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<document id="F0330E661562F2561013925D333C6CA0" ID-DOI="10.5281/zenodo.14681984" ID-ISSN="0253-116X" ID-Zenodo-Dep="14681984" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737168586081" checkinUser="felipe" docAuthor="Kahlen, Manfred, Eckelt, Andreas, C, Alpi &amp; C, Alpi" docDate="2024" docId="03B087C9FFAEB334E1E17917FBEAFEA9" docLanguage="en" docName="LinzerbiolBeitr.56.1.61-67.pdf" docOrigin="Linzer biologische Beiträge 56 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.12:LinzerbiolBeitr.2006-.journal_article.type1" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article.type1" docStyleVersion="12" docTitle="Bryaxis cialanciaensis Kahlen, Eckelt, C &amp; C, 2024, nov.sp." docType="treatment" docVersion="2" lastPageNumber="65" masterDocId="FF89FFB1FFAFB330E1427B27FFC4FF94" masterDocTitle="A new species of the genus Bryaxis from the Northern Cottian Alps (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="67" masterPageNumber="61" pageNumber="62" updateTime="1737168945555" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="51279A6EEF1025B7AAE4B2CA7910132B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="44980FCEB39775829C6B5B3A2B22A91A">
<mods:title id="E864B7D8C4BE3BFFA2574A0D933F90C2">A new species of the genus Bryaxis from the Northern Cottian Alps (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
</mods:titleInfo>
<mods:name id="539EE4917BBB3D5468140333F47B9EA2" type="personal">
<mods:role id="6C2897F7B8047F73F29A19E7C22AD1E6">
<mods:roleTerm id="0B99A6FBE06A47A282DBED8A6681C0D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="450D6422A572CEAA490C2AF6DB3FDDED">Kahlen, Manfred</mods:namePart>
</mods:name>
<mods:name id="0EAFBDE3CDCB8C874E4C7C5AD0D4713A" type="personal">
<mods:role id="60FE7B00D6FB84B17D44340875CBEEBE">
<mods:roleTerm id="552785579D974C425DB328D9AD5C65B6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EAE901EA91AB9B9AEA036D07A97521DA">Eckelt, Andreas</mods:namePart>
</mods:name>
<mods:name id="E5DF4B31351F1D48E165029769AB3B5C" type="personal">
<mods:role id="A130C0E2C339D44F1C0D32C995C6BEE8">
<mods:roleTerm id="CFF2C6C3E99FE41AB27CB81887F34B0D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DFC83A3A6C61BCAD3C067CC01FC67CC9">C, Alpi</mods:namePart>
</mods:name>
<mods:name id="1A8E1CAA67F0ADE4040149DF248D492A" type="personal">
<mods:role id="B30B2ED5A51FACDFD837659692B06E2B">
<mods:roleTerm id="A008BFC81C7EA6D6FE97C8D42D137950">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8844001757B314FFF34B47A210B67EA5">C, Alpi</mods:namePart>
</mods:name>
<mods:typeOfResource id="091CC061D8A1A548CB8EC114D6107F14">text</mods:typeOfResource>
<mods:relatedItem id="50E8A06F0F82B3FB7D042E4C00DAA952" type="host">
<mods:titleInfo id="5D615A58B24202A4D46D321F1C4614BE">
<mods:title id="2D89038978C390BE99DE9D4FE6DE51F5">Linzer biologische Beiträge</mods:title>
</mods:titleInfo>
<mods:part id="DDF8D3955AD5BB0B3BF5356F5E84310F">
<mods:date id="F57A3F276E07CE995EA6EEDD5246699F">2024</mods:date>
<mods:detail id="1F66FB8D6AA90CCAD592B46E5D12650B" type="volume">
<mods:number id="3023CD99B091B5F9E50DC9AFD16C47D2">56</mods:number>
</mods:detail>
<mods:detail id="97AADCED5D8A10862042CEDA2BB37AC1" type="issue">
<mods:number id="5515A2DBA87F8A59DE226FEF15AA613A">1</mods:number>
</mods:detail>
<mods:extent id="157F726D3DB8033709F3D27DE3359E1C" unit="page">
<mods:start id="67C30556C9D1067707CEE810B59844D4">61</mods:start>
<mods:end id="D5DC6FBF3A333AE2804FE5591F9B98B6">67</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="43FB5B497B3B97D92A3B40803A6088A9">journal article</mods:classification>
<mods:identifier id="2C9B6FADA558FF151400C32F97D9E9B2" type="DOI">10.5281/zenodo.14681984</mods:identifier>
<mods:identifier id="300D5DD29BCBB284822F6F4965208F06" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="F562184F18D3BE2C336A2F695C23C294" type="Zenodo-Dep">14681984</mods:identifier>
</mods:mods>
<treatment id="03B087C9FFAEB334E1E17917FBEAFEA9" LSID="urn:lsid:plazi:treatment:03B087C9FFAEB334E1E17917FBEAFEA9" httpUri="http://treatment.plazi.org/id/03B087C9FFAEB334E1E17917FBEAFEA9" lastPageId="4" lastPageNumber="65" pageId="1" pageNumber="62">
<subSubSection id="C3036554FFAEB331E1E17917FD88FDD2" box="[163,588,560,582]" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="8BA636DFFFAEB331E1E17917FD88FDD2" blockId="1.[163,588,560,582]" box="[163,588,560,582]" pageId="1" pageNumber="62">
<heading id="D0EE81B3FFAEB331E1E17917FD88FDD2" box="[163,588,560,582]" fontSize="8" level="2" pageId="1" pageNumber="62" reason="2">
<emphasis id="B96DEACDFFAEB331E1E17917FD88FDD2" bold="true" box="[163,588,560,582]" pageId="1" pageNumber="62">
<taxonomicName id="4C194D5CFFAEB331E1E17917FE49FDD2" authorityName="Kahlen &amp; Eckelt &amp; C &amp; C" authorityYear="2024" box="[163,397,560,582]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="cialanciaensis" status="sp. nov.">
<emphasis id="B96DEACDFFAEB331E1E17917FE49FDD2" bold="true" box="[163,397,560,582]" italics="true" pageId="1" pageNumber="62">Bryaxis cialanciaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B6FFAEB331E0D17917FE24FDD1" box="[403,480,560,581]" pageId="1" pageNumber="62" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="13222A5AFFAEB331E0AF7917FD80FDD2" box="[493,580,560,582]" captionStart="Figs 1-4" captionStartId="2.[163,203,1492,1514]" captionTargetBox="[164,1045,210,1478]" captionTargetId="figure-46@2.[164,1069,210,1478]" captionTargetPageId="2" captionText="Figs 1-4: (1) Bryaxis cialanciaensis nov.sp. Holotype, scale bar 0,5 mm; (2) aedeagus dorsal B. cialanciaensis, scale bar 0,1 mm; (3-4) Habitus (scale bar 0,5 mm) and aedeagus (scale bar 0,1 mm) of B. gallicus." figureDoi="http://doi.org/10.5281/zenodo.14681986" httpUri="https://zenodo.org/record/14681986/files/figure.png" pageId="1" pageNumber="62">Figs 1-2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3036554FFAEB334E1E17971FBEAFEA9" lastPageId="4" lastPageNumber="65" pageId="1" pageNumber="62" type="materials_examined">
<paragraph id="8BA636DFFFAEB331E1E17971FEE4FC87" blockId="1.[163,1071,596,787]" pageId="1" pageNumber="62">
<materialsCitation id="3B713C82FFAEB331E1E17971FED0FD08" collectorName="Kahlen &amp; Brachat" county="Torino" determinerName="Kahlen" elevation="2350" latitude="44.875" location="Punta Freidour" longLatPrecision="55" longitude="7.1217" municipality="Parco Naturale Conca Cialancia" pageId="1" pageNumber="62" specimenCount="1" stateProvince="Piemonte" typeStatus="holotype">
<typeStatus id="54A2887DFFAEB331E1E17971FF31FDF8" box="[163,245,598,620]" pageId="1" pageNumber="62" type="holotype">Holotype</typeStatus>
♁:
<collectingRegion id="49DDF83DFFAEB331E0627971FEB0FDF8" box="[288,372,598,620]" country="Italy" name="Piemonte" pageId="1" pageNumber="62">Piemonte</collectingRegion>
/
<collectingCounty id="62C74E53FFAEB331E0397971FE73FDF8" box="[379,439,598,620]" pageId="1" pageNumber="62">Torino</collectingCounty>
,
<collectingMunicipality id="6BC2ACA5FFAEB331E0867971FD29FDF8" box="[452,749,598,620]" pageId="1" pageNumber="62">Parco Naturale Conca Cialancia</collectingMunicipality>
,
<location id="8EC66004FFAEB331E3B97971FC47FDF8" LSID="urn:lsid:plazi:treatment:03B087C9FFAEB334E1E17917FBEAFEA9:8EC66004FFAEB331E3B97971FC47FDF8" box="[763,899,598,620]" county="Torino" latitude="44.875" longLatPrecision="55" longitude="7.1217" municipality="Parco Naturale Conca Cialancia" name="Punta Freidour" pageId="1" pageNumber="62" stateProvince="Piemonte">Punta Freidour</location>
W-Side,
<quantity id="4CE19B3AFFAEB331E2A27971FBEFFDFF" box="[992,1067,598,620]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="1" pageNumber="62" unit="m" value="2350.0">
<elevation id="0034D1ECFFAEB331E2A27971FBEFFDFF" box="[992,1067,598,620]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="1" pageNumber="62" unit="m" value="2350.0">2350 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D5018FFAEB331E18D7949FEE2FD10" box="[207,294,622,644]" degrees="44.875" direction="north" orientation="latitude" pageId="1" pageNumber="62" precision="55" value="44.875">44.875°N</geoCoordinate>
,
<geoCoordinate id="EE2D5018FFAEB331E06C7949FE47FD17" box="[302,387,622,643]" degrees="7.1217" direction="east" orientation="longitude" pageId="1" pageNumber="62" precision="5" value="7.1217">7.1217°E</geoCoordinate>
, 28.6.202 1, leg.
<collectorName id="26EC5309FFAEB331E3527949FD8BFD10" box="[528,591,622,644]" pageId="1" pageNumber="62">Kahlen</collectorName>
/
<taxonomicName id="4C194D5CFFAEB331E31D7948FCE7FD17" authorityName="Kahlen &amp; Eckelt &amp; C &amp; C" authorityYear="2024" box="[607,803,622,644]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="cialanciaensis" status="sp. nov.">
<emphasis id="B96DEACDFFAEB331E31D7948FCE7FD17" box="[607,803,622,644]" italics="true" pageId="1" pageNumber="62">Bryaxis cialanciaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B6FFAEB331E26A7949FC44FD17" box="[808,896,622,643]" pageId="1" pageNumber="62" rank="species">spec. nov.</taxonomicNameLabel>
♁ det.
<determinerName id="7E7ADA1BFFAEB331E2FC7949FC39FD10" box="[958,1021,622,644]" pageId="1" pageNumber="62">Kahlen</determinerName>
/ vid.
<collectorName id="26EC5309FFAEB331E19279A1FED0FD08" box="[208,276,646,668]" pageId="1" pageNumber="62">Brachat</collectorName>
</materialsCitation>
/
<materialsCitation id="3B713C82FFAEB331E07079A1FE1FFD0F" box="[306,475,646,668]" collectionCode="TLMF" pageId="1" pageNumber="62" specimenCount="1" stateProvince="Piemonte" typeStatus="holotype">
<typeStatus id="54A2887DFFAEB331E07079A1FE40FD08" box="[306,388,646,668]" pageId="1" pageNumber="62" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED08AE1AFFAEB331E0D479A1FE12FD0F" box="[406,470,646,667]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="1" pageNumber="62">TLMF</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713C82FFAEB331E0A879A1FBEEFD27" collectionCode="TLMF" collectorName="Kahlen" county="Torino" elevation="2350" latitude="44.875" location="Punta Freidour" longLatPrecision="55" longitude="7.1217" municipality="Parco Naturale Conca Cialancia" pageId="1" pageNumber="62" specimenCount="4" specimenCount-female="4" stateProvince="Piemonte" typeStatus="paratype">
<typeStatus id="54A2887DFFAEB331E0A879A1FD86FD0F" box="[490,578,646,667]" pageId="1" pageNumber="62" type="paratype">Paratypes</typeStatus>
: 1♁,
<specimenCount id="9D1FFD56FFAEB331E33C79A1FD6FFD0F" box="[638,683,646,667]" count="4" pageId="1" pageNumber="62" type="female">4♀♀</specimenCount>
,
<collectingRegion id="49DDF83DFFAEB331E3FE79A1FCD4FD08" box="[700,784,646,668]" country="Italy" name="Piemonte" pageId="1" pageNumber="62">Piemonte</collectingRegion>
/
<collectingCounty id="62C74E53FFAEB331E25479A1FC96FD08" box="[790,850,646,668]" pageId="1" pageNumber="62">Torino</collectingCounty>
,
<collectingMunicipality id="6BC2ACA5FFAEB331E22379A1FEE0FD20" pageId="1" pageNumber="62">Parco Naturale Conca Cialancia</collectingMunicipality>
,
<location id="8EC66004FFAEB331E06F79B9FE74FD20" LSID="urn:lsid:plazi:treatment:03B087C9FFAEB334E1E17917FBEAFEA9:8EC66004FFAEB331E06F79B9FE74FD20" box="[301,432,670,692]" county="Torino" latitude="44.875" longLatPrecision="55" longitude="7.1217" municipality="Parco Naturale Conca Cialancia" name="Punta Freidour" pageId="1" pageNumber="62" stateProvince="Piemonte">Punta Freidour</location>
W-Side,
<quantity id="4CE19B3AFFAEB331E34679B9FD8DFD27" box="[516,585,670,692]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="1" pageNumber="62" unit="m" value="2350.0">
<elevation id="0034D1ECFFAEB331E34679B9FD8DFD27" box="[516,585,670,692]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="1" pageNumber="62" unit="m" value="2350.0">2350 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D5018FFAEB331E31179B9FD6FFD20" box="[595,683,670,692]" degrees="44.875" direction="north" orientation="latitude" pageId="1" pageNumber="62" precision="55" value="44.875">44.875°N</geoCoordinate>
,
<geoCoordinate id="EE2D5018FFAEB331E3F179B9FCCCFD27" box="[691,776,670,691]" degrees="7.1217" direction="east" orientation="longitude" pageId="1" pageNumber="62" precision="5" value="7.1217">7.1217°E</geoCoordinate>
, 28.6.202 1, leg.
<collectorName id="26EC5309FFAEB331E2DB79B9FC1CFD20" box="[921,984,670,692]" pageId="1" pageNumber="62">Kahlen</collectorName>
(
<collectionCode id="ED08AE1AFFAEB331E2A779B9FBE1FD27" box="[997,1061,670,691]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="1" pageNumber="62">TLMF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713C82FFAEB331E1927993FD72FD77" collectionCode="TLMF" collectorName="Kahlen" county="Torino" elevation="2250" latitude="44.8835" location="Punta Lausarot" longLatPrecision="6" longitude="7.1216" municipality="Parco Naturale Conca Cialancia" pageId="1" pageNumber="62" specimenCount="1" stateProvince="Piemonte" typeStatus="paratype">
1♁
<collectingRegion id="49DDF83DFFAEB331E1BA7991FE88FD58" box="[248,332,694,716]" country="Italy" name="Piemonte" pageId="1" pageNumber="62">Piemonte</collectingRegion>
/
<collectingCounty id="62C74E53FFAEB331E0117991FE4BFD58" box="[339,399,694,716]" pageId="1" pageNumber="62">Torino</collectingCounty>
,
<collectingMunicipality id="6BC2ACA5FFAEB331E0DF7991FD09FD58" box="[413,717,694,716]" pageId="1" pageNumber="62">Parco Naturale Conca Cialancia</collectingMunicipality>
,
<location id="8EC66004FFAEB331E39E7991FCA3FD5F" LSID="urn:lsid:plazi:treatment:03B087C9FFAEB334E1E17917FBEAFEA9:8EC66004FFAEB331E39E7991FCA3FD5F" box="[732,871,694,715]" county="Torino" latitude="44.8835" longLatPrecision="6" longitude="7.1216" municipality="Parco Naturale Conca Cialancia" name="Punta Lausarot" pageId="1" pageNumber="62" stateProvince="Piemonte">Punta Lausarot</location>
N-Rücken,
<quantity id="4CE19B3AFFAEB331E29C7991FBEFFD5F" box="[990,1067,694,716]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="1" pageNumber="62" unit="m" value="2250.0">
<elevation id="0034D1ECFFAEB331E29C7991FBEFFD5F" box="[990,1067,694,716]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="1" pageNumber="62" unit="m" value="2250.0">2250 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D5018FFAEB331E19279E9FEF5FD70" box="[208,305,718,740]" degrees="44.8835" direction="north" orientation="latitude" pageId="1" pageNumber="62" precision="5" value="44.8835">44.8835°N</geoCoordinate>
,
<geoCoordinate id="EE2D5018FFAEB331E07E79E9FE54FD70" box="[316,400,718,740]" degrees="7.1216" direction="east" orientation="longitude" pageId="1" pageNumber="62" precision="5" value="7.1216">7.1216°E</geoCoordinate>
, 28.6.202 1, leg.
<collectorName id="26EC5309FFAEB331E36679E9FDA7FD70" box="[548,611,718,740]" pageId="1" pageNumber="62">Kahlen</collectorName>
(
<collectionCode id="ED08AE1AFFAEB331E33279E9FD74FD77" box="[624,688,718,739]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="1" pageNumber="62">TLMF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713C82FFAEB331E38279EBFED8FC87" collectionCode="TLMF" collectorName="Kahlen" elevation="2040" latitude="44.8872" location="Parco Naturale Conca Cialancia" longLatPrecision="6" longitude="7.1266" municipality="Torino" pageId="1" pageNumber="62" specimenCount="1" specimenCount-female="1" stateProvince="Piemonte" typeStatus="paratype">
1♁,
<specimenCount id="9D1FFD56FFAEB331E3AA79E9FCC0FD77" box="[744,772,718,739]" count="1" pageId="1" pageNumber="62" type="female">1♀</specimenCount>
<collectingRegion id="49DDF83DFFAEB331E24879E9FC9AFD70" box="[778,862,718,740]" country="Italy" name="Piemonte" pageId="1" pageNumber="62">Piemonte</collectingRegion>
/
<collectingMunicipality id="6BC2ACA5FFAEB331E22779E9FC65FD70" box="[869,929,718,740]" pageId="1" pageNumber="62">Torino</collectingMunicipality>
,
<location id="8EC66004FFAEB331E2E879E9FEA5FD68" LSID="urn:lsid:plazi:treatment:03B087C9FFAEB334E1E17917FBEAFEA9:8EC66004FFAEB331E2E879E9FEA5FD68" latitude="44.8872" longLatPrecision="6" longitude="7.1266" municipality="Torino" name="Parco Naturale Conca Cialancia" pageId="1" pageNumber="62" stateProvince="Piemonte">Parco Naturale Conca Cialancia</location>
, Lago Lauson Umg.,
<quantity id="4CE19B3AFFAEB331E35D79C1FD53FD6F" box="[543,663,742,764]" metricMagnitude="3" metricUnit="m" metricValue="2.04" metricValueMax="2.05" metricValueMin="2.03" pageId="1" pageNumber="62" unit="m" value="2040.0" valueMax="2050.0" valueMin="2030.0">
<elevation id="0034D1ECFFAEB331E35D79C1FD53FD6F" box="[543,663,742,764]" metricMagnitude="3" metricUnit="m" metricValue="2.04" metricValueMax="2.05" metricValueMin="2.03" pageId="1" pageNumber="62" unit="m" value="2040.0" valueMax="2050.0" valueMin="2030.0">2030-2050 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D5018FFAEB331E3E279C1FCC5FD6F" box="[672,769,742,763]" degrees="44.8872" direction="north" orientation="latitude" pageId="1" pageNumber="62" precision="5" value="44.8872">44.8872°N</geoCoordinate>
,
<geoCoordinate id="EE2D5018FFAEB331E24879C1FC9BFD68" box="[778,863,742,764]" degrees="7.1266" direction="east" orientation="longitude" pageId="1" pageNumber="62" precision="5" value="7.1266">7.1266°E</geoCoordinate>
, 17.7.202 2, leg.
<collectorName id="26EC5309FFAEB331E2AC79C1FBEAFD68" box="[1006,1070,742,764]" pageId="1" pageNumber="62">Kahlen</collectorName>
(
<collectionCode id="ED08AE1AFFAEB331E19479D9FED2FC87" box="[214,278,766,787]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="1" pageNumber="62">TLMF</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636DFFFAEB331E1E07804FEA1FCC2" blockId="1.[162,1069,802,854]" pageId="1" pageNumber="62">E t y m o l o g y: The specific epithet is derived from the name of the collection area &quot;Conca Cialancia&quot;.</paragraph>
<paragraph id="8BA636DFFFAEB331E1E17842FC72FA81" blockId="1.[163,1070,869,1302]" pageId="1" pageNumber="62">
D e s c r i p t i o n: Habitus as in
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. Body length
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. Body, including appendages, uniformly reddish-brown and glossy. Setae sparse, moderately long, and nearly appressed. Head length (from antennal insertion to neck)
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, head width (across eyes)
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, frons slightly depressed, dull and granular punctate, vertex glossy with widely and finely punctured surface. Pronotum wider than long (W
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, L
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), glossy with very widely spaced punctures, with a sharply marked transverse groove in the last quarter before the base, strongly arched, widest before the middle, slightly heart-shaped. Elytra barely wider than long together (W
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, L 0.67-
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), glossy and with loose but strong punctation. Abdominal tergites glossy with very widely spaced fine punctation. ♁: Antennal segment 1 exactly square (
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), only moderately glossy, slightly asymmetrically humped on the upper side, this elevation towards the center or more towards the inner edge; antennal segment 2 strongly spherical (
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) slightly longer than wide; segments 3-8 narrower, also spherical, 3 and 4 barely wider than the following ones, broadening from segment 9 towards the club (
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). Only males with normal leg development are present. Aedeagus ♁:
<figureCitation id="13222A5AFFAEB331E2367E27FC72FA81" box="[884,950,1280,1302]" captionStart="Figs 1-4" captionStartId="2.[163,203,1492,1514]" captionTargetBox="[164,1045,210,1478]" captionTargetId="figure-46@2.[164,1069,210,1478]" captionTargetPageId="2" captionText="Figs 1-4: (1) Bryaxis cialanciaensis nov.sp. Holotype, scale bar 0,5 mm; (2) aedeagus dorsal B. cialanciaensis, scale bar 0,1 mm; (3-4) Habitus (scale bar 0,5 mm) and aedeagus (scale bar 0,1 mm) of B. gallicus." figureDoi="http://doi.org/10.5281/zenodo.14681986" httpUri="https://zenodo.org/record/14681986/files/figure.png" pageId="1" pageNumber="62">Fig. 2.</figureCitation>
</paragraph>
<paragraph id="8BA636DFFFAEB331E1E17E02FDB8FA24" blockId="1.[163,1070,1317,1456]" pageId="1" pageNumber="62">
C o m p a r a t i v e n o t e s: Based on morphology
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<emphasis id="B96DEACDFFAEB331E3917E02FC72FAAE" box="[723,950,1317,1338]" italics="true" pageId="1" pageNumber="62">Bryaxis cialanciaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B6FFAEB331E2F97E02FBC1FAAE" box="[955,1029,1317,1338]" pageId="1" pageNumber="62" rank="species">nov.sp.</taxonomicNameLabel>
and
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<emphasis id="B96DEACDFFAEB331E1E17E64FE8EFAC3" box="[163,330,1346,1368]" italics="true" pageId="1" pageNumber="62">Bryaxis gallicus</emphasis>
</taxonomicName>
are clearly distinguishable in the shape of antennal segments and the aedeagi. Other features (head, pronotum, elytra, abdomen) do not offer consistent distinguishing characteristics. The
<specimenCount id="9D1FFD56FFAEB331E3407E5AFDECFA06" box="[514,552,1405,1426]" count="1" pageId="1" pageNumber="62" type="female">♀♀</specimenCount>
of the species are distinguishable by measuring the length-width ratio of the 1st antennal segment.
</paragraph>
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<paragraph id="8BA636DFFFADB332E1E17EF3FEE6F98E" blockId="2.[163,1071,1492,1562]" pageId="2" pageNumber="63">
<emphasis id="B96DEACDFFADB332E1E17EF3FF36FA7D" bold="true" box="[163,242,1492,1514]" pageId="2" pageNumber="63">Figs 1-4</emphasis>
: (
<emphasis id="B96DEACDFFADB332E04A7EF3FED7FA7D" bold="true" box="[264,275,1492,1513]" pageId="2" pageNumber="63">1</emphasis>
)
<taxonomicName id="4C194D5CFFADB332E0667EF2FE30FA7D" authorityName="Kahlen &amp; Eckelt &amp; C &amp; C" authorityYear="2024" box="[292,500,1492,1514]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="cialanciaensis" status="sp. nov.">
<emphasis id="B96DEACDFFADB332E0667EF2FE30FA7D" box="[292,500,1492,1514]" italics="true" pageId="2" pageNumber="63">Bryaxis cialanciaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B6FFADB332E0BE7EF2FDFBFA7E" box="[508,575,1493,1514]" pageId="2" pageNumber="63" rank="species">nov.sp.</taxonomicNameLabel>
Holotype, scale bar 0,5 mm; (
<emphasis id="B96DEACDFFADB332E2217EF3FCAAFA7D" bold="true" box="[867,878,1492,1513]" pageId="2" pageNumber="63">2</emphasis>
) aedeagus dorsal
<taxonomicName id="4C194D5CFFADB332E5587EF2FEE1F995" authorityName="Kahlen &amp; Eckelt &amp; C &amp; C" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="cialanciaensis">
<emphasis id="B96DEACDFFADB332E5587EF2FEE1F995" italics="true" pageId="2" pageNumber="63">B. cialanciaensis</emphasis>
</taxonomicName>
, scale bar 0,1 mm; (
<emphasis id="B96DEACDFFADB332E09C7ECBFE38F995" bold="true" box="[478,508,1516,1537]" pageId="2" pageNumber="63">3-4</emphasis>
) Habitus (scale bar 0,5 mm) and aedeagus (scale bar 0,1 mm) of
<taxonomicName id="4C194D5CFFADB332E1F97D22FED8F98D" baseAuthorityName="REITTER" baseAuthorityYear="1887" box="[187,284,1540,1562]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="gallicus">
<emphasis id="B96DEACDFFADB332E1F97D22FED8F98D" box="[187,284,1540,1562]" italics="true" pageId="2" pageNumber="63">B. gallicus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
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<taxonomicName id="4C194D5CFFACB333E1F37BF1FE49FF79" authorityName="Kahlen &amp; Eckelt &amp; C &amp; C" authorityYear="2024" box="[177,397,214,237]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="cialanciaensis" status="sp. nov.">
<emphasis id="B96DEACDFFACB333E1F37BF1FE49FF79" box="[177,397,214,237]" italics="true" pageId="3" pageNumber="64">Bryaxis cialanciaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57B6FFACB333E0D17BF1FE1AFF79" box="[403,478,214,237]" pageId="3" pageNumber="64" rank="species">nov.sp.</taxonomicNameLabel>
<taxonomicName id="4C194D5CFFACB333E3347BF1FCD7FF79" baseAuthorityName="REITTER" baseAuthorityYear="1887" box="[630,787,214,237]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="gallicus">
<emphasis id="B96DEACDFFACB333E3347BF1FCD7FF79" box="[630,787,214,237]" italics="true" pageId="3" pageNumber="64">Bryaxis gallicus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BA636DFFFACB333E1F37BDBFBE4FDE7" blockId="3.[177,1056,251,715]" pageId="3" pageNumber="64">
Antenna ♁: Antennal segment 1 exactly Antenna ♁: Antennal segment 1 almost square (
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), only moderately square, appearing shorter due to a glossy, slightly asymmetrically humped stronger approach to the antennal pit, on the upper side, this elevation towards distinctly glossy, slightly asymmetrically the center or more towards the inner humped on the upper side, but this edge; Antennal segment 2 almost exactly elevation is more clearly towards the spherical (
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), slightly longer outer edge; Antennal segment 2 longer than wide (
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). Antenna
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: Antennal than wide segment 1 about half longer than wide (L (L
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, W
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mm
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),
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mm
</quantity>
, W
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<geoCoordinate id="EE2D5018FFACB333E0987923FDDFFD8F" box="[474,539,516,539]" degrees="0.057" direction="west" orientation="longitude" pageId="3" pageNumber="64" precision="55" value="-0.057">‐0.057</geoCoordinate>
mm
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; strongly asymmetrical (the outer edge 1.53:1.00) curved, the inner edge almost straight Habitus:
<figureCitation id="13222A5AFFACB333E04E7961FE87FDC9" box="[268,323,582,605]" captionStart="Figs 1-4" captionStartId="2.[163,203,1492,1514]" captionTargetBox="[164,1045,210,1478]" captionTargetId="figure-46@2.[164,1069,210,1478]" captionTargetPageId="2" captionText="Figs 1-4: (1) Bryaxis cialanciaensis nov.sp. Holotype, scale bar 0,5 mm; (2) aedeagus dorsal B. cialanciaensis, scale bar 0,1 mm; (3-4) Habitus (scale bar 0,5 mm) and aedeagus (scale bar 0,1 mm) of B. gallicus." figureDoi="http://doi.org/10.5281/zenodo.14681986" httpUri="https://zenodo.org/record/14681986/files/figure.png" pageId="3" pageNumber="64">Fig. 1</figureCitation>
and drawn out in a narrow bulge towards the rear (
<figureCitation id="13222A5AFFACB333E39A797BFCD1FDE7" box="[728,789,604,627]" captionStart="Figs 1-4" captionStartId="2.[163,203,1492,1514]" captionTargetBox="[164,1045,210,1478]" captionTargetId="figure-46@2.[164,1069,210,1478]" captionTargetPageId="2" captionText="Figs 1-4: (1) Bryaxis cialanciaensis nov.sp. Holotype, scale bar 0,5 mm; (2) aedeagus dorsal B. cialanciaensis, scale bar 0,1 mm; (3-4) Habitus (scale bar 0,5 mm) and aedeagus (scale bar 0,1 mm) of B. gallicus." figureDoi="http://doi.org/10.5281/zenodo.14681986" httpUri="https://zenodo.org/record/14681986/files/figure.png" pageId="3" pageNumber="64">Fig. 3</figureCitation>
,
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). Antenna
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: Antennal
</paragraph>
<paragraph id="8BA636DFFFACB333E1F3794BFEB5FD17" blockId="3.[177,1056,251,715]" box="[177,369,619,643]" pageId="3" pageNumber="64">
Aedeagus ♁:
<figureCitation id="13222A5AFFACB333E079794BFEB5FD17" box="[315,369,620,643]" captionStart="Figs 1-4" captionStartId="2.[163,203,1492,1514]" captionTargetBox="[164,1045,210,1478]" captionTargetId="figure-46@2.[164,1069,210,1478]" captionTargetPageId="2" captionText="Figs 1-4: (1) Bryaxis cialanciaensis nov.sp. Holotype, scale bar 0,5 mm; (2) aedeagus dorsal B. cialanciaensis, scale bar 0,1 mm; (3-4) Habitus (scale bar 0,5 mm) and aedeagus (scale bar 0,1 mm) of B. gallicus." figureDoi="http://doi.org/10.5281/zenodo.14681986" httpUri="https://zenodo.org/record/14681986/files/figure.png" pageId="3" pageNumber="64">Fig. 2</figureCitation>
</paragraph>
<paragraph id="8BA636DFFFACB333E334795EFBDBFD39" blockId="3.[177,1056,251,715]" pageId="3" pageNumber="64">
segment 1 about one‐third longer than wide (L
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mm
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, W
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</paragraph>
<paragraph id="8BA636DFFFACB333E3347993FCD0FD5F" blockId="3.[177,1056,251,715]" box="[630,788,692,715]" pageId="3" pageNumber="64">mm; 1.35:1:00)</paragraph>
<paragraph id="8BA636DFFFACB333E33479FEFCCCFD64" blockId="3.[630,776,729,752]" box="[630,776,729,752]" pageId="3" pageNumber="64">
Habitus:
<figureCitation id="13222A5AFFACB333E39079FEFCCCFD64" box="[722,776,729,752]" captionStart="Figs 1-4" captionStartId="2.[163,203,1492,1514]" captionTargetBox="[164,1045,210,1478]" captionTargetId="figure-46@2.[164,1069,210,1478]" captionTargetPageId="2" captionText="Figs 1-4: (1) Bryaxis cialanciaensis nov.sp. Holotype, scale bar 0,5 mm; (2) aedeagus dorsal B. cialanciaensis, scale bar 0,1 mm; (3-4) Habitus (scale bar 0,5 mm) and aedeagus (scale bar 0,1 mm) of B. gallicus." figureDoi="http://doi.org/10.5281/zenodo.14681986" httpUri="https://zenodo.org/record/14681986/files/figure.png" pageId="3" pageNumber="64">Fig. 3</figureCitation>
</paragraph>
<paragraph id="8BA636DFFFACB333E33479D9FC8BFC81" blockId="3.[630,847,766,789]" box="[630,847,766,789]" pageId="3" pageNumber="64">
Aedeagus ♁:
<figureCitation id="13222A5AFFACB333E24279D9FCFEFC81" box="[768,826,766,789]" captionStart="Figs 1-4" captionStartId="2.[163,203,1492,1514]" captionTargetBox="[164,1045,210,1478]" captionTargetId="figure-46@2.[164,1069,210,1478]" captionTargetPageId="2" captionText="Figs 1-4: (1) Bryaxis cialanciaensis nov.sp. Holotype, scale bar 0,5 mm; (2) aedeagus dorsal B. cialanciaensis, scale bar 0,1 mm; (3-4) Habitus (scale bar 0,5 mm) and aedeagus (scale bar 0,1 mm) of B. gallicus." figureDoi="http://doi.org/10.5281/zenodo.14681986" httpUri="https://zenodo.org/record/14681986/files/figure.png" pageId="3" pageNumber="64">Fig. 4</figureCitation>
,
<figureCitation id="13222A5AFFACB333E20079D9FC8BFC81" box="[834,847,766,789]" captionStart="Figs 5-6" captionStartId="3.[163,203,1450,1472]" captionTargetBox="[188,963,808,1440]" captionTargetId="figure-196@3.[161,1060,808,1440]" captionTargetPageId="3" captionText="Figs 5-6: Drawings of antenna and aedeagus of Bryaxis gallicus from JEANNEL 1950." figureDoi="http://doi.org/10.5281/zenodo.14681988" httpUri="https://zenodo.org/record/14681988/files/figure.png" pageId="3" pageNumber="64">6</figureCitation>
</paragraph>
<caption id="DF666657FFACB333E1E17E8DFC68FA54" ID-DOI="http://doi.org/10.5281/zenodo.14681988" ID-Zenodo-Dep="14681988" box="[163,940,1450,1473]" httpUri="https://zenodo.org/record/14681988/files/figure.png" pageId="3" pageNumber="64" startId="3.[163,203,1450,1472]" targetBox="[188,963,808,1440]" targetPageId="3" targetType="figure">
<paragraph id="8BA636DFFFACB333E1E17E8DFC68FA54" blockId="3.[163,940,1450,1473]" box="[163,940,1450,1473]" pageId="3" pageNumber="64">
<emphasis id="B96DEACDFFACB333E1E17E8DFF2BFA2B" bold="true" box="[163,239,1450,1472]" pageId="3" pageNumber="64">Figs 5-6</emphasis>
: Drawings of antenna and aedeagus of
<taxonomicName id="4C194D5CFFACB333E3177E8CFD2CFA54" baseAuthorityName="REITTER" baseAuthorityYear="1887" box="[597,744,1451,1472]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="gallicus">
<emphasis id="B96DEACDFFACB333E3177E8CFD2CFA54" box="[597,744,1451,1472]" italics="true" pageId="3" pageNumber="64">Bryaxis gallicus</emphasis>
</taxonomicName>
from
<bibRefCitation id="EF884B2EFFACB333E25D7E8CFC6CFA54" author="JEANNEL R." box="[799,936,1450,1472]" pageId="3" pageNumber="64" refId="ref1776" refString="JEANNEL R. (1950): Coleopteres Pselaphides. - Faune de France 53: 328 pp." type="book" year="1950">
<smallCapsWord id="8D40A003FFACB333E25D7E8CFCB7FA2B" baselines="1467,1467" box="[799,883,1451,1472]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Jeannel" pageId="3" pageNumber="64">JEANNEL</smallCapsWord>
1950
</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="8BA636DFFFABB334E1E17BF7FBEAFEA9" blockId="4.[163,1070,207,318]" pageId="4" pageNumber="65">
D i s t r i b u t i o n a n d h a b i t a t: The new species has been found exclusively in the northern Cottian Alps, specifically in the Conca Cialancia Nature Park (
<figureCitation id="13222A5AFFABB334E2F97BCAFC37FE96" box="[955,1011,237,259]" captionStart="Fig" captionStartId="4.[163,196,984,1006]" captionTargetBox="[170,1059,349,960]" captionTargetId="figure-174@4.[162,1070,339,971]" captionTargetPageId="4" captionText="Fig. 7: Distribution map of B. cialanciaensis." figureDoi="http://doi.org/10.5281/zenodo.14681990" httpUri="https://zenodo.org/record/14681990/files/figure.png" pageId="4" pageNumber="65">Fig 7</figureCitation>
). The
<typeStatus id="54A2887DFFABB334E1E17A2DFF38FEB4" box="[163,252,266,288]" pageId="4" pageNumber="65" type="holotype">holotype</typeStatus>
and
<typeStatus id="54A2887DFFABB334E06D7A2CFE54FEB4" box="[303,400,267,288]" pageId="4" pageNumber="65" type="paratype">paratypes</typeStatus>
were collected by sifting
<emphasis id="B96DEACDFFABB334E3DF7A2DFD14FE8B" box="[669,720,266,287]" italics="true" pageId="4" pageNumber="65">Salix</emphasis>
and
<emphasis id="B96DEACDFFABB334E2467A2DFC5EFE8B" box="[772,922,266,287]" italics="true" pageId="4" pageNumber="65">Rhododendron</emphasis>
litter between rock crevices (
<figureCitation id="13222A5AFFABB334E0777A0FFE49FEA9" box="[309,397,296,318]" captionStart="Figs 8-9" captionStartId="5.[163,203,1451,1473]" captionTargetBox="[166,1069,206,1427]" captionTargetId="figure-23@5.[166,1069,206,1427]" captionTargetPageId="5" captionText="Figs 8-9: Habitat pictures of B. cialanciaensis. (8) Punta Freidour W-Side and (9) Lago Lauson." figureDoi="http://doi.org/10.5281/zenodo.14681992" httpUri="https://zenodo.org/record/14681992/files/figure.png" pageId="4" pageNumber="65">Figs 8-9</figureCitation>
). The species is currently known to be a local endemic of this area.
</paragraph>
</subSubSection>
</treatment>
</document>
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