName>
holotype (specimen from WAM). Scale bar is 10 mm.
</paragraph>
</caption>
<paragraph id="8BA636DAF27FFFACFF690966AC651D54" blockId="16.[151,1436,1802,2007]" lastBlockId="17.[151,1437,151,1040]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="B96DEAC8F27FFFADFF690966A87C1B68" bold="true" box="[199,332,1802,1827]" pageId="16" pageNumber="17">Coloration.</emphasis>
Simple pattern of light to medium blackish-brown background and six (occasionally eight) longitudinal stripes; on lighter specimens, vertebral stripe darker than other areas of background color; four pale dorsal stripes narrow (&lt;½ scale width), interrupted by encroachment of dark dorsal border of scale giving a dashed appearance; two lateral stripes wider (&gt;½ scale width) and solid, continuing forwards to ear and on to upper labials and below eye; paravertebral stripe usually continues anteriorly to eye; short pale lower lateral stripe below upper lateral stripe from below angle of jaw to forelimb; purplish-black eyes somewhat visible through medial supraocular scales; ear lobules pale; lower labials pale with variable stippling; all ventral surfaces pale and immaculate; ground color of legs, arms, and tail light reddish-brown to tan; arm with 4–6 faint alternating light and dark longitudinal stripes; leg with 6–8 contrasting light and dark stripes; on tail, pale paravertebral stripes usually join anterior to the level of the heel of extended leg; distal portion of tail tan on dorsal surface with stripes only on lateral surfaces.
</paragraph>
</subSubSection>
<subSubSection id="C3036551F27EFFACFF690F4BA8551DE4" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BA636DAF27EFFACFF690F4BABE61D2D" blockId="17.[151,1437,151,1040]" pageId="17" pageNumber="18">
<emphasis id="B96DEAC8F27EFFACFF690F4BA8501D09" bold="true" box="[199,352,295,320]" pageId="17" pageNumber="18">Distribution.</emphasis>
Eastern Pilbara region of 
<collectingRegion id="49DDF838F27EFFACFD320F44AA5C1D09" box="[668,876,295,320]" country="Australia" name="Western Australia" pageId="17" pageNumber="18">
Western 
<collectingCountry id="F30E764AF27EFFACFCAC0F4BAA5C1D09" box="[770,876,295,320]" name="Australia" pageId="17" pageNumber="18">Australia</collectingCountry>
</collectingRegion>
. North and west to near Whim Creek, south to Newman and east to Woodie Woodie mine (
<figureCitation id="13222A5FF27EFFACFD2A0F20ABF91D2D" box="[644,713,332,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,717,739]" captionTargetBox="[202,1404,205,695]" captionTargetId="figure@6.[187,1416,188,704]" captionTargetPageId="6" captionText="FIGURE 4. (A) Geographic distribution of morphological forms within the nominate species ‘ C. piankai ’ (blue triangles, gray circles) and ‘ C. duricola ’ (red circles, yellow triangles). Relative to C. piankai from the interior deserts (blue triangles), northern populations from the Kimberley, Ord, Victoria River and northern Tanami desert (gray circles) are characterized by darker ground coloration, more midbody scale rows, and dorsal stripes that extend anteriorly onto the parietal scales (Figs. 1, 3). (B) Distribution of ‘ clean’ (red circles) and ‘ spotted’ (yellow triangles) morphological forms within ‘ C. duricola ’ in the Pilbara region of Western Australia. The ‘ clean’ morphology (Figs. 2 A, B; Fig. 3 C) is largely restricted to the eastern Pilbara, although several individuals from the westernmost edge of the range of C. duricola (North West Cape and Barrow Island) were observed to have this morphology. Several geographic features of north-western Australia discussed in the text are labeled in Fig. 4 B: NWC, North West Cape peninsula; D, De Grey River; F: Fortescue River Basin, AR, Ashburton River." httpUri="https://zenodo.org/record/840427/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BA636DAF27EFFACFF690F03A8551DE4" blockId="17.[151,1437,151,1040]" pageId="17" pageNumber="18">
<emphasis id="B96DEAC8F27EFFACFF690F03A81A1DC1" bold="true" box="[199,298,367,392]" pageId="17" pageNumber="18">Habitat.</emphasis>
Occurs on rocky slopes with spinifex and associated colluvial surfaces in the Pilbara, including mulga woodlands.
</paragraph>
</subSubSection>
<subSubSection id="C3036551F27EFFACFF690FDBAC721DBC" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BA636DAF27EFFACFF690FDBAC721DBC" blockId="17.[151,1437,151,1040]" pageId="17" pageNumber="18">
<emphasis id="B96DEAC8F27EFFACFF690FDBA87E1D99" bold="true" box="[199,334,439,464]" pageId="17" pageNumber="18">Etymology.</emphasis>
The specific name is from the Latin 
<emphasis id="B96DEAC8F27EFFACFD470FDBAA191D99" box="[745,809,439,464]" italics="true" pageId="17" pageNumber="18">durus</emphasis>
, meaning hard, and the suffix 
<emphasis id="B96DEAC8F27EFFACFBD20FDBAD861D99" box="[1148,1206,439,464]" italics="true" pageId="17" pageNumber="18">-cola</emphasis>
, meaning to inhabit, in reference to this species occurrence and presumed preference for the hard surfaces of the Pilbara region.
</paragraph>
</subSubSection>
<subSubSection id="C3036551F27EFFACFF690F93AC5018AE" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BA636DAF27EFFACFF690F93AC261EB9" blockId="17.[151,1437,151,1040]" pageId="17" pageNumber="18">
<emphasis id="B96DEAC8F27EFFACFF690F93A8091E51" bold="true" box="[199,313,511,536]" pageId="17" pageNumber="18">Remarks.</emphasis>
Of the five paratypes listed by 
<bibRefCitation id="EF884B2BF27EFFACFD040C6CAA051E51" author="Storr" box="[682,821,511,536]" pageId="17" pageNumber="18" refString="Storr, G. M. (1975) The genus Ctenotus (Lacertilia: Scincidae) in the Kimberley and North-west Divisions of Western Australia. Records of the Western Australian Museum, 3, 209 - 243." type="journal article" year="1975">Storr (1975)</bibRefCitation>
, only WAM R12629 from Woodstock in the northern Pilbara is unambiguously a true 
<taxonomicName id="4C194D59F27EFFACFDAA0C49ABB31E74" authorityName="Storr" authorityYear="1975" box="[516,643,548,573]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="duricola">
<emphasis id="B96DEAC8F27EFFACFDAA0C49ABB31E74" box="[516,643,548,573]" italics="true" pageId="17" pageNumber="18">C. duricola</emphasis>
</taxonomicName>
with the clean morphology. The other four specimens we assign to 
<taxonomicName id="4C194D59F27EFFACFA2D0C49A8371E29" authority="Rabosky &amp; Doughty &amp; Huang, 2017" authorityName="Rabosky &amp; Doughty &amp; Huang" authorityYear="2017" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pallasotus" status="sp. nov.">
<emphasis id="B96DEAC8F27EFFACFA2D0C49A8371E29" italics="true" pageId="17" pageNumber="18">C. pallasotus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B3F27EFFACFEBE0C2BA8591E29" box="[272,361,583,608]" pageId="17" pageNumber="18" rank="species">
<emphasis id="B96DEAC8F27EFFACFEBE0C2BA8591E29" bold="true" box="[272,361,583,608]" pageId="17" pageNumber="18">sp. nov.</emphasis>
</taxonomicNameLabel>
: WAM R20115 from Tambery possesses the spotted morphology and occur at the eastern limit; WAM R20117 from Mt Herbert also occurs at the eastern limit and shows the spotted morphology (albeit weakly); for the two specimens from Barrow Island, WAM R45362 shows the clean morphology, whereas WAM R45363 has the spotted morphology. This raises the possibility that Barrow Island has both species, or that it only has 
<taxonomicName id="4C194D59F27EFFACFB480CD9AC451E84" authority="Rabosky &amp; Doughty &amp; Huang, 2017" authorityName="Rabosky &amp; Doughty &amp; Huang" authorityYear="2017" box="[1254,1397,692,717]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pallasotus" status="sp. nov.">
<emphasis id="B96DEAC8F27EFFACFB480CD9AC451E84" box="[1254,1397,692,717]" italics="true" pageId="17" pageNumber="18">C. pallasotus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B3F27EFFACFAD50CD8A9F71EB9" pageId="17" pageNumber="18" rank="species">
<emphasis id="B96DEAC8F27EFFACFAD50CD8A9F71EB9" bold="true" pageId="17" pageNumber="18">sp. nov.</emphasis>
</taxonomicNameLabel>
but with variable expression of the spotted morphology, similar to the North West Cape populations.
</paragraph>
<paragraph id="8BA636DAF27EFFACFF690C90ABC51859" blockId="17.[151,1437,151,1040]" pageId="17" pageNumber="18">
This species is now confined to the eastern Pilbara region. The pattern of allopatric replacement of 
<taxonomicName id="4C194D59F27EFFACFAB20C91ACAC1F5C" authorityName="Storr" authorityYear="1975" box="[1308,1436,764,789]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="duricola">
<emphasis id="B96DEAC8F27EFFACFAB20C91ACAC1F5C" box="[1308,1436,764,789]" italics="true" pageId="17" pageNumber="18">C. duricola</emphasis>
</taxonomicName>
with 
<taxonomicName id="4C194D59F27EFFACFF7E0D4CA8531F71" authority="Rabosky &amp; Doughty &amp; Huang, 2017" authorityName="Rabosky &amp; Doughty &amp; Huang" authorityYear="2017" box="[208,355,799,824]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pallasotus" status="sp. nov.">
<emphasis id="B96DEAC8F27EFFACFF7E0D4CA8531F71" box="[208,355,799,824]" italics="true" pageId="17" pageNumber="18">C. pallasotus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B3F27EFFACFEC50D73A8F41F71" box="[363,452,799,824]" pageId="17" pageNumber="18" rank="species">
<emphasis id="B96DEAC8F27EFFACFEC50D73A8F41F71" bold="true" box="[363,452,799,824]" pageId="17" pageNumber="18">sp. nov.</emphasis>
</taxonomicNameLabel>
within the Pilbara region corresponds to the supposition of 
<bibRefCitation id="EF884B2BF27EFFACFBCD0D4CAC711F71" box="[1123,1345,799,824]" pageId="17" pageNumber="18" refString="Pepper, M., Doughty, P. &amp; Keogh, S. J. (2013) Synthesis: Geodiversity and endemism in the iconic Australian Pilbara region: a review of landscape evolution and biotic response in an ancient refugium. Journal of Biogeography, 40, 1225 - 1239. https: // doi. org / 10.1111 / jbi. 12080" type="journal article">
Pepper 
<emphasis id="B96DEAC8F27EFFACFB160D4DADC01F71" box="[1208,1264,799,824]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
(2013)
</bibRefCitation>
that the Fortescue River system provides a major biogeographic break within the region that can potentially drive allopatric speciation. Other species that also share similar distributional patterns are 
<taxonomicName id="4C194D59F27EFFACFBBC0D05ADE61FC9" authorityName="Doughty, Kealley &amp; donnellan" authorityYear="2011" box="[1042,1238,873,896]" class="Reptilia" family="Scincidae" genus="Egernia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="cygnitos">
<emphasis id="B96DEAC8F27EFFACFBBC0D05ADE61FC9" box="[1042,1238,873,896]" italics="true" pageId="17" pageNumber="18">Egernia cygnitos</emphasis>
</taxonomicName>
vs. 
<taxonomicName id="4C194D59F27EFFACFABE0D05A8A51FED" authority="(Doughty et al. 2011 a)" authorityName="Doughty et al." authorityYear="2011" class="Reptilia" family="Scincidae" genus="Egernia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="epsisolus">
<emphasis id="B96DEAC8F27EFFACFABE0D05ACAC1FC9" box="[1296,1436,871,896]" italics="true" pageId="17" pageNumber="18">E. epsisolus</emphasis>
(
<bibRefCitation id="EF884B2BF27EFFACFF300DE0A8BE1FED" author="Doughty" box="[158,398,908,933]" pageId="17" pageNumber="18" refString="Doughty, P., Kealley, L. &amp; Donnellan, S. C. (2011 a) Revision of the Pygmy Spiny-tailed Skinks (Egernia depressa speciesgroup) from western Australia, with descriptions of three new species. Records of the Western Australian Museum, 26, 115 - 137." type="journal article" year="2011" yearSuffix="a">
Doughty 
<emphasis id="B96DEAC8F27EFFACFEA60DE1A8701FEC" box="[264,320,908,933]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2011a
</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C194D59F27EFFACFE0E0DE0ABA91FED" authorityName="Kluge" authorityYear="1963" box="[416,665,908,933]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="savagei">
<emphasis id="B96DEAC8F27EFFACFE0E0DE0ABA91FED" box="[416,665,908,933]" italics="true" pageId="17" pageNumber="18">Diplodactylus savagei</emphasis>
</taxonomicName>
vs. 
<taxonomicName id="4C194D59F27EFFACFD6B0DE1AD701FED" authority="(Doughty et al. 2010)" authorityName="Doughty et al." authorityYear="2010" box="[709,1088,908,933]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="galaxias">
<emphasis id="B96DEAC8F27EFFACFD6B0DE1AA761FEC" box="[709,838,908,933]" italics="true" pageId="17" pageNumber="18">D. galaxias</emphasis>
(Doughty 
<emphasis id="B96DEAC8F27EFFACFC100DE1AAC51FEC" box="[958,1013,908,933]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2010)
</taxonomicName>
, and 
<taxonomicName id="4C194D59F27EFFACFBD70DE0AC651FEC" authorityName="Storr" authorityYear="1980" box="[1145,1365,908,933]" class="Reptilia" family="Agamidae" genus="Diporiphora" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="valens">
<emphasis id="B96DEAC8F27EFFACFBD70DE0AC651FEC" box="[1145,1365,908,933]" italics="true" pageId="17" pageNumber="18">Diporiphora valens</emphasis>
</taxonomicName>
vs. 
<taxonomicName id="4C194D59F27EFFACFA2F0DE1AB301F81" authority="(Doughty et al. 2012)" authorityName="Doughty et al." authorityYear="2012" class="Reptilia" family="Agamidae" genus="Diporiphora" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="vescus">
<emphasis id="B96DEAC8F27EFFACFA2F0DE1A9D01F81" italics="true" pageId="17" pageNumber="18">D. vescus</emphasis>
(
<bibRefCitation id="EF884B2BF27EFFACFF590DC3A8C81F81" author="Doughty" box="[247,504,943,968]" pageId="17" pageNumber="18" refString="Doughty, P., Kealley, L. &amp; Melville, J. (2012) Taxonomic assessment of Diporiphora (Reptilia: Agamidae) dragon lizards from the western arid zone of Australia. Zootaxa, 3518, 1 - 24." type="journal article" year="2012">
Doughty 
<emphasis id="B96DEAC8F27EFFACFEC50DDDA89C1F81" box="[363,428,943,968]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2012
</bibRefCitation>
)
</taxonomicName>
. Species distributions, however, do not always fall exactly along the same biogeographic breaks, with each taxon showing idiosyncratic patterns between species along the Fortescue River barrier (see 
<bibRefCitation id="EF884B2BF27EFFACFEB20D9BAB3C1859" author="Doughty" box="[284,524,1015,1040]" pageId="17" pageNumber="18" refString="Doughty, P., Rolfe, J. K., Burbidge, A. H., Pearson, D. J. &amp; Kendrick, P. G. (2011 b) Herpetologial assemblages of the Pilbara biogeographic region, Western Australia: ecological associations, biogeographic patterns and conservations. Records of the Western Australian Museum, 78 (Supplement), 315 - 341." type="journal article" year="2011" yearSuffix="b">
Doughty 
<emphasis id="B96DEAC8F27EFFACFE2B0D95A88E1859" box="[389,446,1015,1040]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2011b
</bibRefCitation>
; 
<bibRefCitation id="EF884B2BF27EFFACFDB60D94ABD91859" author="Pepper" box="[536,745,1015,1040]" pageId="17" pageNumber="18" refString="Pepper, M., Doughty, P. &amp; Keogh, S. J. (2013) Synthesis: Geodiversity and endemism in the iconic Australian Pilbara region: a review of landscape evolution and biotic response in an ancient refugium. Journal of Biogeography, 40, 1225 - 1239. https: // doi. org / 10.1111 / jbi. 12080" type="journal article" year="2013">
Pepper 
<emphasis id="B96DEAC8F27EFFACFDC30D95AB951859" box="[621,677,1015,1040]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
<caption id="DF666652F27EFFACFF390A2CA86618FE" pageId="17" pageNumber="18">
<paragraph id="8BA636DAF27EFFACFF390A2CA86618FE" blockId="17.[151,1436,1088,1207]" pageId="17" pageNumber="18">
<emphasis id="B96DEAC8F27EFFACFF390A2CA83A181E" bold="true" box="[151,266,1088,1111]" pageId="17" pageNumber="18">TABLE 1.</emphasis>
Morphological characteristics of the 
<taxonomicName id="4C194D59F27EFFACFD0F0A2CAA30181E" authorityName="Storr" authorityYear="1964" box="[673,768,1088,1111]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B96DEAC8F27EFFACFD0F0A2CAA30181E" box="[673,768,1088,1111]" italics="true" pageId="17" pageNumber="18">Ctenotus</emphasis>
</taxonomicName>
species considered in this study. Number in parentheses after taxon name is number of individuals measured. Definitions for each character are provided in the Materials and Methods. Summaries show means ± standard deviations for each character (range in parentheses). Measurements for continuous characters in mm.
</paragraph>
</caption>
<paragraph id="8BA636DAF27EFFACFF0B0ABEAC5018AE" blockId="17.[165,1376,1234,2020]" box="[165,1376,1234,1256]" pageId="17" pageNumber="18">
Character 
<taxonomicName id="4C194D59F27EFFACFEC00ABEA88618AE" authorityName="Storr" authorityYear="1969" box="[366,438,1234,1255]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="piankai">
<emphasis id="B96DEAC8F27EFFACFEC00ABEA88618AE" box="[366,438,1234,1255]" italics="true" pageId="17" pageNumber="18">piankai</emphasis>
</taxonomicName>
(10) 
<taxonomicName id="4C194D59F27EFFACFDEE0ABEAB9118AE" authority="Rabosky &amp; Doughty &amp; Huang, 2017" authorityName="Rabosky &amp; Doughty &amp; Huang" authorityYear="2017" box="[576,673,1234,1255]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="rhabdotus" status="sp. nov.">
<emphasis id="B96DEAC8F27EFFACFDEE0ABEAB9118AE" box="[576,673,1234,1255]" italics="true" pageId="17" pageNumber="18">rhabdotus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B3F27EFFACFD090ABEABC518A1" box="[679,757,1234,1256]" pageId="17" pageNumber="18" rank="species">
<emphasis id="B96DEAC8F27EFFACFD090ABEABC518A1" bold="true" box="[679,757,1234,1256]" pageId="17" pageNumber="18">sp. nov.</emphasis>
</taxonomicNameLabel>
(14) 
<taxonomicName id="4C194D59F27EFFACFC2A0ABEAAE518AE" authorityName="Storr" authorityYear="1975" box="[900,981,1234,1255]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="duricola">
<emphasis id="B96DEAC8F27EFFACFC2A0ABEAAE518AE" box="[900,981,1234,1255]" italics="true" pageId="17" pageNumber="18">duricola</emphasis>
</taxonomicName>
(20) 
<emphasis id="B96DEAC8F27EFFACFBDF0ABEADED18AE" box="[1137,1245,1234,1255]" italics="true" pageId="17" pageNumber="18">pallassotus</emphasis>
<emphasis id="B96DEAC8F27EFFACFB4D0ABEAC0118A1" bold="true" box="[1251,1329,1234,1256]" pageId="17" pageNumber="18">sp. nov.</emphasis>
(18)
</paragraph>
</subSubSection>
<subSubSection id="C3036551F27EFFACFF0B0A9EAC321BAD" pageId="17" pageNumber="18" type="description">
<paragraph id="8BA636DAF27EFFACFF0B0A9EAC3E196E" blockId="17.[165,1376,1234,2020]" pageId="17" pageNumber="18">SVL 45.6±4.7 53.4±4.6 51.1±3.8 52.1±5.5 (38–53) (44–61) (44–59) (45–65)</paragraph>
<paragraph id="8BA636DAF27EFFACFF0B0B5EAC24192E" blockId="17.[165,1376,1234,2020]" pageId="17" pageNumber="18">RTL 2.3±0.2 2.2±0.2 2.0±0.2 2.1±0.2 (1.9–2.5) (1.9–2.3) (1.6–2.4) (1.7–2.4)</paragraph>
<paragraph id="8BA636DAF27EFFACFF0B0B1DAC2B19EF" blockId="17.[165,1376,1234,2020]" pageId="17" pageNumber="18">HL 8.8±0.6 10.3±0.7 9.8±0.6 10.0±0.7 (8.0–9.6) (8.9–11.1) (8.3–10.7) (9.0–10.9)</paragraph>
<paragraph id="8BA636DAF27EFFACFF0B0BDDAC2519AF" blockId="17.[165,1376,1234,2020]" pageId="17" pageNumber="18">HW 5.3±0.5 6.4±0.6 6.1±0.5 6.3±0.4 (4.7–6.2) (5.1–7.5) (5.1–6.7) (5.6–7.0)</paragraph>
<paragraph id="8BA636DAF27EFFACFF0B0B9DAC241A6F" blockId="17.[165,1376,1234,2020]" pageId="17" pageNumber="18">HD 4.3±0.3 5.0±0.5 4.9±0.4 5.2±0.5 (3.8–4.6) (4.2–5.7) (4.2–5.9) (4.1–5.8)</paragraph>
<paragraph id="8BA636DAF27EFFACFF0B085DAC3E1A2C" blockId="17.[165,1376,1234,2020]" pageId="17" pageNumber="18">MBSR 23.8±0.6 26.3±0.7 27.4±1.1 27.9±1.2 (22–24) (26–28) (26–29) (26–30)</paragraph>
<paragraph id="8BA636DAF27EFFACFF0B081CAC251AEC" blockId="17.[165,1376,1234,2020]" pageId="17" pageNumber="18">4ToeL 7.6±0.4 8.5±0.5 8.6±0.7 8.7±0.4 (7.1–8.1) (7.7–9.2) (7.3–9.9) (7.8–9.4)</paragraph>
<paragraph id="8BA636DAF27EFFACFF0B08DCAC3E1AAC" blockId="17.[165,1376,1234,2020]" pageId="17" pageNumber="18">ToeLam 22.2±1.9 23.3±1.6 23.1±1.8 22.5±1.3 (18–25) (21–26) (20–25) (21–25)</paragraph>
<paragraph id="8BA636DAF27EFFACFF0B089CAC321B6C" blockId="17.[165,1376,1234,2020]" pageId="17" pageNumber="18">SupLab 8.0±0.0 7.9±0.4 8.0±0.3 8.0±0.2 (8–8) (7–9) (7–9) (7–9)</paragraph>
<paragraph id="8BA636DAF27EFFACFF0B0943AC381B2D" blockId="17.[165,1376,1234,2020]" pageId="17" pageNumber="18">InfLab 8.0±0.0 8.1±0.7 8.3±0.7 8.1±0.6 (8–8) (7–9) (7–10) (7–10)</paragraph>
<paragraph id="8BA636DAF27EFFACFF0B0903AC321BED" blockId="17.[165,1376,1234,2020]" pageId="17" pageNumber="18">Supraciliaries 7.8±0.4 7.9±0.5 7.7±0.5 7.7±0.5 (7–8) (7–9) (7–9) (6–9)</paragraph>
<paragraph id="8BA636DAF27EFFACFF0B09C3AC321BAD" blockId="17.[165,1376,1234,2020]" pageId="17" pageNumber="18">EarLob 3.5±0.6 3.3±0.6 3.7±0.8 3.8±0.7 (3–5) (2–5) (2–6) (2–6)</paragraph>
</subSubSection>
</treatment>
</document>