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<document id="654ABDC9CCD5F74937EF77C94B09313B" ID-DOI="10.5852/ejt.2024.926.2469" ID-GBIF-Dataset="4511b4bd-f026-49d3-9d4d-a39875e75cc5" ID-ISSN="2118-9773" ID-Zenodo-Dep="10847806" ID-ZooBank="A00D7669-2151-46AE-A066-9AF4D0F2BEEE" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1711011351298" checkinUser="plazi" docAuthor="Jochum, Adrienne, Michalik, Peter, Inäbnit, Thomas, Kneubühler, Jeannette, Slapnik, Rajko, Vrabec, Marko, Schilthuizen, Menno &amp; Ruthensteiner, Bernhard" docDate="2024" docId="03B0879DFF9BFFE3FE6FD18CFEE1FC7D" docLanguage="en" docName="EJT.2024.926.1-62.pdf" docOrigin="European Journal of Taxonomy 926" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2469/10973" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zospeum tumidum Jochum, Schilthuizen &amp; Ruthensteiner 2024, sp. nov." docType="treatment" docUuid="1F6F1AB3-137B-4AA9-ABB8-46E0AF777F8C" docUuidSource="ZooBank" docVersion="10" lastPageNumber="36" masterDocId="FF89FFE5FFBAFFC0FFFFD700FFD4FFD4" masterDocTitle="3 D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="34" updateTime="1722640713706" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9CB3DF45C980AEBD99FB29485CE3A105">3 D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae)</mods:title>
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<mods:namePart id="1F68B3380A3193C7221A076A0B5531EE">Jochum, Adrienne</mods:namePart>
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<mods:affiliation id="9AD2459FAD9C6E325FD3453A794B2076">Senckenberg Research Institute and Natural History Museum, Senckenberganlage 25, D- 60325 Frankfurt / M., Germany.</mods:affiliation>
<mods:nameIdentifier id="19DD144845BD7CDFCD79C509F875FF35" type="email">nne.jochum@senckenberg.de</mods:nameIdentifier>
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<mods:namePart id="9220C87EABEFE6ACB7D7E570D0A84067">Michalik, Peter</mods:namePart>
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<mods:affiliation id="8274B86DA77DE0E3A2415D9408429082">Zoological Institute and Museum, University of Greifswald, Loitzer Strasse 26, D- 17489 Greifswald, Germany.</mods:affiliation>
<mods:nameIdentifier id="1FF13CAAA30281DAF9092E5D4E40F292" type="email">michalik@uni-greifswald.de</mods:nameIdentifier>
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<mods:namePart id="605455A530F21251862420BCD5984D66">Inäbnit, Thomas</mods:namePart>
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<mods:affiliation id="E95E6F85A11D2720157FBE694ED6222F">Institute for Biochemistry &amp; Biology, University of Potsdam, Karl-Liebknecht-Strasse 24 - 25, House 26, D- 14476 Potsdam, Germany.</mods:affiliation>
<mods:nameIdentifier id="36F03FFC60CE651DBD681B85B19796C1" type="email">inaebnit.thomas@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FB2A2D8731E0911CF004094E60DDB968">Kneubühler, Jeannette</mods:namePart>
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<mods:affiliation id="51180DC489F5F9027015AF3A2ADB7317">Naturhistorisches Museum Bern, Bernastrasse 15, CH- 3005 Bern, Switzerland. &amp; Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, CH- 3012 Bern, Switzerland.</mods:affiliation>
<mods:nameIdentifier id="8A3096EAC33FBC318D80D2FC5FD0A281" type="email">jeannette.kneubuehler@nmbe.ch</mods:nameIdentifier>
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<mods:namePart id="72C258F836EE30CBFFE443C847A7D747">Slapnik, Rajko</mods:namePart>
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<mods:affiliation id="BB428ACC4B139F5CA4AE80FC929A6B8F">ZOSPEUM, Molluscs, Cave &amp; Karst Biological Consulting, Drnovškova pot 2, Mekinje, SI 1240 Kamnik, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="482DA774F5A7EDC31A56F08EE612CBED" type="email">rajko.slapnik@gmail.com</mods:nameIdentifier>
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<mods:namePart id="3CD5E0438B48E47976422067297901F9">Vrabec, Marko</mods:namePart>
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<mods:affiliation id="CE9DB4F918FA6BB5A849EA29AF023FE6">Department of Geology, Faculty of Natural Sciences and Engineering, Aškerčeva 12, University of Ljubljana, 1000 Ljubljana, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="AE9BFA41A756AFEFADB1D3A1ABA845CF" type="email">marko.vrabec@geo.ntf.uni-lj.si</mods:nameIdentifier>
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<mods:namePart id="3CF3AD539533DDBA68CE2D54FB8E4DD0">Schilthuizen, Menno</mods:namePart>
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<mods:affiliation id="01C7A7612504B4E02B535ABC886E2DE3">Naturalis Biodiversity Center and Leiden University, Vondellaan 55, NL- 2332 AA Leiden, The Netherlands.</mods:affiliation>
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<mods:namePart id="8B25251F95E92DD01ED2452D29C66F8F">Ruthensteiner, Bernhard</mods:namePart>
<mods:nameIdentifier id="247D2F2C0139671990AECBEF8269057F" type="LSID">9A8A56B0-19FA-42AB-A85E-4FCA98BF4A6F</mods:nameIdentifier>
<mods:affiliation id="3C057FE670C2C28F9DE28E74E372D240">SNSB - Zoologische Staatssammlung München (ZSM), 81247 München, Germany.</mods:affiliation>
<mods:nameIdentifier id="0D6FA527804EE46D4C0C360E3019C932" type="email">ruthensteiner@snsb.de</mods:nameIdentifier>
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<mods:title id="3E21DA175695C51DE3D211F019321CE8">European Journal of Taxonomy</mods:title>
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<mods:part id="C0D532DC93677624CF5D0282E81E2B71">
<mods:date id="B4C9B49F384177A87390AB14814A6B51">2024</mods:date>
<mods:detail id="ED2B4F91E47CE11CC50357B0AB015A90" type="pubDate">
<mods:number id="36C039A1B45A675757CC55AB9D8166A5">2024-03-20</mods:number>
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<mods:detail id="85FA7B3EF257C0ACB81686329E72673D" type="volume">
<mods:number id="48EE1D5BB8B79582369F5621B40C90BF">926</mods:number>
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<mods:start id="4B9DD8C8187165971EED21A25E5D93E3">1</mods:start>
<mods:end id="AAB7931A5221C8422FB9D77345F82F1C">62</mods:end>
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<mods:url id="F196F2F3601BD0326F25F90342CD725A">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2469/10973</mods:url>
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<mods:classification id="714988259CDE848DA8B06BC4CA110B3B">journal article</mods:classification>
<mods:identifier id="2FC15683A512B7868098E702CC19DE51" type="DOI">10.5852/ejt.2024.926.2469</mods:identifier>
<mods:identifier id="DFCFE1B77C02D8486F99E191B2B00AEF" type="GBIF-Dataset">4511b4bd-f026-49d3-9d4d-a39875e75cc5</mods:identifier>
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<treatment id="03B0879DFF9BFFE3FE6FD18CFEE1FC7D" ID-DOI="http://doi.org/10.5281/zenodo.10847836" ID-GBIF-Taxon="224067515" ID-Zenodo-Dep="10847836" LSID="urn:lsid:zoobank.org:act:1F6F1AB3-137B-4AA9-ABB8-46E0AF777F8C" httpUri="http://treatment.plazi.org/id/03B0879DFF9BFFE3FE6FD18CFEE1FC7D" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
<subSubSection id="C3036500FF9BFFE1FE6FD18CFB70F972" box="[400,1188,1675,1702]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BA6368BFF9BFFE1FE6FD18CFB70F972" blockId="33.[359,1228,1675,1772]" box="[400,1188,1675,1702]" pageId="33" pageNumber="34">
<heading id="D0EE81E7FF9BFFE1FE6FD18CFB70F972" box="[400,1188,1675,1702]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="34" reason="2">
<taxonomicName id="4C194D08FF9BFFE1FE6FD18CFB96F971" ID-CoL="C8NTG" authority="Jochum, Schilthuizen &amp; Ruthensteiner" authorityName="Jochum, Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[400,1090,1675,1702]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="tumidum" status="sp. nov.">
<emphasis id="B96DEA99FF9BFFE1FE6FD18CFDA1F972" bold="true" box="[400,629,1675,1702]" italics="true" pageId="33" pageNumber="34">Zospeum tumidum</emphasis>
Jochum, Schilthuizen &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF9BFFE1FBB6D18CFB70F972" box="[1097,1188,1676,1702]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF9BFFE1FE98D1AFFC97F938" pageId="33" pageNumber="34" type="description">
<paragraph id="8BA6368BFF9BFFE1FE98D1AFFB18F91D" blockId="33.[359,1228,1675,1772]" box="[359,1228,1711,1737]" pageId="33" pageNumber="34">
<uri id="FF883A89FF9BFFE1FE98D1AFFB18F91D" box="[359,1228,1711,1737]" pageId="33" pageNumber="34">
urn:lsid:zoobank.org:act:
<uuid id="FFBF0C5EFF9BFFE1FD72D1AFFB18F91D" box="[653,1228,1711,1737]" pageId="33" pageNumber="34">1F6F1AB3-137B-4AA9-ABB8-46E0AF777F8C</uuid>
</uri>
</paragraph>
<paragraph id="8BA6368BFF9BFFE1FD0FD1D2FC97F938" blockId="33.[359,1228,1675,1772]" box="[752,835,1746,1772]" pageId="33" pageNumber="34">
<figureCitation id="13222A0EFF9BFFE1FD0FD1D2FC97F938" box="[752,835,1746,1772]" captionStart="Fig" captionStartId="34.[192,235,1857,1883]" captionTargetBox="[189,1398,881,1817]" captionTargetId="figure-207@34.[189,1398,881,1817]" captionTargetPageId="34" captionText="Fig. 10. Zospeum tumidum Jochum, Schilthuizen &amp; Ruthensteiner sp. nov., holotype (NMBE 572615). A. Light microscopic images, showing aperture and dorsal views and TxEx Collection label. B. 3D visualizations of Micro-CT data, apical view. C. Ventral view showing deep umbilical depression. D. Aperture view.E. Aperture facing right view showing tapered columella and lamellar bulge. F. Dorsal view. G. Aperture facing left view." figureDoi="http://doi.org/10.5281/zenodo.10847832" httpUri="https://zenodo.org/record/10847832/files/figure.png" pageId="33" pageNumber="34">Fig. 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF9BFFE1FF42D013FE55F811" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BA6368BFF9BFFE1FF42D013FEE2F8F9" blockId="33.[189,310,1811,1837]" box="[189,310,1811,1837]" pageId="33" pageNumber="34">
<heading id="D0EE81E7FF9BFFE1FF42D013FEE2F8F9" bold="true" box="[189,310,1811,1837]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B96DEA99FF9BFFE1FF42D013FEE2F8F9" bold="true" box="[189,310,1811,1837]" pageId="33" pageNumber="34">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF9BFFE1FF42D042FE55F811" blockId="33.[189,1399,1856,1989]" pageId="33" pageNumber="34">
Shell ca
<quantity id="4CE19B6EFF9BFFE1FEDBD042FE56F888" box="[292,386,1858,1885]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="34" unit="mm" value="1.5">1.5 mm</quantity>
, turbinate-conical with 5–5¼ regularly coiled, irregularly formed convex whorls; last coiling of shell twists a fraction beyond the frontal plane ca ⅛ the width of penultimate whorl; columella centrally aligned and moderately fat bearing a lamellar bulge rather than a fine lamella; umbilical depression deep.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF98FFE2FF42D609FBCBFE85" box="[189,1055,265,338]" pageId="34" pageNumber="35" type="etymology">
<paragraph id="8BA6368BFF98FFE2FF42D609FE90FEF7" blockId="34.[189,324,265,291]" box="[189,324,265,291]" pageId="34" pageNumber="35">
<heading id="D0EE81E7FF98FFE2FF42D609FE90FEF7" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B96DEA99FF98FFE2FF42D609FE90FEF7" bold="true" box="[189,324,265,291]" pageId="34" pageNumber="35">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF98FFE2FF42D637FBCBFE85" blockId="34.[189,1055,311,338]" box="[189,1055,311,338]" pageId="34" pageNumber="35">The species is named after the squat and wide, turbinate form of the shell.</paragraph>
</subSubSection>
<subSubSection id="C3036500FF98FFE2FF42D679FAB7FDDC" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8BA6368BFF98FFE2FF42D679FEB8FE46" blockId="34.[189,364,376,403]" box="[189,364,376,403]" pageId="34" pageNumber="35">
<heading id="D0EE81E7FF98FFE2FF42D679FEB8FE46" bold="true" box="[189,364,376,403]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B96DEA99FF98FFE2FF42D679FEB8FE46" bold="true" box="[189,364,376,403]" pageId="34" pageNumber="35">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF98FFE2FF15D6A7FE88FE15" blockId="34.[189,1398,423,520]" box="[234,348,423,449]" pageId="34" pageNumber="35">
<emphasis id="B96DEA99FF98FFE2FF15D6A7FE88FE15" bold="true" box="[234,348,423,449]" pageId="34" pageNumber="35">
<typeStatus id="54A28829FF98FFE2FF15D6A7FE88FE15" box="[234,348,423,449]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA6368BFF98FFE2FF42D6CAFAB7FDDC" blockId="34.[189,1398,423,520]" pageId="34" pageNumber="35">
<materialsCitation id="3B713CD6FF98FFE2FF42D6CAFAB7FDDC" ID-GBIF-Occurrence="4598261327" collectingDate="2018-07-01" collectionCode="NMBE" collectorName="M. Schilthuizen &amp; I. Njunjic" country="Serbia" county="Dokova" latitude="43.0315" location="Piva Region" longLatPrecision="6" longitude="18.8104" municipality="Bajovo Polje" pageId="34" pageNumber="35" specimenCode="NMBE 572615" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F30E761BFF98FFE2FF42D6CAFE5EFE30" box="[189,394,458,484]" name="Serbia" pageId="34" pageNumber="35">MONTENEGRO</collectingCountry>
•
<location id="8EC66050FF98FFE2FE61D6CAFDE4FE30" LSID="urn:lsid:plazi:treatment:03B0879DFF9BFFE3FE6FD18CFEE1FC7D:8EC66050FF98FFE2FE61D6CAFDE4FE30" box="[414,560,458,484]" country="Serbia" county="Dokova" latitude="43.0315" longLatPrecision="6" longitude="18.8104" municipality="Bajovo Polje" name="Piva Region" pageId="34" pageNumber="35">Piva Region</location>
,
<collectingMunicipality id="6BC2ACF1FF98FFE2FDC6D6CAFD05FE30" box="[569,721,458,484]" pageId="34" pageNumber="35">Bajovo Polje</collectingMunicipality>
,
<location id="8EC66050FF98FFE2FD24D6CAFC59FE30" LSID="urn:lsid:plazi:treatment:03B0879DFF9BFFE3FE6FD18CFEE1FC7D:8EC66050FF98FFE2FD24D6CAFC59FE30" box="[731,909,458,484]" country="Serbia" county="Dokova" latitude="43.0315" longLatPrecision="6" longitude="18.8104" municipality="Bajovo Polje" name="Dokova pecina" pageId="34" pageNumber="35">Đokova pećina</location>
, at cave entrance;
<geoCoordinate id="EE2D504CFF98FFE2FBA4D6CAFB37FE31" box="[1115,1251,458,485]" degrees="43.0315" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="5" value="43.0315">43.0315° N</geoCoordinate>
,
<geoCoordinate id="EE2D504CFF98FFE2FB13D6CAFAA5FE31" box="[1260,1393,458,485]" degrees="18.8104" direction="east" orientation="longitude" pageId="34" pageNumber="35" precision="5" value="18.8104">18.8104° E</geoCoordinate>
;
<date id="FFA7104BFF98FFE2FF42D6EEFE96FDDC" box="[189,322,493,520]" pageId="34" pageNumber="35" value="2018-07-01">
<collectingDate id="EFE3E9A3FF98FFE2FF42D6EEFE96FDDC" box="[189,322,493,520]" pageId="34" pageNumber="35" value="2018-07-01">1 Jul. 2018</collectingDate>
</date>
;
<collectorName id="26EC535DFF98FFE2FEB1D6EEFDDEFDD3" box="[334,522,493,520]" pageId="34" pageNumber="35">M. Schilthuizen</collectorName>
and
<collectorName id="26EC535DFF98FFE2FDBDD6EEFD67FDD3" box="[578,691,493,520]" pageId="34" pageNumber="35">I. Njunjić</collectorName>
leg; at cave entrance;
<specimenCode id="DBBF9EF0FF98FFE2FC45D6EEFBBAFDDC" box="[954,1134,493,520]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="34" pageNumber="35">NMBE 572615</specimenCode>
(ex TxEx-DU0025).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF98FFE3FF42D52FFC5AFDB4" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
<paragraph id="8BA6368BFF98FFE2FF42D52FFE9AFD9D" blockId="34.[189,334,559,585]" box="[189,334,559,585]" pageId="34" pageNumber="35">
<heading id="D0EE81E7FF98FFE2FF42D52FFE9AFD9D" bold="true" box="[189,334,559,585]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B96DEA99FF98FFE2FF42D52FFE9AFD9D" bold="true" box="[189,334,559,585]" pageId="34" pageNumber="35">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF98FFE2FF42D55EFEB8FD4E" blockId="34.[188,1398,605,667]" pageId="34" pageNumber="35">
<smallCapsWord id="8D40A057FF98FFE2FF42D55EFEBAFDA2" baselines="625,625" box="[189,366,606,632]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="34" pageNumber="35">MEASUREMENTS</smallCapsWord>
.
<typeStatus id="54A28829FF98FFE2FE83D55DFE3AFDA3" box="[380,494,605,631]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
: sh:
<quantity id="4CE19B6EFF98FFE2FDDED55DFD4EFDAC" box="[545,666,605,632]" metricMagnitude="-3" metricUnit="m" metricValue="1.537" pageId="34" pageNumber="35" unit="mm" value="1.537">1.537 mm</quantity>
; sw:
<quantity id="4CE19B6EFF98FFE2FD2AD55DFC9AFDAC" box="[725,846,605,632]" metricMagnitude="-3" metricUnit="m" metricValue="1.177" pageId="34" pageNumber="35" unit="mm" value="1.177">1.177 mm</quantity>
; ah:
<quantity id="4CE19B6EFF98FFE2FC7BD55DFC29FDAC" box="[900,1021,605,632]" metricMagnitude="-4" metricUnit="m" metricValue="7.31" pageId="34" pageNumber="35" unit="mm" value="0.731">0.731 mm</quantity>
; aw:
<quantity id="4CE19B6EFF98FFE2FBC6D55DFB66FDAC" box="[1081,1202,605,632]" metricMagnitude="-4" metricUnit="m" metricValue="7.119999999999999" pageId="34" pageNumber="35" unit="mm" value="0.712">0.712 mm</quantity>
; hlw:
<quantity id="4CE19B6EFF98FFE2FB07D55DFAA6FDAC" box="[1272,1394,605,632]" metricMagnitude="-3" metricUnit="m" metricValue="1.028" pageId="34" pageNumber="35" unit="mm" value="1.028">1.028 mm</quantity>
; SA: 74.75 deg.
</paragraph>
<paragraph id="8BA6368BFF98FFE3FF42D5C2FC5AFDB4" blockId="34.[189,1399,705,803]" lastBlockId="35.[189,1399,265,608]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
Shell ca
<quantity id="4CE19B6EFF98FFE2FED2D5C2FE5BFD09" box="[301,399,706,733]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="35" unit="mm" value="1.5">1.5 mm</quantity>
, turbinate-conical with 5–5¼ irregular, flatly convex whorls; shell comparatively large; milky translucent; suture shallow with irregular indentations at the junction of the suture and the succeeding whorl growing into course growth lines; teleoconch sculpture of blunt irregular growth lines interspersed with fine, weak thread-like striations; distinct, fine, thread-like axial ribbing extends for a distance immediately behind palatal-basal lip; last whorl large and tumid, encompassing ca ⅔ of shell height; last coiling twists a fraction beyond the frontal plane ca ⅛ the width of body whorl; suture shallow; aperture reniform, somewhat higher than wide; peristome generally thin especially at upper palatal side, slightly expanded; weak notch present at upper junction with parietal shield, columellar side oblique and thickly callused, columellar and palatal-basal lip narrowly reflected; parietal shield well defined, straight, long, angular with course growth lines crossing into it; columella thick and centrally aligned, apically tapered (aperture facing right) with a lamellar bulge and no defined lamella; umbilical depression deep, callused and puckered; columellar side of peristome arches high over and oblique to umbilical depression, alignment of last ¼ whorl not compact.
</paragraph>
</subSubSection>
<caption id="DF666603FF98FFE2FF3FD041FD82F83C" ID-DOI="http://doi.org/10.5281/zenodo.10847832" ID-Zenodo-Dep="10847832" httpUri="https://zenodo.org/record/10847832/files/figure.png" pageId="34" pageNumber="35" startId="34.[192,235,1857,1883]" targetBox="[189,1398,881,1817]" targetPageId="34">
<paragraph id="8BA6368BFF98FFE2FF3FD041FD82F83C" blockId="34.[189,1401,1857,2025]" pageId="34" pageNumber="35">
<emphasis id="B96DEA99FF98FFE2FF3FD041FEC9F88F" bold="true" box="[192,285,1857,1883]" pageId="34" pageNumber="35">Fig. 10.</emphasis>
<taxonomicName id="4C194D08FF98FFE2FEDBD042FC1FF88F" authority="Jochum, Schilthuizen &amp; Ruthensteiner" authorityName="Jochum, Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[292,971,1857,1884]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="tumidum" status="sp. nov.">
<emphasis id="B96DEA99FF98FFE2FEDBD042FE2AF88F" box="[292,510,1857,1883]" italics="true" pageId="34" pageNumber="35">Zospeum tumidum</emphasis>
Jochum, Schilthuizen &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF98FFE2FC2DD042FBFBF888" box="[978,1071,1858,1884]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (NMBE 572615).
<emphasis id="B96DEA99FF98FFE2FF40D064FF00F8AA" bold="true" box="[191,212,1892,1918]" pageId="34" pageNumber="35">A</emphasis>
. Light microscopic images, showing aperture and dorsal views and TxEx Collection label.
<emphasis id="B96DEA99FF98FFE2FAD0D065FA97F8AB" bold="true" box="[1327,1347,1893,1919]" pageId="34" pageNumber="35">B</emphasis>
. 3D visualizations of Micro-CT data, apical view.
<emphasis id="B96DEA99FF98FFE2FD09D088FCDFF876" bold="true" box="[758,779,1928,1954]" pageId="34" pageNumber="35">C</emphasis>
. Ventral view showing deep umbilical depression.
<emphasis id="B96DEA99FF98FFE2FF40D0ABFF00F811" bold="true" box="[191,212,1963,1989]" pageId="34" pageNumber="35">D</emphasis>
. Aperture view.
<emphasis id="B96DEA99FF98FFE2FE71D0ABFE76F811" bold="true" box="[398,418,1963,1989]" pageId="34" pageNumber="35">E</emphasis>
. Aperture facing right view showing tapered columella and lamellar bulge.
<emphasis id="B96DEA99FF98FFE2FAF5D0ABFAC8F811" bold="true" box="[1290,1308,1963,1989]" pageId="34" pageNumber="35">F</emphasis>
. Dorsal view.
<emphasis id="B96DEA99FF98FFE2FEFAD0CEFEC8F83C" bold="true" box="[261,284,1998,2024]" pageId="34" pageNumber="35">G</emphasis>
. Aperture facing left view.
</paragraph>
</caption>
<subSubSection id="C3036500FF99FFE3FF42D587FAA3FD04" box="[189,1399,647,720]" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BA6368BFF99FFE3FF42D587FE82FD75" blockId="35.[189,342,647,673]" box="[189,342,647,673]" pageId="35" pageNumber="36">
<heading id="D0EE81E7FF99FFE3FF42D587FE82FD75" bold="true" box="[189,342,647,673]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B96DEA99FF99FFE3FF42D587FE82FD75" bold="true" box="[189,342,647,673]" pageId="35" pageNumber="36">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF99FFE3FF42D5B6FAA3FD04" blockId="35.[189,1399,694,720]" box="[189,1399,694,720]" pageId="35" pageNumber="36">
This species is only known from its
<typeStatus id="54A28829FF99FFE3FDA8D5B6FD5EFD04" box="[599,650,694,720]" pageId="35" pageNumber="36">type</typeStatus>
locality cave, Đokova pećina in the Bajovo Polje of
<collectingCountry id="F30E761BFF99FFE3FB1FD5B6FAA7FD04" box="[1248,1395,694,720]" name="Serbia" pageId="35" pageNumber="36">Montenegro</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF99FFE3FF42D5F7FEE1FC7D" pageId="35" pageNumber="36" type="discussion">
<paragraph id="8BA6368BFF99FFE3FF42D5F7FEFBFCC5" blockId="35.[189,303,759,785]" box="[189,303,759,785]" pageId="35" pageNumber="36">
<heading id="D0EE81E7FF99FFE3FF42D5F7FEFBFCC5" bold="true" box="[189,303,759,785]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B96DEA99FF99FFE3FF42D5F7FEFBFCC5" bold="true" box="[189,303,759,785]" pageId="35" pageNumber="36">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF99FFE3FF42D426FEE1FC7D" blockId="35.[188,1399,806,937]" pageId="35" pageNumber="36">
The species is the only Southern Balkan species to possess a thick lamellar bulge. It has the widest shell with the greatest aperture height and aperture width as well as the highest last whorl and greatest spire angle of all 13 species (and one morphospecies
<taxonomicName id="4C194D08FF99FFE3FD08D46DFC77FC52" box="[759,931,876,903]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="undefined-2">
<emphasis id="B96DEA99FF99FFE3FD08D46DFCB6FC52" box="[759,866,877,902]" italics="true" pageId="35" pageNumber="36">Zospeum</emphasis>
sp. 2
</taxonomicName>
) (encompassing 31 shells) measured in this study.
</paragraph>
</subSubSection>
</treatment>
</document>
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