386 lines
54 KiB
XML
386 lines
54 KiB
XML
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<mods:title id="59DE0D03932D6A13C0162B3FF1C9B0B2">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:namePart id="564D8A50E204CEB1E06DB830C3C74D50">Merle, Didier</mods:namePart>
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<mods:namePart id="FBFC31413265C52E492EC89C9705C098">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="4CB6F0947B6C4D07DD17DC1AFAB42722">Ledon, Daniel</mods:namePart>
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<mods:affiliation id="51C103118127408ECB12014BB5AEB496">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:nameIdentifier id="CEB1815EC3F8373B7FAEFEB69E9E6DC4" type="email">didier.merle@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="A0F2140CBF1CFAC6A208BFC245BE52E3">Goret, Bernard</mods:namePart>
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<mods:affiliation id="85D3E0B08DCA04FC545704C85707161C">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
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<mods:nameIdentifier id="2560E95EB0D22AE786C7BF554A5B445A" type="email">bernardgoret@hotmail.com</mods:nameIdentifier>
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<mods:title id="360DE2BE33C70DE793A14FAD433B3169">Geodiversitas</mods:title>
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<mods:date id="0DEDB3DFA528E16318EDDF5170E8A0D1">2024</mods:date>
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<mods:number id="8B13C194DA85F5186ADAC1D24F9101A2">2024-10-24</mods:number>
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<mods:number id="09753C25EB91249DB78AAB657BC6444A">46</mods:number>
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<subSubSection id="C33D6547FFF7E417FEBDFEC1FD07FE81" box="[261,646,279,306]" pageId="17" pageNumber="510" type="nomenclature">
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<paragraph id="8B9836CCFFF7E417FEBDFEC1FD07FE81" blockId="17.[261,646,279,306]" box="[261,646,279,306]" pageId="17" pageNumber="510">
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<heading id="D0D081A0FFF7E417FEBDFEC1FD07FE81" box="[261,646,279,306]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="510" reason="2">
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Genus
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<taxonomicName id="4C274D4FFFF7E417FEEAFEC1FD07FE81" ID-CoL="7PJK5" authority="de Gregorio, 1885" authorityName="de Gregorio" authorityYear="1885" box="[338,646,279,306]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
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<emphasis id="B953EADEFFF7E417FEEAFEC1FE38FE82" box="[338,441,279,305]" italics="true" pageId="17" pageNumber="510">Timbellus</emphasis>
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de Gregorio, 1885
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</taxonomicName>
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<subSubSection id="C33D6547FFF7E417FF3CFE86FD10FE31" pageId="17" pageNumber="510" type="materials_examined">
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. —
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<taxonomicName id="4C274D4FFFF7E417FE81FE87FDECFEDB" authority="Bellardi, 1873" authorityName="Bellardi" authorityYear="1873" box="[313,621,336,360]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="latifolius">
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<emphasis id="B953EADEFFF7E417FE81FE87FE52FEDB" box="[313,467,336,360]" italics="true" pageId="17" pageNumber="510">Murex latifolius</emphasis>
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Bellardi, 1873
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</taxonomicName>
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, by subsequent designation (
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<bibRefCitation id="EFB64B3DFFF7E417FEBBFEBCFE1FFE30" author="VOKES E. H." box="[259,414,362,387]" pageId="17" pageNumber="510" pagination="1 - 41" refId="ref48825" refString="VOKES E. H. 1964. - Supraspecific groups in the subfamilies Muricinae and Tritonaliinae (Gastropoda: Muricidae). Malacologia 2 (1): 1 - 41. https: // www. biodiversitylibrary. org / page / 13109078" type="journal article" year="1964">Vokes 1964: 14</bibRefCitation>
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||
). Middle Miocene:
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<collectingCountry id="F330765CFFF7E417FDD8FEBCFD0CFE31" box="[608,653,362,386]" name="Italy" pageId="17" pageNumber="510">Italy</collectingCountry>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33D6547FFF7E417FF3CFE61FB14FD23" pageId="17" pageNumber="510" type="discussion">
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||
<paragraph id="8B9836CCFFF7E417FF3CFE61FDECFE7C" blockId="17.[130,777,439,1070]" box="[132,621,439,465]" pageId="17" pageNumber="510">
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</paragraph>
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||
<paragraph id="8B9836CCFFF7E417FF3CFE00FD0AFB9D" blockId="17.[130,777,439,1070]" pageId="17" pageNumber="510">
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||
Historically, fossil and Recent members of
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||
<taxonomicName id="4C274D4FFFF7E417FDDEFE00FD4FFE43" authorityName="de Gregorio" authorityYear="1885" box="[614,718,470,496]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
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||
<emphasis id="B953EADEFFF7E417FDDEFE00FD4FFE43" box="[614,718,470,496]" italics="true" pageId="17" pageNumber="510">Timbellus</emphasis>
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||
</taxonomicName>
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||
have usually been attributed to the genus
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||
<taxonomicName id="4C274D4FFFF7E417FD9BFE21FF3EFD83" authority="Swainson, 1833" authorityName="Swainson" authorityYear="1833" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
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||
<emphasis id="B953EADEFFF7E417FD9BFE21FD10FDA3" box="[547,657,503,528]" italics="true" pageId="17" pageNumber="510">Pterynotus</emphasis>
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||
Swainson, 1833
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||
</taxonomicName>
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||
(e.g.,
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||
<bibRefCitation id="EFB64B3DFFF7E417FEB0FDC0FE3DFD83" author="COSSMANN M." box="[264,444,534,560]" pageId="17" pageNumber="510" refId="ref43425" refString="COSSMANN M. 1889. - Catalogue illustre des coquilles fossiles de l'Eocene des environs de Paris faisant suite aux travaux paleontologiques de G. - P. Deshayes. 4 eme fascicule. Annales de la Societe royale Malacologique de Belgique 24: 1 - 385. [Also published as Cossmann M. 1889 (December). - Catalogue illustre des coquilles fossiles de l'Eocene des environs de Paris faisant suite aux travaux paleontologiques de G. - P. Deshayes. 4 eme fascicule, Societe royale malacologique de Belgique, Bruxelles, 385 p.]" type="book" year="1889">Cossmann 1889</bibRefCitation>
|
||
;
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||
<bibRefCitation id="EFB64B3DFFF7E417FE74FDC0FC81FD83" author="COSSMANN M. & PISSARRO G." box="[460,768,534,560]" pageId="17" pageNumber="510" pagination="26 - 45" refId="ref43883" refString="COSSMANN M. & PISSARRO G. 1911. - Iconographie complete des coquilles fossiles de l'Eocene du Bassin de Paris. Tome 2. Hermann, Paris: pl. 26 - 45. https: // www. biodiversitylibrary. org / page / 36319938" type="book chapter" year="1911">Cossmann & Pissarro 1911</bibRefCitation>
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||
;
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||
<bibRefCitation id="EFB64B3DFFF7E417FF3CFDE0FE9BFDE3" author="GLIBERT M." box="[132,282,566,592]" pageId="17" pageNumber="510" pagination="1 - 179" refId="ref44626" refString="GLIBERT M. 1963. - Les Muricacea et les Buccinacea du Cenozoique etranger des collections de l'Institut royal des Sciences naturelles de Belgique. Memoires de l'Institut royal des Sciences naturelles de Belgique serie 2, 74: 1 - 179." type="journal article" year="1963">Glibert 1963</bibRefCitation>
|
||
;
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||
<bibRefCitation id="EFB64B3DFFF7E417FE93FDE0FE30FDE3" author="VOKES E. H." box="[299,433,566,592]" pageId="17" pageNumber="510" pagination="1 - 50" refId="ref48869" refString="VOKES E. H. 1970. - Cenozoic Muricidae of the Western Atlantic region. Part V. Pterynotus and Poirieria. Tulane Studies in Geology and Paleontology 8 (1): 1 - 50." type="journal article" year="1970">Vokes 1970</bibRefCitation>
|
||
) in the subfamily
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||
<taxonomicName id="4C274D4FFFF7E417FD33FDE0FC82FDE3" authorityName="Rafinesque" authorityYear="1815" box="[651,771,566,592]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="subFamily" subFamily="Muricinae">Muricinae</taxonomicName>
|
||
. Based on molecular data,
|
||
<bibRefCitation id="EFB64B3DFFF7E417FE27FD80FDE1FDC3" author="BARCO A. & CLAREMONT M. & REID D. G. & HOUART R. & BOUCHET P. & WILLIAMS S. T. & CRUAUD C. & COULOUX A. & OLIVERIO M." box="[415,608,598,624]" pageId="17" pageNumber="510" pagination="1025 - 1039" refId="ref42748" refString="BARCO A., CLAREMONT M., REID D. G., HOUART R., BOUCHET P., WILLIAMS S. T., CRUAUD C., COULOUX A. & OLIVERIO M. 2010. - A molecular phylogenetic framework for the Muricidae, a diverse family of carnivorous gastropods. Molecular Phylogenetics and Evolution 55 (3): 1025 - 1039. https: // doi. org / 10.1016 / j. ympev. 2010.03.008" type="journal article" year="2010">
|
||
Barco
|
||
<emphasis id="B953EADEFFF7E417FE5BFD81FD99FDC3" box="[483,536,598,624]" italics="true" pageId="17" pageNumber="510">et al.</emphasis>
|
||
(2010
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB64B3DFFF7E417FDD7FD80FD29FDC3" author="BARCO A. & SCHIAPARELLI S. & HOUART R. & OLIVERIO M." box="[623,680,598,624]" pageId="17" pageNumber="510" pagination="596 - 616" refId="ref42832" refString="BARCO A., SCHIAPARELLI S., HOUART R. & OLIVERIO M. 2012. - Cenozoic evolution of Muricidae (Mollusca, Neogastropoda) in the Southern Ocean, with the description of a new subfamily. Zoologica Scripta 41 (6): 596 - 616. https: // doi. org / 10.1111 / j. 1463 - 6409.2012.00554. x" type="journal article" year="2012">2012</bibRefCitation>
|
||
) studied the phylogenetic relationships of
|
||
<taxonomicName id="4C274D4FFFF7E417FE51FDA1FC86FD23" authority="Swainson, 1833" authorityName="Swainson" authorityYear="1833" box="[489,775,630,656]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFF7E417FE51FDA1FDD4FD23" box="[489,597,631,656]" italics="true" pageId="17" pageNumber="510">Pterynotus</emphasis>
|
||
Swainson, 1833
|
||
</taxonomicName>
|
||
represented by
|
||
<taxonomicName id="4C274D4FFFF7E417FE93FD40FDEEFD03" authority="(Lightfoot, 1786)" baseAuthorityName="Lightfoot" baseAuthorityYear="1786" box="[299,623,662,688]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="elongatus">
|
||
<emphasis id="B953EADEFFF7E417FE93FD40FE25FD03" box="[299,420,662,688]" italics="true" pageId="17" pageNumber="510">P. elongatus</emphasis>
|
||
(Lightfoot, 1786)
|
||
</taxonomicName>
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||
and
|
||
<taxonomicName id="4C274D4FFFF7E417FD10FD40FE96FD63" authority="Houart, 1988" authorityName="Houart" authorityYear="1988" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="fulgens">
|
||
<emphasis id="B953EADEFFF7E417FD10FD40FC88FD03" box="[680,777,662,688]" italics="true" pageId="17" pageNumber="510">P. fulgens</emphasis>
|
||
Houart, 1988
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C274D4FFFF7E417FEFFFD63FDE8FD63" authority="Houart, 1995" authorityName="Houart" authorityYear="1995" box="[327,617,693,720]" class="Gastropoda" family="Muricidae" genus="Pterymarchia" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFF7E417FEFFFD63FE51FD7C" box="[327,464,693,719]" italics="true" pageId="17" pageNumber="510">Pterymarchia</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFF7E417FE6EFD60FDE8FD63" author="HOUART R." box="[470,617,693,720]" pageId="17" pageNumber="510" pagination="127 - 136" refId="ref44916" refString="HOUART R. 1995. - Pterymarchia n. gen. and Vaughtia n. gen., two new muricid genera (Gastropoda: Muricidae: Muricinae and Ocenebrinae). Apex 10 (4): 127 - 136. https: // www. biodiversitylibrary. org / page / 42257087" type="journal article" year="1995">Houart, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
represented by
|
||
<taxonomicName id="4C274D4FFFF7E417FF3AFD00FE4FFD43" authority="(Roding, 1798)" baseAuthorityName="Roding" baseAuthorityYear="1798" box="[130,462,725,752]" class="Gastropoda" family="Muricidae" genus="Pterymarchia" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="martinetana">
|
||
<emphasis id="B953EADEFFF7E417FF3AFD00FE9AFD5C" box="[130,283,726,751]" italics="true" pageId="17" pageNumber="510">P. martinetana</emphasis>
|
||
(Röding, 1798)
|
||
</taxonomicName>
|
||
. These authors demonstrated firstly, that
|
||
<taxonomicName id="4C274D4FFFF7E417FF45FD20FEE8FCBC" authorityName="Swainson" authorityYear="1833" box="[253,361,758,783]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFF7E417FF45FD20FEE8FCBC" box="[253,361,758,783]" italics="true" pageId="17" pageNumber="510">Pterynotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C274D4FFFF7E417FE25FD23FDA6FCBC" authorityName="Houart" authorityYear="1995" box="[413,551,757,783]" class="Gastropoda" family="Muricidae" genus="Pterymarchia" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFF7E417FE25FD23FDA6FCBC" box="[413,551,757,783]" italics="true" pageId="17" pageNumber="510">Pterymarchia</emphasis>
|
||
</taxonomicName>
|
||
are not phylogenetically close to
|
||
<taxonomicName id="4C274D4FFFF7E417FEAFFCC0FD93FC9C" authority="Houart, 1988" authorityName="Houart" authorityYear="1988" box="[279,530,789,816]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="fulgens">
|
||
<emphasis id="B953EADEFFF7E417FEAFFCC0FEF7FC9C" box="[279,374,789,815]" italics="true" pageId="17" pageNumber="510">P. fulgens</emphasis>
|
||
Houart, 1988
|
||
</taxonomicName>
|
||
and secondly, none of these taxa are included within the clade of the
|
||
<taxonomicName id="4C274D4FFFF7E417FD28FCE3FC85FCFC" authorityName="Rafinesque" authorityYear="1815" box="[656,772,821,847]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="subFamily" subFamily="Muricinae">Muricinae</taxonomicName>
|
||
. According to
|
||
<bibRefCitation id="EFB64B3DFFF7E417FEADFC83FE5FFCDC" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[277,478,853,879]" pageId="17" pageNumber="510" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFF7E417FEADFC83FED5FCDC" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[277,340,853,879]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFF7E417FEE3FC80FE0FFCDC" box="[347,398,853,879]" italics="true" pageId="17" pageNumber="510">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C274D4FFFF7E417FE53FC80FD22FCDC" authorityName="Houart" authorityYear="1988" box="[491,675,853,879]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="fulgens">
|
||
<emphasis id="B953EADEFFF7E417FE53FC80FD22FCDC" box="[491,675,853,879]" italics="true" pageId="17" pageNumber="510">Pterynotus fulgens</emphasis>
|
||
</taxonomicName>
|
||
is closely related to the
|
||
<typeStatus id="549C886EFFF7E417FEAEFCA3FEC5FC3C" box="[278,324,885,911]" pageId="17" pageNumber="510">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C274D4FFFF7E417FE0DFCA3FD9DFC3C" authorityName="de Gregorio" authorityYear="1885" box="[437,540,885,911]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFF7E417FE0DFCA3FD9DFC3C" box="[437,540,885,911]" italics="true" pageId="17" pageNumber="510">Timbellus</emphasis>
|
||
</taxonomicName>
|
||
and is now attributed to that genus. Thus,
|
||
<bibRefCitation id="EFB64B3DFFF7E417FEDCFC43FDAEFC1C" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[356,559,917,943]" pageId="17" pageNumber="510" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFF7E417FEDCFC43FE22FC1C" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[356,419,917,943]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFF7E417FE12FC40FE5FFC1C" box="[426,478,917,943]" italics="true" pageId="17" pageNumber="510">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFB64B3DFFF7E417FDDEFC43FC86FC1C" author="HOUART R." box="[614,775,917,943]" pageId="17" pageNumber="510" pagination="37 - 66" refId="ref45005" refString="HOUART R. 2018. - Historique et classification des especes actuelles de Muricidae (Neogastropoda, Muricoidea). Novapex 19 (2): 37 - 66." type="journal article" year="2018">Houart (2018)</bibRefCitation>
|
||
excluded
|
||
<taxonomicName id="4C274D4FFFF7E417FF54FC63FED2FC7C" authorityName="de Gregorio" authorityYear="1885" box="[236,339,949,975]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFF7E417FF54FC63FED2FC7C" box="[236,339,949,975]" italics="true" pageId="17" pageNumber="510">Timbellus</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<taxonomicName id="4C274D4FFFF7E417FE7BFC63FDB4FC7C" authorityName="Rafinesque" authorityYear="1815" box="[451,565,949,975]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="subFamily" subFamily="Muricinae">Muricinae</taxonomicName>
|
||
, but did not give a subfamilial attribution.
|
||
<bibRefCitation id="EFB64B3DFFF7E417FE39FC02FD0EFC5D" author="RUSSINI V. & FASSIO G. & NOCELLA E. & HOUART R. & BARCO A. & PUILLANDRE N. & MODICA M. V. & M. OLIVERIO" box="[385,655,980,1007]" pageId="17" pageNumber="510" pagination="856 - 868" refId="ref48453" refString="RUSSINI V., FASSIO G., NOCELLA E., HOUART R., BARCO A., PUILLANDRE N., LOZOUET P, MODICA M. V. & M. OLIVERIO 2023. - Whelks, rock-snails, and allied: a new phylogenetic framework for the family Muricidae (Mollusca: Gastropoda). The European Zoological Journal 90 (2): 856 - 868. https: // doi. org / 10.1080 / 24750263.2023.2283517" type="journal article" year="2023">
|
||
Russini
|
||
<emphasis id="B953EADEFFF7E417FE6FFC03FD88FC5D" box="[471,521,980,1006]" italics="true" pageId="17" pageNumber="510">et al.</emphasis>
|
||
(2023: 866)
|
||
</bibRefCitation>
|
||
place
|
||
<taxonomicName id="4C274D4FFFF7E417FD68FC03FF3FFBBD" authorityName="de Gregorio" authorityYear="1885" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFF7E417FD68FC03FF3FFBBD" italics="true" pageId="17" pageNumber="510">Timbellus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<emphasis id="B953EADEFFF7E417FF67FC23FEE2FBBD" box="[223,355,1012,1038]" italics="true" pageId="17" pageNumber="510">incertae sedis</emphasis>
|
||
and considered it likely to represent an independent lineage worthy of subfamilial rank.
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF7E417FF3CFB82FD42FBDD" blockId="17.[130,778,1108,2028]" box="[132,707,1108,1135]" pageId="17" pageNumber="510">
|
||
<smallCapsWord id="8D7EA010FFF7E417FF3CFB82FF57FBDE" baselines="1127,1128" box="[132,214,1108,1134]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Species" pageId="17" pageNumber="510">SPECIES</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFF7E417FF65FB8CFEA7FBDE" baselines="1128" box="[221,294,1114,1133]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="group" pageId="17" pageNumber="510">GROUP</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFF7E417FE95FB8CFEC8FBDE" baselines="1128" box="[301,329,1114,1133]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="17" pageNumber="510">OF</smallCapsWord>
|
||
<taxonomicName id="4C274D4FFFF7E417FEE8FB83FD42FBDD" authority="(LAMARCK, 1822)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[336,707,1108,1135]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF7E417FEE8FB83FEE8FBDD" box="[336,361,1109,1134]" italics="true" pageId="17" pageNumber="510">T.</emphasis>
|
||
<smallCapsWord id="8D7EA010FFF7E417FEC8FB8CFE7CFBDE" baselines="1128" box="[368,509,1114,1133]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="tripteroides" pageId="17" pageNumber="510">
|
||
<emphasis id="B953EADEFFF7E417FEC8FB8CFE7CFBDE" box="[368,509,1114,1133]" italics="true" pageId="17" pageNumber="510">TRIPTEROIDES</emphasis>
|
||
</smallCapsWord>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFF7E417FDB3FB82FD3AFBDD" author="LAMARCK J. - B." box="[523,699,1108,1135]" pageId="17" pageNumber="510" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">
|
||
<smallCapsWord id="8D7EA010FFF7E417FDB3FB82FDF5FBDE" baselines="1127,1128" box="[523,628,1108,1134]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Lamarck" pageId="17" pageNumber="510">LAMARCK</smallCapsWord>
|
||
, 1822
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF7E417FF3CFBA3FB14FD23" blockId="17.[130,778,1108,2028]" lastBlockId="17.[811,1457,215,656]" pageId="17" pageNumber="510">
|
||
In order to delineate different lineages included in
|
||
<taxonomicName id="4C274D4FFFF7E417FD22FBA3FC87FB3D" authorityName="Swainson" authorityYear="1833" box="[666,774,1141,1166]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFF7E417FD22FBA3FC87FB3D" box="[666,774,1141,1166]" italics="true" pageId="17" pageNumber="510">Pterynotus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B953EADEFFF7E417FF35FB43FF2BFB1D" box="[141,170,1173,1198]" italics="true" pageId="17" pageNumber="510">s.s.</emphasis>
|
||
),
|
||
<bibRefCitation id="EFB64B3DFFF7E417FF78FB42FE7AFB1D" author="HARASEWYCH M. G. & JENSEN R. H." box="[192,507,1172,1198]" pageId="17" pageNumber="510" pagination="1 - 16" refId="ref44794" refString="HARASEWYCH M. G. & JENSEN R. H. 1979. - Review of the subgenus Pterynotus (Gastropoda: Muricidae) in the Western Atlantic. Nemouria 22: 1 - 16. https: // www. biodiversitylibrary. org / page / 45974443" type="journal article" year="1979">Harasewych & Jensen (1979)</bibRefCitation>
|
||
and more recently
|
||
<bibRefCitation id="EFB64B3DFFF7E417FD7EFB42FF4FFB7D" author="VOKES E. H." pageId="17" pageNumber="510" pagination="1 - 108" refId="ref48996" refString="VOKES E. H. 1992. - Cenozoic Muricidae of the Western Atlantic region, Part IX - Pterynotus, Poirieria, Aspella, Dermomurex, Calotrophon, Acantolabia, and Attiliosa; additions and corrections. Tulane Studies in Geology and Paleontology 25 (1 - 3): 1 - 108." type="journal article" year="1992">Vokes (1992)</bibRefCitation>
|
||
defined five “species groups” for the Western Atlantic Region. With the exception of the typical group of
|
||
<taxonomicName id="4C274D4FFFF7E417FD22FB02FC84FB5E" authorityName="Swainson" authorityYear="1833" box="[666,773,1236,1261]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFF7E417FD22FB02FC84FB5E" box="[666,773,1236,1261]" italics="true" pageId="17" pageNumber="510">Pterynotus</emphasis>
|
||
</taxonomicName>
|
||
[species group of
|
||
<taxonomicName id="4C274D4FFFF7E417FE86FB22FD53FABD" authority="(Swainson, 1822)" baseAuthorityName="Swainson" baseAuthorityYear="1822" box="[318,722,1267,1294]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="pinnatus">
|
||
<emphasis id="B953EADEFFF7E417FE86FB22FD8BFABE" box="[318,522,1268,1293]" italics="true" pageId="17" pageNumber="510">Pterynotus pinnatus</emphasis>
|
||
(Swainson, 1822)
|
||
</taxonomicName>
|
||
], the four other species groups are members of the genus
|
||
<taxonomicName id="4C274D4FFFF7E417FD6AFAC2FE15FAFE" authority="(Merle et al. 2011)" baseAuthorityName="Merle" baseAuthorityYear="2011" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFF7E417FD6AFAC2FF3FFAFE" italics="true" pageId="17" pageNumber="510">Timbellus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFF7E417FF77FAE5FE0DFAFE" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[207,396,1331,1357]" pageId="17" pageNumber="510" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
Merle
|
||
<emphasis id="B953EADEFFF7E417FEAFFAE2FECAFAFE" box="[279,331,1331,1357]" italics="true" pageId="17" pageNumber="510">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFB64B3DFFF7E417FE1BFAE5FD2BFAFE" author="MERLE D. & GARRIGUES B. & POINTIER J. - P." box="[419,682,1331,1358]" pageId="17" pageNumber="510" refId="ref47234" refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
|
||
<taxonomicName id="4C274D4FFFF7E417FE1BFAE5FE63FAFE" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[419,482,1331,1357]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
<emphasis id="B953EADEFFF7E417FE53FAE2FD9EFAFE" box="[491,543,1331,1357]" italics="true" pageId="17" pageNumber="510">et al.</emphasis>
|
||
(2011: 130)
|
||
</bibRefCitation>
|
||
added a fifth group to
|
||
<taxonomicName id="4C274D4FFFF7E417FE98FA85FE07FADE" box="[288,390,1363,1389]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFF7E417FE98FA85FE07FADE" box="[288,390,1363,1389]" italics="true" pageId="17" pageNumber="510">Timbellus</emphasis>
|
||
</taxonomicName>
|
||
: the species group of
|
||
<taxonomicName id="4C274D4FFFF7E417FDCDFA82FEC0FA3E" authority="(Lamarck, 1822)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF7E417FDCDFA82FD07FADE" box="[629,646,1364,1389]" italics="true" pageId="17" pageNumber="510">T</emphasis>
|
||
.
|
||
<emphasis id="B953EADEFFF7E417FD2DFA85FC86FADE" box="[661,775,1363,1389]" italics="true" pageId="17" pageNumber="510">tripteroides</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFF7E417FF33FAA5FEB8FA3E" author="LAMARCK J. - B." box="[139,313,1395,1421]" pageId="17" pageNumber="510" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Members of that group display a strong development of the internal denticles within the outer lip and their shell shape is narrower than those belonging to the group of
|
||
<taxonomicName id="4C274D4FFFF7E417FF52FA02FDA3FA5E" authority="(Roding, 1798)" baseAuthorityName="Roding" baseAuthorityYear="1798" box="[234,546,1491,1517]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="crenulatus">
|
||
<emphasis id="B953EADEFFF7E417FF52FA02FEF1FA5E" box="[234,368,1491,1517]" italics="true" pageId="17" pageNumber="510">T. crenulatus</emphasis>
|
||
(Röding, 1798)
|
||
</taxonomicName>
|
||
. Fine growing lamellae forming a scabrous surface are usually missing and only reported in
|
||
<taxonomicName id="4C274D4FFFF7E417FF47F9C5FD54F99E" authority="Pacaud, Ledon & Goret, 2017" authorityName="Pacaud, Ledon & Goret" authorityYear="2017" box="[255,725,1554,1581]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="capitaneus">
|
||
<emphasis id="B953EADEFFF7E417FF47F9C5FE07F99F" box="[255,390,1555,1580]" italics="true" pageId="17" pageNumber="510">T. capitaneus</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFF7E417FE35F9C4FD54F99E" author="PACAUD J. - M. & LEDON D. & GORET B." box="[397,725,1554,1581]" pageId="17" pageNumber="510" pagination="58 - 61" refId="ref48127" refString="PACAUD J. - M., LEDON D. & GORET B. 2017. - Timbellus capitaneus n. sp., une espece nouvelle de Muricidae (Mollusca, Gastropoda) du Lutetien (Eocene moyen) du bassin de Paris. Fossiles. Revue francaise de paleontologie 32: 58 - 61." type="journal article" year="2017">Pacaud, Ledon & Goret, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
. The following new species described herein,
|
||
<taxonomicName id="4C274D4FFFF7E417FD8BF9E5FD3FF9FE" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[563,702,1587,1613]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="magnificus" status="n. sp.">
|
||
<emphasis id="B953EADEFFF7E417FD8BF9E5FD3FF9FE" box="[563,702,1587,1613]" italics="true" pageId="17" pageNumber="510">T. magnificus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF7E417FD7DF9E5FC83F9FE" box="[709,770,1587,1613]" pageId="17" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C274D4FFFF7E417FF3CF985FE97F9DF" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[132,278,1618,1644]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="occidentalis" status="n. sp.">
|
||
<emphasis id="B953EADEFFF7E417FF3CF985FE97F9DF" box="[132,278,1618,1644]" italics="true" pageId="17" pageNumber="510">T. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF7E417FEA6F985FEDAF9DE" box="[286,347,1619,1645]" pageId="17" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C274D4FFFF7E417FED0F985FE5FF9DF" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[360,478,1618,1644]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="calciacus" status="n. sp.">
|
||
<emphasis id="B953EADEFFF7E417FED0F985FE5FF9DF" box="[360,478,1618,1644]" italics="true" pageId="17" pageNumber="510">T. calciacus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF7E417FE5DF985FDA3F9DE" box="[485,546,1619,1645]" pageId="17" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C274D4FFFF7E417FD97F985FD44F9DF" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[559,709,1618,1644]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="longicanalis" status="n. sp.">
|
||
<emphasis id="B953EADEFFF7E417FD97F985FD44F9DF" box="[559,709,1618,1644]" italics="true" pageId="17" pageNumber="510">T. longicanalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF7E417FD74F985FC89F9DE" box="[716,776,1619,1645]" pageId="17" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C274D4FFFF7E417FF0DF9A5FE9FF93F" box="[181,286,1651,1676]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="magnei">
|
||
<emphasis id="B953EADEFFF7E417FF0DF9A5FE9FF93F" box="[181,286,1651,1676]" italics="true" pageId="17" pageNumber="510">T. magnei</emphasis>
|
||
</taxonomicName>
|
||
n. name (for
|
||
<taxonomicName id="4C274D4FFFF7E417FE03F9A5FC82F93E" authority="Rouault, 1850" authorityName="sensu Rouault" authorityYear="1850" box="[443,771,1650,1677]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="trigonus">
|
||
<emphasis id="B953EADEFFF7E417FE03F9A5FDDAF93F" box="[443,603,1651,1676]" italics="true" pageId="17" pageNumber="510">Murex trigonus</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFF7E417FDD9F9A4FC82F93E" author="ROUAULT A." box="[609,771,1650,1677]" pageId="17" pageNumber="510" pagination="457 - 502" refId="ref48421" refString="ROUAULT A. 1850. - Description des fossiles eocenes des environs de Pau. Memoires de la Societe geologique de France, serie 2, 3: 457 - 502." type="journal article" year="1850">Rouault, 1850</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B953EADEFFF7E417FF3EF945FF2CF91F" box="[134,173,1683,1708]" italics="true" pageId="17" pageNumber="510">non</emphasis>
|
||
<taxonomicName id="4C274D4FFFF7E417FF0AF945FE69F91F" authority="Gmelin, 1791" authorityName="Gmelin" authorityYear="1791" box="[178,488,1682,1708]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="trigonus">
|
||
<emphasis id="B953EADEFFF7E417FF0AF945FECCF91F" box="[178,333,1683,1708]" italics="true" pageId="17" pageNumber="510">Murex trigonus</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFF7E417FEE9F944FE69F91F" author="GMELIN J. F." box="[337,488,1682,1708]" pageId="17" pageNumber="510" pagination="3021 - 3910" refId="ref44669" refString="GMELIN J. F. 1791 (ed.). - Caroli a Linnaei Systema Naturae per Regna Tria Naturae, part Vermes. Ed. 13. Tome 1 (6). G. E. Beer, Leipzig: 3021 - 3910. https: // doi. org / 10.5962 / bhl. title. 36932" type="book chapter" year="1791">Gmelin, 1791</bibRefCitation>
|
||
</taxonomicName>
|
||
) display features allowing a placement in this species group. Several of these new species (
|
||
<taxonomicName id="4C274D4FFFF7E417FD4AF965FF78F95F" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="occidentalis" status="n. sp.">
|
||
<emphasis id="B953EADEFFF7E417FD4AF965FF78F95F" italics="true" pageId="17" pageNumber="510">T. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF7E417FF47F904FEBAF95F" box="[255,315,1746,1772]" pageId="17" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C274D4FFFF7E417FEFCF905FE39F95F" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[324,440,1746,1772]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="calciacus" status="n. sp.">
|
||
<emphasis id="B953EADEFFF7E417FEFCF905FE39F95F" box="[324,440,1746,1772]" italics="true" pageId="17" pageNumber="510">T. calciacus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF7E417FE78F904FE7AF95F" box="[448,507,1746,1772]" pageId="17" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C274D4FFFF7E417FDBCF905FD19F95F" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[516,664,1746,1772]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="longicanalis" status="n. sp.">
|
||
<emphasis id="B953EADEFFF7E417FDBCF905FD19F95F" box="[516,664,1746,1772]" italics="true" pageId="17" pageNumber="510">T. longicanalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF7E417FD26F904FD59F95F" box="[670,728,1746,1772]" pageId="17" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C274D4FFFF7E417FF3BF925FF6BF8BF" box="[131,234,1779,1804]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="magnei">
|
||
<emphasis id="B953EADEFFF7E417FF3BF925FF6BF8BF" box="[131,234,1779,1804]" italics="true" pageId="17" pageNumber="510">T. magnei</emphasis>
|
||
</taxonomicName>
|
||
n. name) are superficially similar to, and mistaken for,
|
||
<taxonomicName id="4C274D4FFFF7E417FF16F8C4FEBDF89F" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[174,316,1810,1836]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF7E417FF16F8C4FEBDF89F" box="[174,316,1810,1836]" italics="true" pageId="17" pageNumber="510">T. tripteroides</emphasis>
|
||
</taxonomicName>
|
||
in the past. Moreover, the type material of
|
||
<taxonomicName id="4C274D4FFFF7E417FF3CF8E4FE92F8F8" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[132,275,1841,1867]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF7E417FF3CF8E4FE92F8F8" box="[132,275,1841,1867]" italics="true" pageId="17" pageNumber="510">T. tripteroides</emphasis>
|
||
</taxonomicName>
|
||
is lost, as it is missing in the Lamarck’s collections in Geneva and Paris. In order to avoid future taxonomic mistakes, we designate a
|
||
<typeStatus id="549C886EFFF7E417FE3EF8A4FE5AF83F" box="[390,475,1906,1932]" pageId="17" pageNumber="510" type="neotype">neotype</typeStatus>
|
||
herein (MNHN.F.A90546, coll. of the Club géologique d’Île-de-France,
|
||
<figureCitation id="131C2A49FFF7E417FDE3F847FD4AF818" box="[603,715,1937,1964]" captionStart="FIG" captionStartId="16.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-382@16.[698,1267,772,1332]" captionTargetPageId="16" captionText="FIG. 6. — Neotype, historical figures, spiral cords and denticles of the outer lip in T. tripteroides (Lamarck, 1822): A-D, morphotype with rounded varices and poorly distinguishable P1 spine; E-H, morphotype with easily distinguishable P1 spine: A, original figure of Lamarck (1805: pl. 3, fig. 4); B, C, neotype designated herein, MNHN.F.A90546 (Club géologique d'Île-de-France leg.), Thiverval-Grignon (Falunière), Yvelines, Paris Basin, France, middle Eocene (Lutetian); D, MNHN.F.A90552 (Pacaud coll.), Chaussy (Les Garennes),Paris Basin, France,middle Eocene (Lutetian); E, F, figured specimen of Deshayes (1835: pl. 82, fig. 1-2); G, H, figured specimen of Cossmann & Pissarro (1911: pl. 35, fig. 169-1; MNHN.F.J10350 (Cossmann coll.), Chaussy (Les Garennes). Scale bars: 20 mm. Credits: B-D, P. Loubry MNHN/CNRS; G, H, J. Mouchart." figureDoi="http://doi.org/10.5281/zenodo.13985772" httpUri="https://zenodo.org/record/13985772/files/figure.png" pageId="17" pageNumber="510">Fig. 6B, C</figureCitation>
|
||
). The type material of Lamarck was collected at Grignon (middle Lutetian) and the
|
||
<typeStatus id="549C886EFFF7E417FEEBF804FE2AF85F" box="[339,427,2002,2028]" pageId="17" pageNumber="510" type="neotype">neotype</typeStatus>
|
||
is selected from the same type locality (precisely Falunière of Grignon; Calcaire à Orbitolites complanatus Formation, biozone NP15, see
|
||
<bibRefCitation id="EFB64B3DFFF7E417FAB5FF21FCE5FE82" author="GELY J. - P. & LORENZ C." pageId="17" pageNumber="510" pagination="713 - 747" refId="ref44561" refString="GELY J. - P. & LORENZ C. 1991. - Analyse sequentielle de l'Eocene et de l'Oligocene du Bassin parisien (France). Revue de l'institut francais du petrole 46 (6): 713 - 747." type="journal article" year="1991">Gély & Lorenz 1991</bibRefCitation>
|
||
). The
|
||
<typeStatus id="549C886EFFF7E417FC1FFECEFC7DFE81" box="[935,1020,280,306]" pageId="17" pageNumber="510" type="neotype">neotype</typeStatus>
|
||
was chosen to closely represent the specimen illustrated by
|
||
<bibRefCitation id="EFB64B3DFFF7E417FC42FEE1FB21FEE1" author="LAMARCK J. - B." box="[1018,1184,311,338]" pageId="17" pageNumber="510" pagination="222 - 228" refId="ref45602" refString="LAMARCK J. - B. 1805. - Sur les fossiles des environs de Paris. Explication des planches relatives aux coquilles fossiles des environs de Paris. Annales du Museum d'Histoire Naturelle 6 (33): 222 - 228." type="journal article" year="1805">Lamarck (1805</bibRefCitation>
|
||
: pl. 3, fig. 4;
|
||
<figureCitation id="131C2A49FFF7E417FA8FFEE1FA0AFEE2" box="[1335,1419,311,337]" captionStart="FIG" captionStartId="16.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-382@16.[698,1267,772,1332]" captionTargetPageId="16" captionText="FIG. 6. — Neotype, historical figures, spiral cords and denticles of the outer lip in T. tripteroides (Lamarck, 1822): A-D, morphotype with rounded varices and poorly distinguishable P1 spine; E-H, morphotype with easily distinguishable P1 spine: A, original figure of Lamarck (1805: pl. 3, fig. 4); B, C, neotype designated herein, MNHN.F.A90546 (Club géologique d'Île-de-France leg.), Thiverval-Grignon (Falunière), Yvelines, Paris Basin, France, middle Eocene (Lutetian); D, MNHN.F.A90552 (Pacaud coll.), Chaussy (Les Garennes),Paris Basin, France,middle Eocene (Lutetian); E, F, figured specimen of Deshayes (1835: pl. 82, fig. 1-2); G, H, figured specimen of Cossmann & Pissarro (1911: pl. 35, fig. 169-1; MNHN.F.J10350 (Cossmann coll.), Chaussy (Les Garennes). Scale bars: 20 mm. Credits: B-D, P. Loubry MNHN/CNRS; G, H, J. Mouchart." figureDoi="http://doi.org/10.5281/zenodo.13985772" httpUri="https://zenodo.org/record/13985772/files/figure.png" pageId="17" pageNumber="510">Fig. 6A</figureCitation>
|
||
). It displays rounded wings, P1 spine is included in the wings and is not protruding. This morphology of the anterior part of the wings (
|
||
<figureCitation id="131C2A49FFF7E417FC43FE40FBD0FE02" box="[1019,1105,406,433]" captionStart="FIG" captionStartId="16.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-382@16.[698,1267,772,1332]" captionTargetPageId="16" captionText="FIG. 6. — Neotype, historical figures, spiral cords and denticles of the outer lip in T. tripteroides (Lamarck, 1822): A-D, morphotype with rounded varices and poorly distinguishable P1 spine; E-H, morphotype with easily distinguishable P1 spine: A, original figure of Lamarck (1805: pl. 3, fig. 4); B, C, neotype designated herein, MNHN.F.A90546 (Club géologique d'Île-de-France leg.), Thiverval-Grignon (Falunière), Yvelines, Paris Basin, France, middle Eocene (Lutetian); D, MNHN.F.A90552 (Pacaud coll.), Chaussy (Les Garennes),Paris Basin, France,middle Eocene (Lutetian); E, F, figured specimen of Deshayes (1835: pl. 82, fig. 1-2); G, H, figured specimen of Cossmann & Pissarro (1911: pl. 35, fig. 169-1; MNHN.F.J10350 (Cossmann coll.), Chaussy (Les Garennes). Scale bars: 20 mm. Credits: B-D, P. Loubry MNHN/CNRS; G, H, J. Mouchart." figureDoi="http://doi.org/10.5281/zenodo.13985772" httpUri="https://zenodo.org/record/13985772/files/figure.png" pageId="17" pageNumber="510">Fig. 6D</figureCitation>
|
||
) differs from that of the
|
||
<taxonomicName id="4C274D4FFFF7E417FAD5FE41FC21FE63" baseAuthorityName="Lamarck" baseAuthorityYear="1822" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="510" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF7E417FAD5FE41FC21FE63" italics="true" pageId="17" pageNumber="510">Murex tripteroides</emphasis>
|
||
</taxonomicName>
|
||
illustrated by
|
||
<bibRefCitation id="EFB64B3DFFF7E417FBFAFE60FB6EFE62" author="DESHAYES G. - P." box="[1090,1263,438,465]" pageId="17" pageNumber="510" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel & Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes (1835</bibRefCitation>
|
||
: pl. 82, figs 1, 2;
|
||
<figureCitation id="131C2A49FFF7E417FC96FE00FC16FE43" box="[814,919,470,497]" captionStart="FIG" captionStartId="16.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-382@16.[698,1267,772,1332]" captionTargetPageId="16" captionText="FIG. 6. — Neotype, historical figures, spiral cords and denticles of the outer lip in T. tripteroides (Lamarck, 1822): A-D, morphotype with rounded varices and poorly distinguishable P1 spine; E-H, morphotype with easily distinguishable P1 spine: A, original figure of Lamarck (1805: pl. 3, fig. 4); B, C, neotype designated herein, MNHN.F.A90546 (Club géologique d'Île-de-France leg.), Thiverval-Grignon (Falunière), Yvelines, Paris Basin, France, middle Eocene (Lutetian); D, MNHN.F.A90552 (Pacaud coll.), Chaussy (Les Garennes),Paris Basin, France,middle Eocene (Lutetian); E, F, figured specimen of Deshayes (1835: pl. 82, fig. 1-2); G, H, figured specimen of Cossmann & Pissarro (1911: pl. 35, fig. 169-1; MNHN.F.J10350 (Cossmann coll.), Chaussy (Les Garennes). Scale bars: 20 mm. Credits: B-D, P. Loubry MNHN/CNRS; G, H, J. Mouchart." figureDoi="http://doi.org/10.5281/zenodo.13985772" httpUri="https://zenodo.org/record/13985772/files/figure.png" pageId="17" pageNumber="510">Fig. 6E, F</figureCitation>
|
||
) and
|
||
<bibRefCitation id="EFB64B3DFFF7E417FC6CFE00FA84FE43" author="COSSMANN M. & PISSARRO G." box="[980,1285,470,496]" pageId="17" pageNumber="510" pagination="26 - 45" refId="ref43883" refString="COSSMANN M. & PISSARRO G. 1911. - Iconographie complete des coquilles fossiles de l'Eocene du Bassin de Paris. Tome 2. Hermann, Paris: pl. 26 - 45. https: // www. biodiversitylibrary. org / page / 36319938" type="book chapter" year="1911">Cossmann & Pissarro (1911</bibRefCitation>
|
||
: pl. 34, 169-1), which display a well individualized P1 spine (
|
||
<figureCitation id="131C2A49FFF7E417FA9CFE20FA23FDA3" box="[1316,1442,502,528]" captionStart="FIG" captionStartId="16.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-382@16.[698,1267,772,1332]" captionTargetPageId="16" captionText="FIG. 6. — Neotype, historical figures, spiral cords and denticles of the outer lip in T. tripteroides (Lamarck, 1822): A-D, morphotype with rounded varices and poorly distinguishable P1 spine; E-H, morphotype with easily distinguishable P1 spine: A, original figure of Lamarck (1805: pl. 3, fig. 4); B, C, neotype designated herein, MNHN.F.A90546 (Club géologique d'Île-de-France leg.), Thiverval-Grignon (Falunière), Yvelines, Paris Basin, France, middle Eocene (Lutetian); D, MNHN.F.A90552 (Pacaud coll.), Chaussy (Les Garennes),Paris Basin, France,middle Eocene (Lutetian); E, F, figured specimen of Deshayes (1835: pl. 82, fig. 1-2); G, H, figured specimen of Cossmann & Pissarro (1911: pl. 35, fig. 169-1; MNHN.F.J10350 (Cossmann coll.), Chaussy (Les Garennes). Scale bars: 20 mm. Credits: B-D, P. Loubry MNHN/CNRS; G, H, J. Mouchart." figureDoi="http://doi.org/10.5281/zenodo.13985772" httpUri="https://zenodo.org/record/13985772/files/figure.png" pageId="17" pageNumber="510">Fig. 6G, H</figureCitation>
|
||
). We examined numerous specimens from different localities in the Paris Basin and found transitional forms between the two morphotypes. Thus, we consider this difference to fall within the intraspecific variation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |