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<document id="FF71A03A9A6FBFC36660719CFDADE19E" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFD7E52450C6831B67FEF855" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) hastatus Fauvel 1904" docType="treatment" docVersion="11" lastPageNumber="139" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="139" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="2D7022345339B5D76BA89A3BB9F9C10D">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="112A65F375697C9EF5979E7D38DD1D04">Assing, Volker</mods:namePart>
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<mods:title id="8557836D4941636D1240EDDAB834FAAB">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="46F5AA9AED161F3526EC71488F894B5B">2017</mods:date>
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<mods:number id="07B05E2B112B0B134EB6EC6F7EE4F8E5">2017-06-30</mods:number>
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<treatment id="03A787BAFFD7E52450C6831B67FEF855" ID-DOI="http://doi.org/10.5281/zenodo.5888511" ID-GBIF-Taxon="192498172" ID-Zenodo-Dep="5888511" LSID="urn:lsid:plazi:treatment:03A787BAFFD7E52450C6831B67FEF855" httpUri="http://treatment.plazi.org/id/03A787BAFFD7E52450C6831B67FEF855" lastPageNumber="139" pageId="22" pageNumber="139">
<subSubSection id="C3146527FFD7E52450C6831B6159FB9A" box="[227,594,1043,1070]" pageId="22" pageNumber="139" type="nomenclature">
<paragraph id="8BB136ACFFD7E52450C6831B6159FB9A" blockId="22.[227,594,1043,1103]" box="[227,594,1043,1070]" pageId="22" pageNumber="139">
<heading id="D0F981C0FFD7E52450C6831B6159FB9A" box="[227,594,1043,1070]" centered="true" fontSize="12" level="2" pageId="22" pageNumber="139" reason="2">
<taxonomicName id="4C0E4D2FFFD7E52450C6831B6159FB9A" ID-CoL="5DDCS" authority="FAUVEL, 1904" authorityName="Fauvel" authorityYear="1904" box="[227,594,1043,1070]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="139" phylum="Arthropoda" rank="species" species="hastatus" subGenus="Zyras">
<emphasis id="B97AEABEFFD7E52450C6831B6212FB9A" box="[227,281,1043,1070]" italics="true" pageId="22" pageNumber="139">Zyras</emphasis>
(
<emphasis id="B97AEABEFFD7E5245102831B6256FB9A" box="[295,349,1043,1070]" italics="true" pageId="22" pageNumber="139">Zyras</emphasis>
)
<emphasis id="B97AEABEFFD7E524514F831B62C8FB9A" box="[362,451,1043,1070]" italics="true" pageId="22" pageNumber="139">hastatus</emphasis>
<bibRefCitation id="EF9F4B5DFFD7E52451EF831B6159FB9A" author="FAUVEL, A." box="[458,594,1043,1070]" pageId="22" pageNumber="139" pagination="43 - 70" refId="ref57899" refString="FAUVEL, A. 1904: Staphylinides de l'Hindoustan et de la Birmanie. - Revue d'Entomologie 23: 43 - 70." type="journal article" year="1904">FAUVEL, 1904</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD7E52450C68333629EFBFB" box="[227,405,1083,1103]" pageId="22" pageNumber="139" type="description">
<paragraph id="8BB136ACFFD7E52450C68333629EFBFB" blockId="22.[227,594,1043,1103]" box="[227,405,1083,1103]" pageId="22" pageNumber="139">
(
<figureCitation id="13352A29FFD7E52450C28333622BFBFB" box="[231,288,1083,1103]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="22" pageNumber="139">Figs 16</figureCitation>
,
<figureCitation id="13352A29FFD7E524510283336237FBFB" box="[295,316,1083,1103]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="22" pageNumber="139">54</figureCitation>
,
<figureCitation id="13352A29FFD7E524516683336253FBFB" box="[323,344,1083,1103]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="22" pageNumber="139">90</figureCitation>
,
<figureCitation id="13352A29FFD7E524517A8333629AFBFB" box="[351,401,1083,1103]" captionStart="Map 7" captionStartId="67.[151,196,1246,1267]" captionTargetBox="[351,1198,232,1238]" captionTargetId="figure-79@67.[351,1198,232,1238]" captionTargetPageId="67" captionText="Map 7: Revised distributions of Zyras gardneri (black circles), Z. hirsutiventris (white diamonds), Z. parageminus (black triangles), Z. hastatus (white triangle), Z. hirtus (white circles), and Z. latilobatus (star) in India and adjacent regions." figureDoi="http://doi.org/10.5281/zenodo.5742413" httpUri="https://zenodo.org/record/5742413/files/figure.png" pageId="22" pageNumber="139">Map 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD7E5245098836462FEFB35" box="[189,501,1131,1153]" pageId="22" pageNumber="139" type="reference_group">
<paragraph id="8BB136ACFFD7E5245098836462FEFB35" blockId="22.[189,501,1131,1153]" box="[189,501,1131,1153]" pageId="22" pageNumber="139">
<treatmentCitationGroup id="AB1E1182FFD7E5245098836462FEFB35" box="[189,501,1131,1153]" pageId="22" pageNumber="139">
<treatmentCitation id="0AAF10BDFFD7E5245098836462FBFB35" author="FAUVEL, A." box="[189,496,1131,1153]" page="64" pageId="22" pageNumber="139" year="1904">
<taxonomicName id="4C0E4D2FFFD7E5245098836462FBFB35" ID-CoL="5DDCS" authority="FAUVEL, 1904: 64" authorityName="Fauvel" authorityPageNumber="64" authorityYear="1904" box="[189,496,1131,1153]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="139" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B97AEABEFFD7E52450988364624EFB34" box="[189,325,1131,1153]" italics="true" pageId="22" pageNumber="139">Zyras hastatus</emphasis>
<bibRefCitation id="EF9F4B5DFFD7E524516F836362FBFB35" author="FAUVEL, A." box="[330,496,1131,1153]" pageId="22" pageNumber="139" pagination="43 - 70" refId="ref57899" refString="FAUVEL, A. 1904: Staphylinides de l'Hindoustan et de la Birmanie. - Revue d'Entomologie 23: 43 - 70." type="journal article" year="1904">FAUVEL, 1904: 64</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD7E524509883A16261FAD5" pageId="22" pageNumber="139" type="materials_examined">
<paragraph id="8BB136ACFFD7E524509883A16261FAD5" blockId="22.[189,794,1193,1377]" pageId="22" pageNumber="139">
<emphasis id="B97AEABEFFD7E524509883A162DCFB75" bold="true" box="[189,471,1193,1217]" pageId="22" pageNumber="139">Type material examined</emphasis>
:
<materialsCitation id="3B663CF1FFD7E52451C983A1614FFAB5" ID-GBIF-Occurrence="3450162320" collectionCode="IRSNB" collectorName="V. Assing" location="Belgaum /" municipality="Belgaum /" pageId="22" pageNumber="139" specimenCount="1" typeStatus="syntype">
<typeStatus id="54B5880EFFD7E52451C983A16146FB75" box="[492,589,1193,1217]" pageId="22" pageNumber="139" type="syntype">Syntypes</typeStatus>
:
<specimenCount id="9D08FD25FFD7E524524583A1618EFB75" box="[608,645,1193,1217]" pageId="22" pageNumber="139">1 ♀</specimenCount>
: “
<collectingMunicipality id="6BD5ACD6FFD7E524528683A16011FB75" box="[675,794,1193,1217]" pageId="22" pageNumber="139">Belgaum /</collectingMunicipality>
<emphasis id="B97AEABEFFD7E524509883C16201FB55" box="[189,266,1225,1249]" italics="true" pageId="22" pageNumber="139">hastata</emphasis>
Fvl. / Coll. I.R.Sc.N.B. / Syntypus
<taxonomicName id="4C0E4D2FFFD7E52452A483C2620FFAB5" authority="Fauvel" authorityName="Fauvel" authorityYear="1904" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="139" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B97AEABEFFD7E52452A483C26011FB55" box="[641,794,1225,1249]" italics="true" pageId="22" pageNumber="139">Zyras hastatus</emphasis>
Fauvel
</taxonomicName>
, rev.
<collectorName id="26FB537AFFD7E524511283E26291FAB5" box="[311,410,1257,1281]" pageId="22" pageNumber="139">V. Assing</collectorName>
2016” (
<collectionCode id="ED1FAE69FFD7E52451CC83E16132FAB5" box="[489,569,1257,1281]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="22" pageNumber="139">IRSNB</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B663CF1FFD7E524526C83E1626DFAD5" ID-GBIF-Occurrence="3450162411" collectionCode="BMNH" collectorName="V. Assing" location="Zyras" municipality="Belgaum /" pageId="22" pageNumber="139" specimenCount="1" typeStatus="syntype">
1 ♀ [damaged; both antennae missing]: “Cotype /
<collectingMunicipality id="6BD5ACD6FFD7E52452208201617EFA95" box="[517,629,1289,1313]" pageId="22" pageNumber="139">Belgaum /</collectingMunicipality>
1266 /
<emphasis id="B97AEABEFFD7E52452E882016011FA95" box="[717,794,1289,1313]" italics="true" pageId="22" pageNumber="139">hastata</emphasis>
Fvl. /
<typeStatus id="54B5880EFFD7E524512782216268FAF5" box="[258,355,1321,1345]" pageId="22" pageNumber="139" type="syntype">Syntypus</typeStatus>
♀
<taxonomicName id="4C0E4D2FFFD7E52451A982226172FAF5" authority="Fauvel" authorityName="Fauvel" authorityYear="1904" box="[396,633,1321,1345]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="139" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B97AEABEFFD7E52451A982226123FAF5" box="[396,552,1321,1345]" italics="true" pageId="22" pageNumber="139">
<location id="8ED16077FFD7E52451A9822262CDFAF5" LSID="urn:lsid:plazi:treatment:03A787BAFFD7E52450C6831B67FEF855:8ED16077FFD7E52451A9822262CDFAF5" box="[396,454,1322,1345]" municipality="Belgaum /" name="Zyras" pageId="22" pageNumber="139">Zyras</location>
hastatus
</emphasis>
<location id="8ED16077FFD7E524521782216172FAF5" LSID="urn:lsid:plazi:treatment:03A787BAFFD7E52450C6831B67FEF855:8ED16077FFD7E524521782216172FAF5" box="[562,633,1321,1345]" municipality="Belgaum /" name="Fauvel" pageId="22" pageNumber="139">Fauvel</location>
</taxonomicName>
, rev.
<collectorName id="26FB537AFFD7E524529282226012FAF5" box="[695,793,1321,1345]" pageId="22" pageNumber="139">V. Assing</collectorName>
2016” (
<collectionCode id="ED1FAE69FFD7E524512D82416255FAD5" box="[264,350,1353,1377]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="22" pageNumber="139">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD7E524509882816138F995" pageId="22" pageNumber="139" type="discussion">
<paragraph id="8BB136ACFFD7E524509882816138F995" blockId="22.[189,794,1417,1570]" pageId="22" pageNumber="139">
<emphasis id="B97AEABEFFD7E524509882816227FA15" bold="true" box="[189,300,1417,1441]" pageId="22" pageNumber="139">Comment</emphasis>
: The original description is based on at least
<specimenCount id="9D08FD25FFD7E524509882A26241FA76" box="[189,330,1450,1474]" pageId="22" pageNumber="139" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
(“Sexus differentia latet”) from “Belgaum” (
<bibRefCitation id="EF9F4B5DFFD7E52450E382C16252FA55" author="FAUVEL, A." box="[198,345,1481,1505]" pageId="22" pageNumber="139" pagination="43 - 70" refId="ref57899" refString="FAUVEL, A. 1904: Staphylinides de l'Hindoustan et de la Birmanie. - Revue d'Entomologie 23: 43 - 70." type="journal article" year="1904">FAUVEL 1904</bibRefCitation>
). One
<typeStatus id="54B5880EFFD7E524518082C262FCFA56" box="[421,503,1482,1506]" pageId="22" pageNumber="139" type="syntype">syntype</typeStatus>
was located in the collections of the IRSNB and another in the collections of the BMNH. Both
<typeStatus id="54B5880EFFD7E5245176810262A4F996" box="[339,431,1546,1570]" pageId="22" pageNumber="139" type="syntype">syntypes</typeStatus>
are females.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD7E524509881416713F835" pageId="22" pageNumber="139" type="description">
<paragraph id="8BB136ACFFD7E524509881416163F855" blockId="22.[189,794,1609,2081]" pageId="22" pageNumber="139">
<emphasis id="B97AEABEFFD7E524509881416251F9D5" bold="true" box="[189,346,1609,1633]" pageId="22" pageNumber="139">Redescription</emphasis>
: Body length 4.1–6.0 mm; length of forebody
<quantity id="4CF69B49FFD7E52451258161628EF935" box="[256,389,1641,1665]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.8" metricValueMin="2.1" pageId="22" pageNumber="139" unit="mm" value="2.45" valueMax="2.8" valueMin="2.1">2.1–2.8 mm</quantity>
. Coloration (
<figureCitation id="13352A29FFD7E52452058161617AF935" box="[544,625,1641,1665]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="22" pageNumber="139">Figs 16</figureCitation>
,
<figureCitation id="13352A29FFD7E52452A781616195F935" box="[642,670,1641,1665]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="22" pageNumber="139">54</figureCitation>
,
<figureCitation id="13352A29FFD7E524528A816161C6F935" box="[687,717,1641,1665]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="22" pageNumber="139">90</figureCitation>
): head black; pronotum pale-reddish; elytra reddish-yellow with the postero-lateral portion extensively blackish (leaving only the scutellum and its vicinity, the anterior margin, and the suture reddish-yellow); abdomen bicoloured with tergites II–V pale-reddish, tergite VI black with the anterior margin and the antero-lateral portions reddish, and tergites VII–VIII black; legs pale-yellowish; antennae dark-brown with antennomeres I–III dark-reddish and the apical half of antennomere XI dark-reddish; maxillary palpi reddish with the terminal palpomere yellowish.
</paragraph>
<paragraph id="8BB136ACFFD7E524509880E167D3FE55" blockId="22.[189,794,1609,2081]" lastBlockId="22.[831,1437,233,1665]" pageId="22" pageNumber="139">
Head (
<figureCitation id="13352A29FFD7E524512F80E1625EF7B5" box="[266,341,2025,2049]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="22" pageNumber="139">Fig. 54</figureCitation>
) moderately transverse, median portion extensively impunctate; punctures in lateral portions moderately dense and large, but shallow. Eyes moderately large, but weakly convex, much longer than postocular region in dorsal view. Antenna (
<figureCitation id="13352A29FFD7E52454FA86216622FEF5" box="[1247,1321,297,321]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="22" pageNumber="139">Fig. 16</figureCitation>
)
<quantity id="4CF69B49FFD7E524551B86216696FEF5" box="[1342,1437,297,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="22" pageNumber="139" unit="mm" value="1.75">1.75 mm</quantity>
long; antennomeres IV approximately as long as broad, V weakly transverse, VI–X of gradually increasing width and increasingly transverse, X approximately 1.5 times as broad as long, and XI conspicuously elongate, nearly as long as the combined length of VII–X.
</paragraph>
<paragraph id="8BB136ACFFD7E524531A86E2670DFD55" blockId="22.[831,1437,233,1665]" pageId="22" pageNumber="139">
Pronotum (
<figureCitation id="13352A29FFD7E524539B86E16702FDB5" box="[958,1033,489,513]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="22" pageNumber="139">Fig. 54</figureCitation>
) distinctly transverse, approximately 1.2 times as broad as long and 1.3 times as broad as head, broadest anterior to middle; lateral margins smoothly convex (dorsal view); punctation not very coarse, rather sparse, and somewhat irregularly distributed; midline narrowly impunctate; lateral margins each with four long, black setae, anterior margin with an additional long black seta on either side.
</paragraph>
<paragraph id="8BB136ACFFD7E524531A85E1675FFC15" blockId="22.[831,1437,233,1665]" pageId="22" pageNumber="139">
Elytra (
<figureCitation id="13352A29FFD7E52453B785E160D7FCB5" box="[914,988,745,769]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="22" pageNumber="139">Fig. 54</figureCitation>
) 0.85 times as long as pronotum; punctation moderately coarse, defined, rather dense, and regularly distributed; interstices on average distinctly broader than diameter of punctures. Hind wings fully developed. Metatarsomere I approximately as long as the combined length of II–IV.
</paragraph>
<paragraph id="8BB136ACFFD7E524531A84A16788F9D5" blockId="22.[831,1437,233,1665]" pageId="22" pageNumber="139">
Abdomen (
<figureCitation id="13352A29FFD7E52453E784A16707FC75" box="[962,1036,937,961]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="22" pageNumber="139">Fig. 90</figureCitation>
) narrower than elytra, with deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a transverse row of non-setiferous punctures; tergite III with a lateral long brown seta on either side, posterior margin with three long brown setae on either side and with a median pair of fine short yellowish setae; tergites IV–V with a median pair of punctures, with a lateral seta on either side, posterior margins with four long brown setae on either side, and with a median pair of minute pale setae; tergite VI anteriorly only with few shallow non-setiferous punctures laterally (without such punctures in the middle), with a median pair of punctures, and with approximately 12 setiferous punctures at posterior margin, otherwise impunctate (except for scattered micropunctures); tergite VII anteriorly with a transverse row (not band!) of sparse non-setiferous punctures, with a transverse row of six setiferous punctures at posterior third, and with additional setiferous punctures at posterior margin; tergite VIII with rather sparse long setae only in posterior portion, posterior margin convex, in the middle with indistinct and small concavity.
</paragraph>
<paragraph id="8BB136ACFFD7E524531A816260C5F935" blockId="22.[831,1437,233,1665]" box="[831,974,1641,1665]" pageId="22" pageNumber="139">♂: unknown.</paragraph>
<paragraph id="8BB136ACFFD7E524531A81A16713F835" blockId="22.[831,1437,1705,1921]" pageId="22" pageNumber="139">
<emphasis id="B97AEABEFFD7E524531A81A1671BF975" bold="true" box="[831,1040,1705,1729]" pageId="22" pageNumber="139">Comparative notes</emphasis>
: Among the species of similar coloration (e.g.,
<taxonomicName id="4C0E4D2FFFD7E52453ED81C267E5F955" authority="PACE, 1993" authorityName="PACE" authorityYear="1993" box="[968,1262,1737,1761]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="139" phylum="Arthropoda" rank="species" species="beijingensis">
<emphasis id="B97AEABEFFD7E52453ED81C26750F955" box="[968,1115,1737,1761]" italics="true" pageId="22" pageNumber="139">Z. beijingensis</emphasis>
<bibRefCitation id="EF9F4B5DFFD7E524544E81C167E5F955" author="PACE, R." box="[1131,1262,1737,1761]" pageId="22" pageNumber="139" pagination="69 - 126" refId="ref58430" refString="PACE, R. 1993: Aleocharinae della China (Coleoptera, Staphylinidae). - Bollettino del Museo Civico di Storia Naturale di Verona 17 [1990]: 69 - 126." type="journal article" year="1993">PACE, 1993</bibRefCitation>
</taxonomicName>
),
<taxonomicName id="4C0E4D2FFFD7E524552C81C26677F955" authorityName="Fauvel" authorityYear="1904" box="[1289,1404,1737,1761]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="139" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B97AEABEFFD7E524552C81C26677F955" box="[1289,1404,1737,1761]" italics="true" pageId="22" pageNumber="139">Z. hastatus</emphasis>
</taxonomicName>
is characterized by slender antennae with a conspicuously elongate antennomere XI, a relatively strongly transverse pronotum with smoothly convex lateral margins, and the punctation pattern of the abdomen (particularly of tergites VI and VII).
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFD7E524531A80A167FEF855" pageId="22" pageNumber="139" type="distribution">
<paragraph id="8BB136ACFFD7E524531A80A167FEF855" blockId="22.[831,1436,1961,2018]" pageId="22" pageNumber="139">
<emphasis id="B97AEABEFFD7E524531A80A160C0F875" bold="true" box="[831,971,1961,1985]" pageId="22" pageNumber="139">Distribution</emphasis>
: This species is currently known only from the
<typeStatus id="54B5880EFFD7E524534D80C2609CF856" box="[872,919,1994,2018]" pageId="22" pageNumber="139">type</typeStatus>
locality in South
<collectingCountry id="F319763CFFD7E524547280C1679BF855" box="[1111,1168,1993,2017]" name="India" pageId="22" pageNumber="139">India</collectingCountry>
(
<figureCitation id="13352A29FFD7E52454BB80C167ECF855" box="[1182,1255,1993,2017]" captionStart="Map 7" captionStartId="67.[151,196,1246,1267]" captionTargetBox="[351,1198,232,1238]" captionTargetId="figure-79@67.[351,1198,232,1238]" captionTargetPageId="67" captionText="Map 7: Revised distributions of Zyras gardneri (black circles), Z. hirsutiventris (white diamonds), Z. parageminus (black triangles), Z. hastatus (white triangle), Z. hirtus (white circles), and Z. latilobatus (star) in India and adjacent regions." figureDoi="http://doi.org/10.5281/zenodo.5742413" httpUri="https://zenodo.org/record/5742413/files/figure.png" pageId="22" pageNumber="139">Map 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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