157 lines
18 KiB
XML
157 lines
18 KiB
XML
<document id="77E8A2E779222482CD42E0E26EC3562F" ID-CLB-Dataset="21707" ID-DOI="10.11646/zootaxa.4790.2.10" ID-GBIF-Dataset="3f492afe-37c4-4bc9-b7d1-5f8d804cf5bb" ID-ISSN="1175-5326" ID-Zenodo-Dep="3889408" ID-ZooBank="7F4A9B99-C12E-473B-B258-7BE5697F4986" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591859385935" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2020" docId="038E87D4FF85FFA2FF1579E1FBEF7179" docLanguage="en" docName="zootaxa.4790.2.10.pdf" docOrigin="Zootaxa 4790 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus latipennis Perkins 2020, new species" docType="treatment" docVersion="8" lastPageNumber="365" masterDocId="FFB7FFACFF81FFA5FF827F44FFB87066" masterDocTitle="Taxonomy of Ecuadorian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="374" masterPageNumber="358" pageNumber="362" updateTime="1698842324292" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="4EB6D9A225870E0CF2C0A1D3F03BAF8E">
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<mods:title id="5FC9F6444A4ECA28CF098CD8CA433F15">Taxonomy of Ecuadorian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<mods:name id="5580A59EDE6A9C7584A26C8A03BA6676" type="personal">
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<mods:namePart id="6ADAF8FC7E1F289FB499905E0E095DD4">Perkins, Philip D.</mods:namePart>
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<mods:title id="8CBEEC58FCB7C2E874DE1123C8003ED3">Zootaxa</mods:title>
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<mods:part id="A258C39506C8016FCE7F9A84FA640E19">
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<mods:date id="EF2BC962D24DD2994683063DECCCB56E">2020</mods:date>
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<mods:number id="A0C6B6D8A977FBA7A8299BC36BF8C4FF">2020-06-11</mods:number>
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<mods:number id="464545F304F2712F5212B686CEC51977">4790</mods:number>
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<treatment id="038E87D4FF85FFA2FF1579E1FBEF7179" ID-DOI="http://doi.org/10.5281/zenodo.5918720" ID-GBIF-Taxon="164501159" ID-Zenodo-Dep="5918720" LSID="urn:lsid:plazi:treatment:038E87D4FF85FFA2FF1579E1FBEF7179" httpUri="http://treatment.plazi.org/id/038E87D4FF85FFA2FF1579E1FBEF7179" lastPageId="7" lastPageNumber="365" pageId="4" pageNumber="362">
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<subSubSection id="C33D6549FF85FFA1FF1579E1FD8276D9" box="[151,570,1701,1728]" pageId="4" pageNumber="362" type="nomenclature">
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<paragraph id="8B9836C2FF85FFA1FF1579E1FD8276D9" blockId="4.[151,570,1701,1763]" box="[151,570,1701,1728]" pageId="4" pageNumber="362">
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<heading id="D0D081AEFF85FFA1FF1579E1FD8276D9" bold="true" box="[151,570,1701,1728]" fontSize="11" level="1" pageId="4" pageNumber="362" reason="1">
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<emphasis id="B953EAD0FF85FFA1FF1579E1FD8276D9" bold="true" box="[151,570,1701,1728]" pageId="4" pageNumber="362">
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<taxonomicName id="4C274D41FF85FFA1FF1579E1FE2376A6" ID-CoL="8Y4QG" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[151,411,1701,1728]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="362" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
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<emphasis id="B953EAD0FF85FFA1FF1579E1FE2376A6" bold="true" box="[151,411,1701,1728]" italics="true" pageId="4" pageNumber="362">Hydrochus latipennis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26057ABFF85FFA1FE2879E1FD8276D9" box="[426,570,1701,1727]" pageId="4" pageNumber="362" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D6549FF85FFA1FF15798DFF477685" box="[151,255,1737,1763]" pageId="4" pageNumber="362" type="description">
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<paragraph id="8B9836C2FF85FFA1FF15798DFF477685" blockId="4.[151,570,1701,1763]" box="[151,255,1737,1763]" pageId="4" pageNumber="362">
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<figureCitation id="131C2A47FF85FFA1FF15798DFF5B7685" box="[151,227,1737,1763]" captionStart="FIGURE 1" captionStartId="2.[151,250,2019,2044]" captionTargetBox="[192,1405,448,1963]" captionTargetId="figure@2.[151,1435,417,1993]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3889410" httpUri="https://zenodo.org/record/3889410/files/figure.png" pageId="4" pageNumber="362">Figs. 1</figureCitation>
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,
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<figureCitation id="131C2A47FF85FFA1FF73798DFF477685" box="[241,255,1737,1763]" captionStart="FIGURE 4" captionStartId="6.[151,250,1617,1642]" captionTargetBox="[186,1411,197,1556]" captionTargetId="figure@6.[151,1435,166,1598]" captionTargetPageId="6" captionText="FIGURE 4. Hydrochus latipennis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889416" httpUri="https://zenodo.org/record/3889416/files/figure.png" pageId="4" pageNumber="362">4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D6549FF85FFA1FF157855FAD6774D" box="[151,1390,1808,1836]" pageId="4" pageNumber="362" type="materials_examined">
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<paragraph id="8B9836C2FF85FFA1FF157855FAD6774D" blockId="4.[151,1436,1808,1944]" box="[151,1390,1808,1836]" pageId="4" pageNumber="362">
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<emphasis id="B953EAD0FF85FFA1FF157855FEF0774D" bold="true" box="[151,328,1808,1835]" pageId="4" pageNumber="362">Type Material.</emphasis>
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<materialsCitation id="3B4F3C9FFF85FFA1FECD7855FAD6774D" ID-GBIF-Occurrence="2632886325" box="[335,1390,1809,1836]" collectingDate="1977-06-14" collectionCode="NMNH" collectorName="W. E. Steiner" country="Ecuador" location="Limoncocha" pageId="4" pageNumber="362" specimenCount="1" stateProvince="Napo" typeStatus="holotype">
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<typeStatus id="549C8860FF85FFA1FECD7855FE0F774D" box="[335,439,1809,1835]" pageId="4" pageNumber="362" type="holotype">Holotype</typeStatus>
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(male): “
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<emphasis id="B953EAD0FF85FFA1FDA07855FD31774D" bold="true" box="[546,649,1809,1835]" pageId="4" pageNumber="362">
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<collectingCountry id="F3307652FF85FFA1FDA07855FD31774D" box="[546,649,1809,1835]" name="Ecuador" pageId="4" pageNumber="362">Ecuador</collectingCountry>
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</emphasis>
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:
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<collectingRegion id="49E3F820FF85FFA1FD1A7855FD6F774D" box="[664,727,1809,1835]" country="Ecuador" name="Napo" pageId="4" pageNumber="362">Napo</collectingRegion>
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,
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<location id="8EF86019FF85FFA1FD617855FCCA774D" LSID="urn:lsid:plazi:treatment:038E87D4FF85FFA2FF1579E1FBEF7179:8EF86019FF85FFA1FD617855FCCA774D" box="[739,882,1809,1835]" country="Ecuador" name="Limoncocha" pageId="4" pageNumber="362" stateProvince="Napo">Limoncocha</location>
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,
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<date id="FF991002FF85FFA1FCFF7855FBAA774D" box="[893,1042,1809,1835]" pageId="4" pageNumber="362" value="1977-06-14">
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<collectingDate id="EFDDE9EAFF85FFA1FCFF7855FBAA774D" box="[893,1042,1809,1835]" pageId="4" pageNumber="362" value="1977-06-14">14 June 1977</collectingDate>
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</date>
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BLT.
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<collectorName id="26D25314FF85FFA1FBD57855FB56774D" box="[1111,1262,1809,1835]" pageId="4" pageNumber="362">W. E. Steiner</collectorName>
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” (
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<collectionCode id="ED36AE07FF85FFA1FA847855FADA774D" box="[1286,1378,1809,1835]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="362" type="Museum">NMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D6549FF85FFA1FF457870FD3E77FE" pageId="4" pageNumber="362" type="diagnosis">
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<paragraph id="8B9836C2FF85FFA1FF457870FD3E77FE" blockId="4.[151,1436,1808,1944]" pageId="4" pageNumber="362">
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<emphasis id="B953EAD0FF85FFA1FF457870FE6E7729" bold="true" box="[199,470,1844,1871]" pageId="4" pageNumber="362">Differential Diagnosis.</emphasis>
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Among Ecuadorian species, recognized by the combination of small size (ca.
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<quantity id="4CDF9B27FF85FFA1FAE97870FF7F7715" metricMagnitude="-3" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="362" unit="mm" value="2.54">2.54 mm</quantity>
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), the shallow pronotal impressions, the wide elytra with arcuate lateral margins and large serial punctures, and the male genitalia (
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<figureCitation id="131C2A47FF85FFA1FEEC7839FE0B77F1" box="[366,435,1917,1943]" captionStart="FIGURE 4" captionStartId="6.[151,250,1617,1642]" captionTargetBox="[186,1411,197,1556]" captionTargetId="figure@6.[151,1435,166,1598]" captionTargetPageId="6" captionText="FIGURE 4. Hydrochus latipennis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889416" httpUri="https://zenodo.org/record/3889416/files/figure.png" pageId="4" pageNumber="362">Fig. 4</figureCitation>
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; described below).
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</paragraph>
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</subSubSection>
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<caption id="DF58664AFF84FFA0FF1579C3FC0376C6" ID-DOI="http://doi.org/10.5281/zenodo.3889414" ID-Zenodo-Dep="3889414" box="[151,955,1671,1696]" httpUri="https://zenodo.org/record/3889414/files/figure.png" pageId="5" pageNumber="363" startId="5.[151,250,1671,1696]" targetBox="[191,1423,216,1604]" targetPageId="5">
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<paragraph id="8B9836C2FF84FFA0FF1579C3FC0376C6" blockId="5.[151,955,1671,1696]" box="[151,955,1671,1696]" pageId="5" pageNumber="363">
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<emphasis id="B953EAD0FF84FFA0FF1579C3FEAB76C6" bold="true" box="[151,275,1671,1696]" pageId="5" pageNumber="363">FIGURE 3.</emphasis>
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<taxonomicName id="4C274D41FF84FFA0FE9B79C3FE5C76F9" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[281,484,1671,1695]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="363" phylum="Arthropoda" rank="species" species="hamulus" status="sp. nov.">
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<emphasis id="B953EAD0FF84FFA0FE9B79C3FE5C76F9" box="[281,484,1671,1695]" italics="true" pageId="5" pageNumber="363">Hydrochus hamulus</emphasis>
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</taxonomicName>
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<emphasis id="B953EAD0FF84FFA0FE6879CCFD9B76C6" bold="true" box="[490,547,1672,1696]" pageId="5" pageNumber="363">
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<taxonomicNameLabel id="A26057ABFF84FFA0FE6879CCFD9B76C6" box="[490,547,1672,1696]" pageId="5" pageNumber="363" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, habitus and male genitalia of holotype.
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</paragraph>
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</caption>
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<subSubSection id="C33D6549FF84FFA2FF457989FEC5709D" lastPageId="7" lastPageNumber="365" pageId="5" pageNumber="363" type="description">
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<paragraph id="8B9836C2FF84FFA0FF457989FC68771E" blockId="5.[151,1437,1741,1984]" pageId="5" pageNumber="363">
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<emphasis id="B953EAD0FF84FFA0FF457989FEE9768E" bold="true" box="[199,337,1741,1768]" pageId="5" pageNumber="363">Description</emphasis>
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. Habitus as illustrated (
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<figureCitation id="131C2A47FF84FFA0FDDF798AFD1B768E" box="[605,675,1742,1768]" captionStart="FIGURE 4" captionStartId="6.[151,250,1617,1642]" captionTargetBox="[186,1411,197,1556]" captionTargetId="figure@6.[151,1435,166,1598]" captionTargetPageId="6" captionText="FIGURE 4. Hydrochus latipennis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889416" httpUri="https://zenodo.org/record/3889416/files/figure.png" pageId="5" pageNumber="363">Fig. 4</figureCitation>
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). Size:
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<typeStatus id="549C8860FF84FFA0FD75798AFCE1768E" box="[759,857,1742,1768]" pageId="5" pageNumber="363" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 2.54/0.94; head width 0.62; pronotum l/w 0.60/0.60; PA 0.56; PB 0.48; elytra 1.54/0.94. Body size small (ca.
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<quantity id="4CDF9B27FF84FFA0FAE979B5FF7F7756" metricMagnitude="-3" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="363" unit="mm" value="2.54">2.54 mm</quantity>
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). Dorsum black, with slight iridescence, elytra without black spots. Punctation on elytra ca. 2–4x that of pronotum. Elytra interstriae narrow walls to ca. 0.5x strial puncture diameter. Interstria 9
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<superScript id="7C529B8AFF84FFA0FBE5787DFBCC7721" attach="left" box="[1127,1140,1849,1863]" fontSize="6" pageId="5" pageNumber="363">th</superScript>
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more convex than others, overhanging 10
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<superScript id="7C529B8AFF84FFA0FEC47819FEEB770D" attach="left" box="[326,339,1885,1899]" fontSize="6" pageId="5" pageNumber="363">th</superScript>
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interstria. Usual area of callus on 5
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<superScript id="7C529B8AFF84FFA0FD637819FD56770D" attach="left" box="[737,750,1885,1899]" fontSize="6" pageId="5" pageNumber="363">th</superScript>
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very weakly raised.
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</paragraph>
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<paragraph id="8B9836C2FF84FFA0FF4578C6FB0C77A6" blockId="5.[151,1437,1741,1984]" pageId="5" pageNumber="363">Pronotum length and width equal, widest slightly behind anterior margin, narrowed at base, sides arcuate or very weakly sinuate, smooth; anterior margin slightly bisinuate; depressions moderately deep.</paragraph>
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<caption id="DF58664AFF87FFA3FF157915FC71760C" ID-DOI="http://doi.org/10.5281/zenodo.3889416" ID-Zenodo-Dep="3889416" box="[151,969,1617,1642]" httpUri="https://zenodo.org/record/3889416/files/figure.png" pageId="6" pageNumber="364" startId="6.[151,250,1617,1642]" targetBox="[186,1411,197,1556]" targetPageId="6">
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<paragraph id="8B9836C2FF87FFA3FF157915FC71760C" blockId="6.[151,969,1617,1642]" box="[151,969,1617,1642]" pageId="6" pageNumber="364">
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<emphasis id="B953EAD0FF87FFA3FF157915FEAB760C" bold="true" box="[151,275,1617,1642]" pageId="6" pageNumber="364">FIGURE 4.</emphasis>
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<taxonomicName id="4C274D41FF87FFA3FE9B7915FE4A760F" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[281,498,1617,1641]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="latipennis" status="sp. nov.">
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<emphasis id="B953EAD0FF87FFA3FE9B7915FE4A760F" box="[281,498,1617,1641]" italics="true" pageId="6" pageNumber="364">Hydrochus latipennis</emphasis>
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</taxonomicName>
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<emphasis id="B953EAD0FF87FFA3FE7A7916FD89760C" bold="true" box="[504,561,1618,1642]" pageId="6" pageNumber="364">
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<taxonomicNameLabel id="A26057ABFF87FFA3FE7A7916FD89760C" box="[504,561,1618,1642]" pageId="6" pageNumber="364" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, habitus and male genitalia of holotype.
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</paragraph>
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</caption>
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<paragraph id="8B9836C2FF87FFA3FF4579DCFEAD769C" blockId="6.[151,1437,1688,2038]" pageId="6" pageNumber="364">Elytra widest at ca. posterior 1/3, where lateral margin notably arcuate; spaces between strial punctures narrow walls to ca. 0.5x their diameter; apices separately rounded in dorsal view, in lateral view outer margin without angulation.</paragraph>
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<paragraph id="8B9836C2FF87FFA3FF457840FA9F7724" blockId="6.[151,1437,1688,2038]" pageId="6" pageNumber="364">Ventral characters: Mentum coarsely punctate, with large and deep median fovea. Submentum with two large foveae, separated by narrow midlongitudinal ridge. Legs brown to testaceous, femora darker than tibiae.</paragraph>
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<paragraph id="8B9836C2FF87FFA3FF457808FB477790" blockId="6.[151,1437,1688,2038]" pageId="6" pageNumber="364">
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Male genitalia (
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<figureCitation id="131C2A47FF87FFA3FEFF7808FE7B7700" box="[381,451,1868,1894]" captionStart="FIGURE 4" captionStartId="6.[151,250,1617,1642]" captionTargetBox="[186,1411,197,1556]" captionTargetId="figure@6.[151,1435,166,1598]" captionTargetPageId="6" captionText="FIGURE 4. Hydrochus latipennis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889416" httpUri="https://zenodo.org/record/3889416/files/figure.png" pageId="6" pageNumber="364">Fig. 4</figureCitation>
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) general characters: genitalia small; paramere tips moderately large, each as wide, or almost as wide, as widest part of aedeagus, very lightly sclerotized, tips rounded and curving toward midline, but not contacting one another, tips wider on ventral than on dorsal surface, on dorsal surface pdmm arcuate away from midline, parameres more evidently sclerotized for short distance just proximal to tips; aedeagus widest at about distal ¼; basal piece as long as parameres, widest at about distal 1/3 to ¼, orifice weakly sclerotized.
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</paragraph>
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<paragraph id="8B9836C2FF86FFA2FF457FDDFD0770D5" blockId="7.[151,1436,153,287]" box="[199,703,153,179]" pageId="7" pageNumber="365">Dorsal surface: adtl lacking; adbl very small.</paragraph>
|
||
<paragraph id="8B9836C2FF86FFA2FF457FF9FEC5709D" blockId="7.[151,1436,153,287]" pageId="7" pageNumber="365">Ventral surface: pvmm slightly overlapping basal part of aedeagus; pseudoparameres extending to about ¾ length of parameres.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6549FF86FFA2FF457E41FBEF7179" box="[199,1111,261,287]" pageId="7" pageNumber="365" type="etymology">
|
||
<paragraph id="8B9836C2FF86FFA2FF457E41FBEF7179" blockId="7.[151,1436,153,287]" box="[199,1111,261,287]" pageId="7" pageNumber="365">
|
||
<emphasis id="B953EAD0FF86FFA2FF457E41FEF67179" bold="true" box="[199,334,261,287]" pageId="7" pageNumber="365">Etymology.</emphasis>
|
||
Named in reference to the wide, arcuate lateral margins of the elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |