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<document id="1F41617815719CDF680C782A226C4CDA" ID-CLB-Dataset="46304" ID-DOI="10.5281/zenodo.205904" ID-GBIF-Dataset="62376e09-f30b-4fb0-a635-8f8f43e008fd" ID-ISSN="1175-5326" ID-Zenodo-Dep="205904" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460391967158" checkinUser="plazi" docAuthor="Sasakawa, Mitsuhiro" docDate="2011" docId="03A787A1FFC6FFAEFF5EFCC7EB3E1BFD" docLanguage="en" docName="zt03038p028.pdf" docOrigin="Zootaxa 3038" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Phylloclusia bivittata Sasakawa, 2011, sp. nov." docType="treatment" docVersion="14" lastPageNumber="4" masterDocId="FF9EFFD9FFC4FFADFFC9FFB5EF5D1878" masterDocTitle="Oriental species of clusiid flies (Diptera: Clusiidae: Clusiinae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="3" updateTime="1698298315006" updateUser="ExternalLinkService">
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<mods:title id="842F801D35D06190D9EE092413EFA2D1">Oriental species of clusiid flies (Diptera: Clusiidae: Clusiinae)</mods:title>
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<mods:name id="AA1F0419B5D1480C995F94B482B3DA1A" type="personal">
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<mods:namePart id="5DB55535BE0E6B21A0F57E7F7C723A2F">Sasakawa, Mitsuhiro</mods:namePart>
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<mods:title id="8A71669D8F0CDAF8CC8DD4CD48965C1B">Zootaxa</mods:title>
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<mods:date id="ED82EC026F6D6B88286971F420A921D6">2011</mods:date>
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<treatment id="03A787A1FFC6FFAEFF5EFCC7EB3E1BFD" ID-DOI="http://doi.org/10.5281/zenodo.3505792" ID-GBIF-Taxon="119416985" ID-Zenodo-Dep="3505792" LSID="urn:lsid:plazi:treatment:03A787A1FFC6FFAEFF5EFCC7EB3E1BFD" httpUri="http://treatment.plazi.org/id/03A787A1FFC6FFAEFF5EFCC7EB3E1BFD" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="C314653CFFC6FFAFFF5EFCC7EE571BD6" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BB136B7FFC6FFAFFF5EFCC7EEB71BF3" blockId="2.[151,490,882,942]" box="[151,490,882,907]" pageId="2" pageNumber="3">
<heading id="D0F981DBFFC6FFAFFF5EFCC7EEB71BF3" bold="true" box="[151,490,882,907]" fontSize="10" level="2" pageId="2" pageNumber="3" reason="2">
<emphasis id="B97AEAA5FFC6FFAFFF5EFCC7EEB71BF3" bold="true" box="[151,490,882,907]" pageId="2" pageNumber="3">
<taxonomicName id="4C0E4D34FFC6FFAFFF5EFCC7EED71BF3" ID-CoL="8DC4S" box="[151,394,882,907]" class="Insecta" family="Clusiidae" genus="Phylloclusia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bivittata" status="sp. nov.">
<emphasis id="B97AEAA5FFC6FFAFFF5EFCC7EED71BF3" bold="true" box="[151,394,882,907]" italics="true" pageId="2" pageNumber="3">Phylloclusia bivittata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957DEFFC6FFAFFE59FCC6EEB71BF3" box="[400,490,883,907]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB136B7FFC6FFAFFF5EFC23EE571BD6" blockId="2.[151,490,882,942]" box="[151,266,918,942]" pageId="2" pageNumber="3">
(
<figureCitation id="13352A32FFC6FFAFFF56FC23EE5C1BD6" box="[159,257,918,942]" captionStart="FIGURES 1 5" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[254,1383,198,1888]" captionTargetId="figure@4.[210,1387,195,1894]" captionTargetPageId="4" captionText="FIGURES 1 5. Male genitalia of Phylloclusia bivittata sp. nov. (1 3, paratype) and P. sternopleuralis sp. nov. (4, 5, paratype). 1, 2, 4, 5, epandrium (E), surstylus (S), cercus (C), and proctiger (P), lateral view (1,4) and anterior view (2,5); 3, 4, hypandrium (H), phallapodeme (PA), and phallus, lateral view (parts of phallus: BP, basiphallus; DP, distiphallus; EA, ejaculatory apodeme; EP, epiphallus; MP, mesophallus; PP, paraphallus). Scale = 0.1 mm." httpUri="https://zenodo.org/record/205905/files/figure.png" pageId="2" pageNumber="3">Figs 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFC6FFAFFF5EFC6AEF8D1CFE" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BB136B7FFC6FFAFFF5EFC6AEF8D1CFE" blockId="2.[151,1437,990,2028]" pageId="2" pageNumber="3">
<emphasis id="B97AEAA5FFC6FFAFFF5EFC6AEE181B8F" bold="true" box="[151,325,990,1015]" pageId="2" pageNumber="3">
<typeStatus id="54B58815FFC6FFAFFF5EFC6AEF8D1B8F" box="[151,208,991,1015]" pageId="2" pageNumber="3">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54B58815FFC6FFAFFE85FC6BEEE91B8E" box="[332,436,990,1014]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED1FAE72FFC6FFAFFDCBFC6BED0C1B8E" LSID="urn:lsid:biocol.org:col:34697" box="[514,593,990,1014]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="3">BPBM</collectionCode>
17240),
<collectingCountry id="F3197627FFC6FFAFFD7FFC6AEDB21B8E" box="[694,751,991,1014]" name="Laos" pageId="2" pageNumber="3">Laos</collectingCountry>
, Vientiane Prov., Ban Van Eue,
<date id="FFB01077FFC6FFAFFB91FC6BEB801B8E" box="[1112,1245,990,1014]" pageId="2" pageNumber="3" value="1967-05-15">15. V. 1967</date>
, native collector.
<typeStatus id="54B58815FFC6FFAFFF5EFBB6EE551C63" box="[151,264,1027,1051]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
: 63, same locality as
<typeStatus id="54B58815FFC6FFAFFE35FBB7ED3C1C62" box="[508,609,1026,1050]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
, 15 &amp; 29. IV. and 15.
<date id="FFB01077FFC6FFAFFC9DFBB6EC861C62" box="[852,987,1026,1050]" pageId="2" pageNumber="3" value="1966-05-30">V. 1966, 30.</date>
VIII. and 30. IX. 1967, native collector, light trap; 3,
<collectingCountry id="F3197627FFC6FFAFFEFCFB93EECA1C46" box="[309,407,1062,1086]" name="Vietnam" pageId="2" pageNumber="3">Vietnam</collectingCountry>
,
<quantity id="4CF69B52FFC6FFAFFE6CFB93EEBD1C46" box="[421,480,1062,1086]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="3" unit="km" value="6.0">6 km</quantity>
S of Dalat,
<quantity id="4CF69B52FFC6FFAFFDB9FB93EC521C46" box="[624,783,1062,1086]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="2" pageNumber="3" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">14001500 m</quantity>
,
<date id="FFB01077FFC6FFAFFCD4FB93ECA31C46" box="[797,1022,1062,1086]" pageId="2" pageNumber="3" value="1961-06-09" valueMax="1961-07-07" valueMin="1961-06-09">9. VI.7. VII. 1961</date>
, N. R. Spencer; Ƥ, same locality as
<typeStatus id="54B58815FFC6FFAFFF5EFBFFEFA11C1A" box="[151,252,1098,1122]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
,
<date id="FFB01077FFC6FFAFFEC1FBFFEEC61C1A" box="[264,411,1098,1122]" pageId="2" pageNumber="3" value="1965-09-30">30. IX. 1965</date>
, native collector, Ƥ, S.
<collectingCountry id="F3197627FFC6FFAFFD6FFBFFEDB11C1A" box="[678,748,1098,1122]" name="China" pageId="2" pageNumber="3">China</collectingCountry>
, Fukien, Shaowu, Taohulau,
<quantity id="4CF69B52FFC6FFAFFBFFFBFFEBD31C1A" box="[1078,1166,1098,1122]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="m" value="1000.0">1000 m</quantity>
,
<date id="FFB01077FFC6FFAFFB52FBFFEA051C1A" box="[1179,1368,1098,1122]" pageId="2" pageNumber="3" value="1945-08-18" valueMax="1945-08-20" valueMin="1945-08-18">18-20 VIII 1945</date>
, T. C. Maa.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFC6FFAFFF0CFB27ED571CB6" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BB136B7FFC6FFAFFF0CFB27ED571CB6" blockId="2.[151,1437,990,2028]" pageId="2" pageNumber="3">
<emphasis id="B97AEAA5FFC6FFAFFF0CFB27EE621CD3" bold="true" box="[197,319,1170,1195]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
This species has two brown stripes on the yellow mesoscutum, a laterally protrudent surstylus, and a long, membranous distiphallus.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFC6FFAEFF0EFB6FEEDD1AF1" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
<paragraph id="8BB136B7FFC6FFAFFF0EFB6FED6C1EB4" blockId="2.[151,1437,990,2028]" pageId="2" pageNumber="3">
<emphasis id="B97AEAA5FFC6FFAFFF0EFB6FEE041C8B" bold="true" box="[199,345,1242,1267]" pageId="2" pageNumber="3">Description.</emphasis>
MALE. Head yellow; frons shiny, faintly brown-tinged on dorsal half; ocellar triangle black; face with carina pale brown or black at middle (colored part 1/31/2 as long as whole length of carina). Antenna yellow to brownish yellow; first flagellomere brown on dorsal 1/31/4, with whitish pile; second and third flagellomeres blackish brown; arista brownish yellow; palpus yellow. Thorax brownish yellow; mesoscutum matte, with two brown stripes just laterad of dc-lines throughout entire length of scutum, these distinctly broadened before transverse suture (about 1/2 as wide as distance between dc- and sa-lines, about twice as wide as postsutural stripe), sometimes connecting with each other along anterior margin of scutum or narrowly interrupted on transverse suture, posteriorly extending to lateral margins of scutellum and ending before bases of asc; lsc situated on edge of yellow area of scutellum; postpronotal lobe and pleura yellow; anatergite entirely and mediotergite centrally brown. Wing hyaline, with pale brown marking on apical 1/31/4 of R2+3, this marking extending anteriorly to costa, but not reaching to R4+5; calypter pale gray, with margin and fringe brown; halter yellow. Legs yellow; fore tarsus brown. Abdomen shining; T13 brownish yellow to pale brown, more or less darkened centrally as fasciae; T46 and S8 black; epandrium, surstylus, cercus, and sternites yellowish brown, but epandrium blackened dorsomesally. Bristles brownish yellow.
</paragraph>
<paragraph id="8BB136B7FFC6FFAFFF0CF96CEC5F1FDC" blockId="2.[151,1437,990,2028]" pageId="2" pageNumber="3">Frons as wide as eye, diverging ventrally; frontalia bare; parafrontalia not projecting above eye margin in profile, dorsal parafacialia narrowly projecting beyond that; first and second or subequal in length, third or longer than upper two; oh minute, in row; oc slightly shorter than first or; poc as long as oc; facial carina distinctly swollen below antennal bases; gena narrow, 1/10 as high as eye; pm five. Antennae separated at bases by distance equal to diameter of scape; arista arising just behind dorsal tip of first flagellomere, densely pubescent. Palpus broad, almost as wide as base of first flagellomere, setulose ventrally.</paragraph>
<paragraph id="8BB136B7FFC6FFAEFF0CF804EC4D1882" blockId="2.[151,1437,990,2028]" lastBlockId="3.[151,1437,151,902]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">Mesoscutum with two postsutural dc, anterior dc about 1/2 as long as posterior dc and situated behind level of sa-base; six rows of acr; prsc subequal to anterior dc in length and situated before level of posterior dc; scutellum with lsc 2/3 as long as asc. Wing 3.44.7 (4.0) mm long; three costal sections in proportion of 8085: 15: 910, rm almost at basal 1/3 of cell dm, M1 ratio 1.82.2, ultimate section of CuA1 about 1/3 as long as penultimate section. Mid tibia with one long and one or two short spurs.</paragraph>
<paragraph id="8BB136B7FFC7FFAEFF0CFEB2EEAF19D7" blockId="3.[151,1437,151,902]" pageId="3" pageNumber="4">
S8 slightly longer than epandrium. Genitalia: epandrium (
<figureCitation id="13352A32FFC7FFAEFC9AFEB2ECE41967" box="[851,953,263,287]" captionStart="FIGURES 1 5" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[254,1383,198,1888]" captionTargetId="figure@4.[210,1387,195,1894]" captionTargetPageId="4" captionText="FIGURES 1 5. Male genitalia of Phylloclusia bivittata sp. nov. (1 3, paratype) and P. sternopleuralis sp. nov. (4, 5, paratype). 1, 2, 4, 5, epandrium (E), surstylus (S), cercus (C), and proctiger (P), lateral view (1,4) and anterior view (2,5); 3, 4, hypandrium (H), phallapodeme (PA), and phallus, lateral view (parts of phallus: BP, basiphallus; DP, distiphallus; EA, ejaculatory apodeme; EP, epiphallus; MP, mesophallus; PP, paraphallus). Scale = 0.1 mm." httpUri="https://zenodo.org/record/205905/files/figure.png" pageId="3" pageNumber="4">Figs 1, 2</figureCitation>
) with surstylus roundly protruding anterolaterally, slightly projecting ventrally, and sparsely setulose; cercus longer than 1/2 of epandrial height, with long posterior setae; hypandrium (
<figureCitation id="13352A32FFC7FFAEFE20FEFAED72191F" box="[489,559,335,359]" captionStart="FIGURES 1 5" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[254,1383,198,1888]" captionTargetId="figure@4.[210,1387,195,1894]" captionTargetPageId="4" captionText="FIGURES 1 5. Male genitalia of Phylloclusia bivittata sp. nov. (1 3, paratype) and P. sternopleuralis sp. nov. (4, 5, paratype). 1, 2, 4, 5, epandrium (E), surstylus (S), cercus (C), and proctiger (P), lateral view (1,4) and anterior view (2,5); 3, 4, hypandrium (H), phallapodeme (PA), and phallus, lateral view (parts of phallus: BP, basiphallus; DP, distiphallus; EA, ejaculatory apodeme; EP, epiphallus; MP, mesophallus; PP, paraphallus). Scale = 0.1 mm." httpUri="https://zenodo.org/record/205905/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
) 440 µm long; phallapodeme 530 µm long; basiphallus short; mesophallus tubular at base but divided into two narrow sclerites distally; distiphallus 590 µm long; paraphallus pointed at base. Body length 3.65.0 (4.6) mm.
</paragraph>
<paragraph id="8BB136B7FFC7FFAEFF0EFE0EEEDD1AF1" blockId="3.[151,1437,151,902]" pageId="3" pageNumber="4">
FEMALE. Similar to male, but brown stripes on mesoscutum never interrupted on transverse suture, notopleuron pale brown, anepisternum brown on anterodorsal 1/3, mediotergite entirely blackish brown except for central pale line, apical marking of R2+3 larger, extending between costa and M1, abdomen brownish black, only T3 brown but anterolaterally yellow (1/3 as long as whole tergal length), T6 with yellowish brown posterior margin, T7 brownish black on basal half but distally yellowish brown; prsc behind level of posterior dc; wing length 4.0 mm; body length
<quantity id="4CF69B52FFC7FFAEFEEAFDC4EE211AF1" box="[291,380,625,649]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="3" pageNumber="4" unit="mm" value="4.9">4.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFC7FFAEFF0EFD20EDDD1AD5" box="[199,640,661,686]" pageId="3" pageNumber="4" type="distribution">
<paragraph id="8BB136B7FFC7FFAEFF0EFD20EDDD1AD5" blockId="3.[151,1437,151,902]" box="[199,640,661,686]" pageId="3" pageNumber="4">
<emphasis id="B97AEAA5FFC7FFAEFF0EFD20EE3D1AD6" bold="true" box="[199,352,661,686]" pageId="3" pageNumber="4">Distribution.</emphasis>
<collectingCountry id="F3197627FFC7FFAEFEAEFD23EEFD1AD5" box="[359,416,662,685]" name="Laos" pageId="3" pageNumber="4">Laos</collectingCountry>
,
<collectingCountry id="F3197627FFC7FFAEFE65FD20ED531AD5" box="[428,526,661,685]" name="Vietnam" pageId="3" pageNumber="4">Vietnam</collectingCountry>
, S.
<collectingCountry id="F3197627FFC7FFAEFDFFFD20ED201AD5" box="[566,637,661,685]" name="China" pageId="3" pageNumber="4">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFC7FFAEFF0EFD0FEE7F1B19" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8BB136B7FFC7FFAEFF0EFD0FEE7F1B19" blockId="3.[151,1437,151,902]" pageId="3" pageNumber="4">
<emphasis id="B97AEAA5FFC7FFAEFF0EFD0FEE661AAA" bold="true" box="[199,315,698,722]" pageId="3" pageNumber="4">Remarks.</emphasis>
This new species differs clearly from
<taxonomicName id="4C0E4D34FFC7FFAEFD2CFD0CEB7F1AA9" box="[741,1058,697,721]" class="Insecta" family="Clusiidae" genus="Phylloclusia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sternopleuralis" status="sp. nov.">
<emphasis id="B97AEAA5FFC7FFAEFD2CFD0CEB7F1AA9" box="[741,1058,697,721]" italics="true" pageId="3" pageNumber="4">Phylloclusia sternopleuralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957DEFFC7FFAEFBE1FD0FEBDC1AAA" box="[1064,1153,698,722]" pageId="3" pageNumber="4" rank="species">
<emphasis id="B97AEAA5FFC7FFAEFBE1FD0FEBDC1AAA" bold="true" box="[1064,1153,698,722]" pageId="3" pageNumber="4">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C0E4D34FFC7FFAEFB7FFD0FEA791AA9" box="[1206,1316,697,721]" class="Insecta" family="Clusiidae" genus="Phylloclusia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="hendeli">
<emphasis id="B97AEAA5FFC7FFAEFB7FFD0FEB991AA9" box="[1206,1220,698,721]" italics="true" pageId="3" pageNumber="4">P</emphasis>
.
<emphasis id="B97AEAA5FFC7FFAEFB1BFD0CEA791AA9" box="[1234,1316,697,721]" italics="true" pageId="3" pageNumber="4">hendeli</emphasis>
</taxonomicName>
in the coloration of the thorax and shape of the surstylus as stated in the key to the species. It is similar to the Japanese
<taxonomicName id="4C0E4D34FFC7FFAEFA96FD6BED621B61" authority="Sueyoshi, 2006" authorityName="Sueyoshi" authorityYear="2006" class="Insecta" family="Clusiidae" genus="Phylloclusia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="quadrivittata">
<emphasis id="B97AEAA5FFC7FFAEFA96FD6BEED51B61" italics="true" pageId="3" pageNumber="4">Phylloclusia quadrivittata</emphasis>
<bibRefCitation id="EF9F4B46FFC7FFAEFE46FCB4ED621B61" author="Sueyoshi" box="[399,575,769,793]" pageId="3" pageNumber="4" refString="Sueyoshi, M. (2006) Species diversity of Japanese Clusiidae (Diptera: Acalyptrata) with description of 12 new species. Annales de la Societe Entomologique de France (n. s.), 42, 1 - 26." type="journal article" year="2006">Sueyoshi, 2006</bibRefCitation>
</taxonomicName>
, which also has a laterally protrudent surstylus, but is easily distinguished by its bivittate mesoscutum, dark abdominal tergites, long mesophallus, and simple paraphallus (see
<bibRefCitation id="EF9F4B46FFC7FFAEFB24FC90EAC51B45" author="Sueyoshi" box="[1261,1432,805,829]" pageId="3" pageNumber="4" refString="Sueyoshi, M. (2006) Species diversity of Japanese Clusiidae (Diptera: Acalyptrata) with description of 12 new species. Annales de la Societe Entomologique de France (n. s.), 42, 1 - 26." type="journal article" year="2006">Sueyoshi 2006</bibRefCitation>
: fig. 27DF).
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFC7FFAEFF0CFCDBEB3E1BFD" box="[197,1123,877,902]" pageId="3" pageNumber="4" type="etymology">
<paragraph id="8BB136B7FFC7FFAEFF0CFCDBEB3E1BFD" blockId="3.[151,1437,151,902]" box="[197,1123,877,902]" pageId="3" pageNumber="4">
<emphasis id="B97AEAA5FFC7FFAEFF0CFCDBEE111BFE" bold="true" box="[197,332,878,902]" pageId="3" pageNumber="4">Etymology.</emphasis>
The specific name refers to the two brown stripes on the mesoscutum.
</paragraph>
</subSubSection>
</treatment>
</document>