treatments-xml/data/03/8E/87/038E87BEFFC3FFE7FF10FDF3994CFA13.xml

<document id="D633714CCF65469107D5C1658280D4D6" ID-CLB-Dataset="22471" ID-DOI="10.5281/zenodo.3776456" ID-GBIF-Dataset="5dd36ea3-7c58-4644-a362-7dcac8e05a15" ID-ISSN="0253-116X" ID-Zenodo-Dep="3776456" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588185290416" checkinUser="valdenar" docAuthor="Assing, Volker" docDate="2018" docId="038E87BEFFC3FFE7FF10FDF3994CFA13" docLanguage="en" docName="Linzer.biol.Beitr.50.2.969-976.pdf.imf" docOrigin="Linzer biologische Beiträge 50 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Caucanthobium mirandum Assing 2018, nov.sp." docType="treatment" docUuid="E9B63742-8A55-44B6-B976-1B6C4A6DB1E0" docUuidSource="ZooBank" docVersion="6" lastPageNumber="975" masterDocId="FFB7FFC6FFC6FFE1FFA2FF919A37FFCC" masterDocTitle="An enigmatic new genus and species of Anthophagini from Georgia (Coleoptera: Staphylinidae: Omaliinae)" masterLastPageNumber="976" masterPageNumber="969" pageNumber="974" updateTime="1698838069146" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="E618A739DFE7FEC1EF643CB442DC9A5F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3C9381DDA98F2EDDF1E6406A395F164E">
<mods:title id="32ED5CB0A244B9E70273F7D42EF5E82D">An enigmatic new genus and species of Anthophagini from Georgia (Coleoptera: Staphylinidae: Omaliinae)</mods:title>
</mods:titleInfo>
<mods:name id="A173FC9FE9E192183986E49E6A91AFFD" type="personal">
<mods:role id="092B5E43BFB42BC6D803EB6C965BD96B">
<mods:roleTerm id="3809019482974B2963E6DEB0E7AA9B0D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="534B5FB2E1DAA8F5A9DD2B6F289041BC">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="4CCC7785C05FC3E7DC444FEA910C1E45">text</mods:typeOfResource>
<mods:relatedItem id="C97166594E8C1799DD31105A552731F7" type="host">
<mods:titleInfo id="6802D1B80B12DB56FEA0F2346C8004C7">
<mods:title id="290A801EAD9B5DB728E862A25F1DE618">Linzer biologische Beiträge</mods:title>
</mods:titleInfo>
<mods:part id="71A2DE5B79D90C4264F71127D27F3766">
<mods:date id="C43357A61C98385E3D1473A12954464A">2018</mods:date>
<mods:detail id="D38F56C3FFC5B54144181D17C735A9AE" type="pubDate">
<mods:number id="06007ECCF5887CC058678D08715A0C1F">2018-12-17</mods:number>
</mods:detail>
<mods:detail id="D71E2A6035E6A318E50C612AD435CB8B" type="volume">
<mods:number id="C88D8861ED8E042278CC5FD26D4DDCE8">50</mods:number>
</mods:detail>
<mods:detail id="BAF7BD7E07C53E818812A56CF7DDF08C" type="issue">
<mods:number id="99E64A297089C3BD14055BC00202EA2E">2</mods:number>
</mods:detail>
<mods:extent id="3E4894C6C765FE5F3E2CC5ED7F7C5DD9" unit="page">
<mods:start id="4CC8E0225EA1D17B856B1856065CE956">969</mods:start>
<mods:end id="9E4B26896DA5FF46864987E3E118EFC1">976</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="722E4A0B90D256979DBB385E48D57361">journal article</mods:classification>
<mods:identifier id="4321CAC50C27E178BAF535C7E9F787EF" type="CLB-Dataset">22471</mods:identifier>
<mods:identifier id="98B6BC33C964B12A01F270E94C3A1DD9" type="DOI">10.5281/zenodo.3776456</mods:identifier>
<mods:identifier id="990202F6B59D4E3B9DADCB83D6D9B547" type="GBIF-Dataset">5dd36ea3-7c58-4644-a362-7dcac8e05a15</mods:identifier>
<mods:identifier id="6EDEFC85F5544591464C9CDDF0A2DDD8" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="D2BD92AB6197C5938CB6BB992AE14664" type="Zenodo-Dep">3776456</mods:identifier>
<mods:identifier id="250B64E2E3BBC0BFF24606037D90863C" type="ZooBank">65D37C1A-778E-47C5-85FC-DDA7FCC25F9C</mods:identifier>
</mods:mods>
<treatment id="038E87BEFFC3FFE7FF10FDF3994CFA13" ID-DOI="http://doi.org/10.5281/zenodo.3796205" ID-GBIF-Taxon="163876804" ID-Zenodo-Dep="3796205" LSID="urn:lsid:zoobank.org:act:E9B63742-8A55-44B6-B976-1B6C4A6DB1E0" httpUri="http://treatment.plazi.org/id/038E87BEFFC3FFE7FF10FDF3994CFA13" lastPageId="6" lastPageNumber="975" pageId="5" pageNumber="974">
<subSubSection id="C33D6523FFC3FFE4FF10FDF39841FCF7" pageId="5" pageNumber="974" type="nomenclature">
<paragraph id="8B9836A8FFC3FFE4FF10FDF398DAFDB4" blockId="5.[178,749,610,632]" box="[178,749,610,632]" pageId="5" pageNumber="974">
<heading id="D0D081C4FFC3FFE4FF10FDF398DAFDB4" box="[178,749,610,632]" fontSize="8" level="2" pageId="5" pageNumber="974" reason="2">
<emphasis id="B953EABAFFC3FFE4FF10FDF398DAFDB4" bold="true" box="[178,749,610,632]" pageId="5" pageNumber="974">
<taxonomicName id="4C274D2BFFC3FFE4FF10FDF39BF8FDB4" ID-CoL="8XWRW" authority="Assing, 2018" authorityName="Assing" authorityYear="2018" box="[178,463,610,632]" class="Insecta" family="Staphylinidae" genus="Caucanthobium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="974" phylum="Arthropoda" rank="species" species="mirandum" status="sp. nov.">
<emphasis id="B953EABAFFC3FFE4FF10FDF39BF8FDB4" bold="true" box="[178,463,610,632]" italics="true" pageId="5" pageNumber="974">Caucanthobium mirandum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057C1FFC3FFE4FE77FDF39816FDBB" box="[469,545,610,631]" pageId="5" pageNumber="974" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="131C2A2DFFC3FFE4FD8CFDF398A4FDBB" box="[558,659,610,632]" captionStart-0="Figs 1-3" captionStart-1="Figs 4-8" captionStart-2="Figs 9-11" captionStartId-0="2.[178,218,1506,1528]" captionStartId-1="3.[178,218,1514,1536]" captionStartId-2="4.[178,218,896,918]" captionTargetBox-0="[192,1076,210,1483]" captionTargetBox-1="[182,1089,212,1497]" captionTargetBox-2="[189,1085,204,856]" captionTargetId-0="figure@2.[179,1086,201,1486]" captionTargetId-1="figure@3.[182,1089,212,1497]" captionTargetId-2="figure@4.[179,1086,203,878]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figs 1-3: Caucanthobium mirandum nov.sp.: (1) habitus; (2) antenna; (3) male protarsus. Scale bars: 1-2: 1.0 mm; 3: 0.2 mm." captionText-1="Figs 4-8: Caucanthobium mirandum nov.sp.: (4) head and pronotum; (5) head in dorso-lateral view; (6) head in ventral view; (7) thorax in ventral view; (8) apical portion of abdomen. Scale bars: 7: 1.0 mm; 4-6, 8: 0.5 mm." captionText-2="Figs 9-11: Caucanthobium mirandum nov.sp.: (9-10) aedeagus in lateral and in ventral view; (11) female genital segment. Scale bars: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3776458" figureDoi-1="http://doi.org/10.5281/zenodo.3776460" figureDoi-2="http://doi.org/10.5281/zenodo.3776462" httpUri-0="https://zenodo.org/record/3776458/files/figure.png" httpUri-1="https://zenodo.org/record/3776460/files/figure.png" httpUri-2="https://zenodo.org/record/3776462/files/figure.png" pageId="5" pageNumber="974">Figs 1-11</figureCitation>
, 
<figureCitation id="131C2A2DFFC3FFE4FD3DFDF398D3FDB4" box="[671,740,610,632]" captionStart="Map 1" captionStartId="6.[178,224,1224,1245]" captionTargetBox="[178,1086,722,1201]" captionTargetId="figure@6.[178,1086,722,1201]" captionTargetPageId="6" captionText="Map 1: Distribution of Caucanthobium mirandum." figureDoi="http://doi.org/10.5281/zenodo.3776464" httpUri="https://zenodo.org/record/3776464/files/figure.png" pageId="5" pageNumber="974">Map 1</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9836A8FFC3FFE4FF10FD169841FCF7" blockId="5.[178,1085,647,827]" pageId="5" pageNumber="974">
<materialsCitation id="3B4F3CF5FFC3FFE4FF10FD169B44FD06" ID-GBIF-Occurrence="2609444603" collectingDate="2018-05-24" collectionCode="T, V" country="Georgia" elevation="1280" location="Gobroneti" longitude="41.671112" pageId="5" pageNumber="974" specimenCount="2" specimenCount-male="2" stateProvince="Adjara" typeStatus="holotype">
<collectionCode id="ED36AE6DFFC3FFE4FF10FD169AF7FD50" box="[178,192,647,668]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="974">T</collectionCode>
y p e m a t e r i a l: 
<typeStatus id="549C880AFFC3FFE4FE05FD169BCCFD51" box="[423,507,647,669]" pageId="5" pageNumber="974" type="holotype">Holotype</typeStatus>
<specimenCount id="9D21FD21FFC3FFE4FDA6FD16982FFD50" box="[516,536,647,668]" pageId="5" pageNumber="974" type="male">♂</specimenCount>
: &quot; 
<geoCoordinate id="EE13506FFFC3FFE4FD8FFD1698CDFD50" box="[557,762,647,669]" pageId="5" pageNumber="974">N41°40'16 E42°02'37</geoCoordinate>
, 
<collectingCountry id="F3307638FFC3FFE4FCA5FD16996CFD51" box="[775,859,647,669]" name="Georgia" pageId="5" pageNumber="974">Georgien</collectingCountry>
<collectingRegion id="49E3F84AFFC3FFE4FCC7FD169994FD51" box="[869,931,647,669]" country="Georgia" name="Ajaria" pageId="5" pageNumber="974">Adjara</collectingRegion>
(62), 
<location id="8EF86073FFC3FFE4FC43FD169E0AFD51" LSID="urn:lsid:plazi:treatment:038E87BEFFC3FFE7FF10FDF3994CFA13:8EF86073FFC3FFE4FC43FD169E0AFD51" box="[993,1085,647,669]" country="Georgia" longitude="41.671112" name="Gobroneti" pageId="5" pageNumber="974" stateProvince="Adjara">Gobroneti</location>
<quantity id="4CDF9B4DFFC3FFE4FF10FD0C9ACCFD7F" box="[178,251,669,691]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="5" pageNumber="974" unit="m" value="1280.0">
<elevation id="000AD19BFFC3FFE4FF10FD0C9ACCFD7F" box="[178,251,669,691]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="5" pageNumber="974" unit="m" value="1280.0">1280 m</elevation>
</quantity>
, 
<date id="FF991068FFC3FFE4FEA4FD0C9B55FD7F" box="[262,354,669,691]" pageId="5" pageNumber="974" value="2018-05-24">
<collectingDate id="EFDDE980FFC3FFE4FEA4FD0C9B55FD7F" box="[262,354,669,691]" pageId="5" pageNumber="974" value="2018-05-24">24.5.2018</collectingDate>
</date>
, Brachat &amp; Meybohm / 
<typeStatus id="549C880AFFC3FFE4FDEAFD0C9891FD7F" box="[584,678,669,691]" pageId="5" pageNumber="974" type="holotype">Holotypus</typeStatus>
<specimenCount id="9D21FD21FFC3FFE4FD0CFD0F9888FD7F" box="[686,703,670,691]" pageId="5" pageNumber="974" type="male">♂</specimenCount>
<taxonomicName id="4C274D2BFFC3FFE4FD64FD0C998BFD7E" authority="Assing, 2018" authorityName="Assing" authorityYear="2018" box="[710,956,669,690]" class="Insecta" family="Staphylinidae" genus="Caucanthobium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="974" phylum="Arthropoda" rank="species" species="mirandum" status="sp. nov.">
<emphasis id="B953EABAFFC3FFE4FD64FD0C998BFD7E" box="[710,956,669,690]" italics="true" pageId="5" pageNumber="974">Caucanthobium mirandum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057C1FFC3FFE4FC66FD0F99C0FD7F" box="[964,1015,670,691]" pageId="5" pageNumber="974" rank="species">sp. n.</taxonomicNameLabel>
det. 
<collectionCode id="ED36AE6DFFC3FFE4FB85FD0F9E0DFD7F" box="[1063,1082,670,691]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="974" type="Museum">V</collectionCode>
. Assing 2018&quot; (cAss)
</materialsCitation>
. 
<materialsCitation id="3B4F3CF5FFC3FFE4FEDFFD259984FD3B" ID-GBIF-Occurrence="2609444602" collectingDate="2018-06-05" collectingDateMax="2018-06-08" collectingDateMin="2018-06-05" collectionCode="NHMD, ZMUC" collectorName="A. K. Hansen" country="Georgia" elevation="1950" latitude="41.888725" location="Chokhatauri" longLatPrecision="1" longitude="42.361164" municipality="Chokhatauri" pageId="5" pageNumber="974" specimenCount="1" specimenCount-female="1" stateProvince="Guria" typeStatus="paratype">
<typeStatus id="549C880AFFC3FFE4FEDFFD259BEEFD05" box="[381,473,692,713]" pageId="5" pageNumber="974" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D21FD21FFC3FFE4FE40FD259BC9FD05" box="[482,510,692,713]" pageId="5" pageNumber="974" type="female">1♀</specimenCount>
[in ethanol]: &quot;AKH-GE18-2f, 
<collectingCountry id="F3307638FFC3FFE4FCB9FD259953FD06" box="[795,868,692,714]" name="Georgia" pageId="5" pageNumber="974">Georgia</collectingCountry>
<collectingRegion id="49E3F84AFFC3FFE4FCC8FD259997FD06" box="[874,928,692,714]" country="Georgia" name="Guria" pageId="5" pageNumber="974">Guria</collectingRegion>
, 
<collectingMunicipality id="6BFCACD2FFC3FFE4FC09FD259E2AFD06" box="[939,1053,692,714]" pageId="5" pageNumber="974">Chokhatauri</collectingMunicipality>
, 
<locationDeviation id="9A2A2F8AFFC3FFE4FB85FD259B18FD2D" location="Bakhmaro" pageId="5" pageNumber="974">W of Bakhmaro</locationDeviation>
, 
<geoCoordinate id="EE13506FFFC3FFE4FE98FD5B9B80FD2C" box="[314,439,714,736]" degrees="41" direction="north" minutes="50" orientation="latitude" pageId="5" pageNumber="974" precision="1" seconds="50.0" value="41.84722">41°50'50.0''N</geoCoordinate>
, 
<geoCoordinate id="EE13506FFFC3FFE4FE61FD5A980AFD2C" box="[451,573,715,736]" degrees="42" direction="east" minutes="17" orientation="longitude" pageId="5" pageNumber="974" precision="1" seconds="10.2" value="42.286167">42°17'10.2''E</geoCoordinate>
, 
<date id="FF991068FFC3FFE4FDE8FD5B989FFD2C" box="[586,680,714,736]" pageId="5" pageNumber="974" value="2018-06-05">
<collectingDate id="EFDDE980FFC3FFE4FDE8FD5B989FFD2C" box="[586,680,714,736]" pageId="5" pageNumber="974" value="2018-06-05">5.VI.2018</collectingDate>
</date>
, h 
<quantity id="4CDF9B4DFFC3FFE4FD62FD5B9936FD2C" box="[704,769,714,736]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="5" pageNumber="974" unit="m" value="1950.0">
<elevation id="000AD19BFFC3FFE4FD62FD5B9936FD2C" box="[704,769,714,736]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="5" pageNumber="974" unit="m" value="1950.0">1950m</elevation>
</quantity>
, sifting alder litter, grazed fields, rhododendron, scrubs, and forest patches, leg. 
<collectorName id="26D2537EFFC3FFE4FDD9FD7098C1FD3B" box="[635,758,737,759]" pageId="5" pageNumber="974">A.K. Hansen</collectorName>
<collectionCode id="ED36AE6DFFC3FFE4FCA0FD70997EFD3A" box="[770,841,737,758]" collectionName="NHMD" pageId="5" pageNumber="974">NHMD</collectionCode>
&quot; (
<collectionCode id="ED36AE6DFFC3FFE4FCC0FD70999EFD3B" box="[866,937,737,759]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="5" pageNumber="974">ZMUC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B4F3CF5FFC3FFE4FC1CFD709845FCF7" ID-GBIF-Occurrence="2609444601" collectingDate="2018-06-05" collectingDateMax="2018-06-08" collectingDateMin="2018-06-05" collectionCode="NHMD, ZMUC" collectorName="A. K. Hansen &amp; Pontic" country="Georgia" elevation="1950" latitude="41.888725" location="Chokhatauri" longLatPrecision="1" longitude="42.361164" municipality="Chokhatauri" pageId="5" pageNumber="974" specimenCount="3" specimenCount-female="3" stateProvince="Guria" typeStatus="paratype">
<specimenCount id="9D21FD21FFC3FFE4FC1CFD7099DBFD3A" box="[958,1004,737,758]" pageId="5" pageNumber="974" type="female">2♀♀</specimenCount>
[one in ethanol]: &quot;AKH-GE18-5b, 
<collectingCountry id="F3307638FFC3FFE4FE67FD699839FCC2" box="[453,526,760,782]" name="Georgia" pageId="5" pageNumber="974">Georgia</collectingCountry>
<collectingRegion id="49E3F84AFFC3FFE4FD80FD69986FFCC2" box="[546,600,760,782]" country="Georgia" name="Guria" pageId="5" pageNumber="974">Guria</collectingRegion>
, 
<location id="8EF86073FFC3FFE4FDCDFD6998D6FCC2" LSID="urn:lsid:plazi:treatment:038E87BEFFC3FFE7FF10FDF3994CFA13:8EF86073FFC3FFE4FDCDFD6998D6FCC2" box="[623,737,760,782]" country="Georgia" latitude="41.888725" longLatPrecision="1" longitude="42.361164" municipality="Chokhatauri" name="Chokhatauri" pageId="5" pageNumber="974" stateProvince="Guria">Chokhatauri</location>
, 
<locationDeviation id="9A2A2F8AFFC3FFE4FD5BFD699992FCC2" box="[761,933,760,782]" location="Bakhmaro" pageId="5" pageNumber="974">N of Bakhmaro</locationDeviation>
, 
<geoCoordinate id="EE13506FFFC3FFE4FC1EFD699E0FFCC2" box="[956,1080,760,782]" degrees="41" direction="north" minutes="53" orientation="latitude" pageId="5" pageNumber="974" precision="1" seconds="19.4" value="41.888725">41°53'19.4''N</geoCoordinate>
, 
<geoCoordinate id="EE13506FFFC3FFE4FF10FC9E9B1BFCE8" box="[178,300,783,804]" degrees="42" direction="east" minutes="21" orientation="longitude" pageId="5" pageNumber="974" precision="1" seconds="40.2" value="42.361164">42°21'40.2''E</geoCoordinate>
, 
<date id="FF991068FFC3FFE4FE98FC9E9BAFFCE8" box="[314,408,783,804]" pageId="5" pageNumber="974" value="2018-06-08">
<collectingDate id="EFDDE980FFC3FFE4FE98FC9E9BAFFCE8" box="[314,408,783,804]" pageId="5" pageNumber="974" value="2018-06-08">8.VI.2018</collectingDate>
</date>
, h 
<quantity id="4CDF9B4DFFC3FFE4FE13FC9E9BC5FCE9" box="[433,498,783,805]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="5" pageNumber="974" unit="m" value="1640.0">
<elevation id="000AD19BFFC3FFE4FE13FC9E9BC5FCE9" box="[433,498,783,805]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="5" pageNumber="974" unit="m" value="1640.0">1640m</elevation>
</quantity>
, sifting oak litter and mouse nest in hollow tree, 
<collectorName id="26D2537EFFC3FFE4FC7AFC9E9E25FCE9" box="[984,1042,783,805]" pageId="5" pageNumber="974">Pontic</collectorName>
oak forest, leg. 
<collectorName id="26D2537EFFC3FFE4FEBAFCB49BBAFCF7" box="[280,397,805,827]" pageId="5" pageNumber="974">A.K. Hansen</collectorName>
<collectionCode id="ED36AE6DFFC3FFE4FE31FCB49BEDFCF6" box="[403,474,805,826]" pageId="5" pageNumber="974">NHMD</collectionCode>
&quot; (
<collectionCode id="ED36AE6DFFC3FFE4FE4FFCB49805FCF6" box="[493,562,805,826]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="5" pageNumber="974">ZMUC</collectionCode>
, cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6523FFC3FFE7FF10FCDB994CFA13" lastPageId="6" lastPageNumber="975" pageId="5" pageNumber="974" type="description">
<paragraph id="8B9836A8FFC3FFE4FF10FCDB9E3DFCB1" blockId="5.[178,1084,841,893]" pageId="5" pageNumber="974">E t y m o l o g y: The specific epithet (Latin, gerundivum of the verb mirare: to be admired) alludes to the conspicuous appearance (size and coloration) of this species.</paragraph>
<paragraph id="8B9836A8FFC3FFE4FF10FC1D9812FC73" blockId="5.[178,1085,908,959]" pageId="5" pageNumber="974">
C o m m e n t: The two 
<typeStatus id="549C880AFFC3FFE4FE18FC1D982CFC6D" box="[442,539,908,929]" pageId="5" pageNumber="974" type="paratype">paratypes</typeStatus>
preserved in concentrated ethanol were not used for the measurements in the description.
</paragraph>
<paragraph id="8B9836A8FFC3FFE4FF10FC5E98AEFB59" blockId="5.[178,1085,975,1173]" pageId="5" pageNumber="974">
D e s c r i p t i o n: Body length 4.7-5.0 mm; length of forebody 
<quantity id="4CDF9B4DFFC3FFE4FCFCFC5E9999FC28" box="[862,942,975,996]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="5" pageNumber="974" unit="mm" value="4.1">4.1 mm</quantity>
. Habitus as in 
<figureCitation id="131C2A2DFFC3FFE4FF10FC7D9AD8FBCD" box="[178,239,1004,1026]" captionStart="Figs 1-3" captionStartId="2.[178,218,1506,1528]" captionTargetBox="[192,1076,210,1483]" captionTargetId="figure@2.[179,1086,201,1486]" captionTargetPageId="2" captionText="Figs 1-3: Caucanthobium mirandum nov.sp.: (1) habitus; (2) antenna; (3) male protarsus. Scale bars: 1-2: 1.0 mm; 3: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3776458" httpUri="https://zenodo.org/record/3776458/files/figure.png" pageId="5" pageNumber="974">Fig. 1</figureCitation>
. Coloration: head blackish-brown with all, or the anterior portion of, the frons and the lateral portions of the posterior constriction reddish-yellow to red; pronotum uniformly dark-yellowish or dark-yellowish with the median portion slightly darker; elytra dark-yellowish, each with two large infuscate spots, one slightly anterior to middle and one in posterior portion; abdomen reddish-yellow; legs yellow; antennae reddish with antennomeres V-IX slightly to distinctly darker.
</paragraph>
<paragraph id="8B9836A8FFC3FFE4FF10FB359B49FAE3" blockId="5.[178,1085,1188,1327]" pageId="5" pageNumber="974">
Head (
<figureCitation id="131C2A2DFFC3FFE4FF54FB359B79FB76" box="[246,334,1188,1210]" captionStart="Figs 4-8" captionStartId="3.[178,218,1514,1536]" captionTargetBox="[182,1089,212,1497]" captionTargetId="figure@3.[182,1089,212,1497]" captionTargetPageId="3" captionText="Figs 4-8: Caucanthobium mirandum nov.sp.: (4) head and pronotum; (5) head in dorso-lateral view; (6) head in ventral view; (7) thorax in ventral view; (8) apical portion of abdomen. Scale bars: 7: 1.0 mm; 4-6, 8: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3776460" httpUri="https://zenodo.org/record/3776460/files/figure.png" pageId="5" pageNumber="974">Figs 4-6</figureCitation>
) transverse, median portion somewhat elevated and with irregularly distributed punctation, between this elevation and dorsal margins of eyes with a deep and densely punctate impression on either side; frons with transverse microsculpture and with very fine and rather sparse punctation. Antenna (
<figureCitation id="131C2A2DFFC3FFE4FD7EFB6D992DFADE" box="[732,794,1276,1298]" captionStart="Figs 1-3" captionStartId="2.[178,218,1506,1528]" captionTargetBox="[192,1076,210,1483]" captionTargetId="figure@2.[179,1086,201,1486]" captionTargetPageId="2" captionText="Figs 1-3: Caucanthobium mirandum nov.sp.: (1) habitus; (2) antenna; (3) male protarsus. Scale bars: 1-2: 1.0 mm; 3: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3776458" httpUri="https://zenodo.org/record/3776458/files/figure.png" pageId="5" pageNumber="974">Fig. 2</figureCitation>
) 1.9-2.0 mm long; antennomeres IV-X oblong.
</paragraph>
<paragraph id="8B9836A8FFC3FFE4FF10FAAE9876F9EE" blockId="5.[178,1085,1343,1570]" pageId="5" pageNumber="974">
Pronotum (
<figureCitation id="131C2A2DFFC3FFE4FE87FAAE9B52FA99" box="[293,357,1343,1365]" captionStart="Figs 4-8" captionStartId="3.[178,218,1514,1536]" captionTargetBox="[182,1089,212,1497]" captionTargetId="figure@3.[182,1089,212,1497]" captionTargetPageId="3" captionText="Figs 4-8: Caucanthobium mirandum nov.sp.: (4) head and pronotum; (5) head in dorso-lateral view; (6) head in ventral view; (7) thorax in ventral view; (8) apical portion of abdomen. Scale bars: 7: 1.0 mm; 4-6, 8: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3776460" httpUri="https://zenodo.org/record/3776460/files/figure.png" pageId="5" pageNumber="974">Fig. 4</figureCitation>
) approximately 1.4 times as broad as long and 1.55 times as broad as head; lateral margins broadly explanate and weakly serrulate, sinuate near the sharply marked posterior angles; antero-lateral angles rather weakly projecting anteriad; median portion in anterior three-fifths with an oblong median impression and an oblong elevation on either side of this impression, in posterior portion with a median impression and with an elevation on either side of this impression; punctation very coarse and rather dense, somewhat irregularly distributed in median portion; interstices with micropunctation, otherwise without microsculpture.
</paragraph>
<paragraph id="8B9836A8FFC3FFE7FF10F9A09991FE85" blockId="5.[178,1085,1585,1608]" lastBlockId="6.[178,1085,219,329]" lastPageId="6" lastPageNumber="975" pageId="5" pageNumber="974">
Elytra (
<figureCitation id="131C2A2DFFC3FFE4FF5DF9A09B0BF984" box="[255,316,1585,1608]" captionStart="Figs 1-3" captionStartId="2.[178,218,1506,1528]" captionTargetBox="[192,1076,210,1483]" captionTargetId="figure@2.[179,1086,201,1486]" captionTargetPageId="2" captionText="Figs 1-3: Caucanthobium mirandum nov.sp.: (1) habitus; (2) antenna; (3) male protarsus. Scale bars: 1-2: 1.0 mm; 3: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3776458" httpUri="https://zenodo.org/record/3776458/files/figure.png" pageId="5" pageNumber="974">Fig. 1</figureCitation>
) approximately 1.05 times as long as combined width, about 1.2 times as broad and 1.8 times as long as pronotum; lateral margins broadly explanate and serrulate; punctation coarse and dense, laterally and posteriorly seriate. Hind wings completely reduced. Legs long; metatibia 
<quantity id="4CDF9B4DFFC0FFE7FE55FE8498BDFEE7" box="[503,650,277,299]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="6" pageNumber="975" unit="mm" value="1.275" valueMax="1.3" valueMin="1.25">1.25-1.30 mm</quantity>
long. Abdomen (
<figureCitation id="131C2A2DFFC0FFE7FCEBFE8499BEFEE0" box="[841,905,277,300]" captionStart="Figs 4-8" captionStartId="3.[178,218,1514,1536]" captionTargetBox="[182,1089,212,1497]" captionTargetId="figure@3.[182,1089,212,1497]" captionTargetPageId="3" captionText="Figs 4-8: Caucanthobium mirandum nov.sp.: (4) head and pronotum; (5) head in dorso-lateral view; (6) head in ventral view; (7) thorax in ventral view; (8) apical portion of abdomen. Scale bars: 7: 1.0 mm; 4-6, 8: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3776460" httpUri="https://zenodo.org/record/3776460/files/figure.png" pageId="6" pageNumber="975">Fig. 8</figureCitation>
) with dense and distinct punctation; posterior margin of tergite VII without palisade fringe.
</paragraph>
<paragraph id="8B9836A8FFC0FFE7FF10FEC89E07FE2F" blockId="6.[178,1085,344,483]" pageId="6" pageNumber="975">
<specimenCount id="9D21FD21FFC0FFE7FF10FEC89AFFFEA2" box="[178,200,345,366]" pageId="6" pageNumber="975" type="male">♂</specimenCount>
: protarsomeres I-IV (
<figureCitation id="131C2A2DFFC0FFE7FE06FEC99BD6FEA2" box="[420,481,344,366]" captionStart="Figs 1-3" captionStartId="2.[178,218,1506,1528]" captionTargetBox="[192,1076,210,1483]" captionTargetId="figure@2.[179,1086,201,1486]" captionTargetPageId="2" captionText="Figs 1-3: Caucanthobium mirandum nov.sp.: (1) habitus; (2) antenna; (3) male protarsus. Scale bars: 1-2: 1.0 mm; 3: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3776458" httpUri="https://zenodo.org/record/3776458/files/figure.png" pageId="6" pageNumber="975">Fig. 3</figureCitation>
) distinctly dilated; posterior margin of tergite VIII broadly concave; aedeagus (
<figureCitation id="131C2A2DFFC0FFE7FE23FEE49BD2FE40" box="[385,485,373,396]" captionStart="Figs 9-11" captionStartId="4.[178,218,896,918]" captionTargetBox="[189,1085,204,856]" captionTargetId="figure@4.[179,1086,203,878]" captionTargetPageId="4" captionText="Figs 9-11: Caucanthobium mirandum nov.sp.: (9-10) aedeagus in lateral and in ventral view; (11) female genital segment. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3776462" httpUri="https://zenodo.org/record/3776462/files/figure.png" pageId="6" pageNumber="975">Figs 9-10</figureCitation>
) of highly derived morphology, small in relation to body size, 
<quantity id="4CDF9B4DFFC0FFE7FF45FE029B75FE64" box="[231,322,403,425]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="6" pageNumber="975" unit="mm" value="0.67">0.67 mm</quantity>
long (including parameres), weakly sclerotized, strongly asymmetric with the right paramere much longer and larger than the left paramere (ventral view), and with a conspicuous dark internal structure apically extending beyond apex of ventral process.
</paragraph>
<paragraph id="8B9836A8FFC0FFE7FF10FE629B0CFDAC" blockId="6.[178,1085,499,609]" pageId="6" pageNumber="975">
<specimenCount id="9D21FD21FFC0FFE7FF10FE629AF0FDC4" box="[178,199,499,520]" pageId="6" pageNumber="975" type="female">♀</specimenCount>
: protarsomeres I-IV weakly dilated, much less so than in male; posterior margin of tergite VIII truncate; posterior margin of sternite VIII broadly convex, in the middle weakly concave; gonocoxites (
<figureCitation id="131C2A2DFFC0FFE7FE51FDBC9809FD88" box="[499,574,557,580]" captionStart="Figs 9-11" captionStartId="4.[178,218,896,918]" captionTargetBox="[189,1085,204,856]" captionTargetId="figure@4.[179,1086,203,878]" captionTargetPageId="4" captionText="Figs 9-11: Caucanthobium mirandum nov.sp.: (9-10) aedeagus in lateral and in ventral view; (11) female genital segment. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3776462" httpUri="https://zenodo.org/record/3776462/files/figure.png" pageId="6" pageNumber="975">Fig. 11</figureCitation>
) short and broad, with short styli and very long terminal seta.
</paragraph>
<paragraph id="8B9836A8FFC0FFE7FF10FDE1995AFD0D" blockId="6.[178,1084,624,705]" pageId="6" pageNumber="975">
C o m p a r a t i v e n o t e s: 
<taxonomicName id="4C274D2BFFC0FFE7FDB8FDE19905FD49" authorityName="Assing" authorityYear="2018" box="[538,818,624,645]" class="Insecta" family="Staphylinidae" genus="Caucanthobium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="975" phylum="Arthropoda" rank="species" species="mirandum">
<emphasis id="B953EABAFFC0FFE7FDB8FDE19905FD49" box="[538,818,624,645]" italics="true" pageId="6" pageNumber="975">Caucanthobium mirandum</emphasis>
</taxonomicName>
is readily distinguished from other 
<taxonomicName id="4C274D2BFFC0FFE7FE87FD1C9B87FD6F" authorityName="C.G. Thomson" authorityYear="1859" box="[293,432,653,675]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="975" phylum="Arthropoda" rank="tribe" tribe="Anthophagini">Anthophagini</taxonomicName>
distributed in the Caucasus region by the generic characters, as well as by its coloration and the unique morphology of the aedeagus.
</paragraph>
<caption id="DF586620FFC0FFE7FF10FB5998B6FB11" ID-DOI="http://doi.org/10.5281/zenodo.3776464" ID-Zenodo-Dep="3776464" box="[178,641,1224,1246]" httpUri="https://zenodo.org/record/3776464/files/figure.png" pageId="6" pageNumber="975" startId="6.[178,224,1224,1245]" targetBox="[178,1086,722,1201]" targetPageId="6">
<paragraph id="8B9836A8FFC0FFE7FF10FB5998B6FB11" blockId="6.[178,641,1224,1246]" box="[178,641,1224,1246]" pageId="6" pageNumber="975">
<emphasis id="B953EABAFFC0FFE7FF10FB599AC7FB11" bold="true" box="[178,240,1224,1245]" pageId="6" pageNumber="975">Map 1</emphasis>
: Distribution of 
<taxonomicName id="4C274D2BFFC0FFE7FE2AFB59984CFB11" authorityName="Assing" authorityYear="2018" box="[392,635,1224,1245]" class="Insecta" family="Staphylinidae" genus="Caucanthobium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="975" phylum="Arthropoda" rank="species" species="mirandum">
<emphasis id="B953EABAFFC0FFE7FE2AFB59984CFB11" box="[392,635,1224,1245]" italics="true" pageId="6" pageNumber="975">Caucanthobium mirandum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B9836A8FFC0FFE7FF10FB6C994CFA13" blockId="6.[178,1085,1276,1503]" pageId="6" pageNumber="975">
D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: This species is currently known from three localities in 
<collectingRegion id="49E3F84AFFC0FFE7FE5AFA88980AFAE3" box="[504,573,1305,1327]" country="Georgia" name="Ajaria" pageId="6" pageNumber="975">Adjara</collectingRegion>
and 
<collectingRegion id="49E3F84AFFC0FFE7FDD7FA889898FAE3" box="[629,687,1305,1327]" country="Georgia" name="Guria" pageId="6" pageNumber="975">Guria</collectingRegion>
regions, Southwest 
<collectingCountry id="F3307638FFC0FFE7FC27FA8899EDFAE3" box="[901,986,1305,1327]" name="Georgia" pageId="6" pageNumber="975">Georgia</collectingCountry>
. It seems remarkable that a conspicuous species like 
<taxonomicName id="4C274D2BFFC0FFE7FDDEFAA69932FA80" authorityName="Assing" authorityYear="2018" box="[636,773,1335,1356]" class="Insecta" family="Staphylinidae" genus="Caucanthobium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="975" phylum="Arthropoda" rank="species" species="mirandum">
<emphasis id="B953EABAFFC0FFE7FDDEFAA69932FA80" box="[636,773,1335,1356]" italics="true" pageId="6" pageNumber="975">C. mirandum</emphasis>
</taxonomicName>
should have remained undiscovered up until today. Possible explanations could be a very restricted distribution and/or a special habitat. The 
<typeStatus id="549C880AFFC0FFE7FE47FAE09809FA4B" box="[485,574,1393,1415]" pageId="6" pageNumber="975" type="holotype">holotype</typeStatus>
was sifted from leaf litter in a mixed deciduous forest with beech, chestnut, hornbeam, alder, hazelnut, laurel, and rhododendron undergrowth (MEYBOHM pers. comm.). Two of the 
<typeStatus id="549C880AFFC0FFE7FD03FA3C9935FA0E" box="[673,770,1453,1474]" pageId="6" pageNumber="975" type="paratype">paratypes</typeStatus>
were sifted from litter and a mouse nest in a hollow tree. The altitudes range from 
<quantity id="4CDF9B4DFFC0FFE7FD77FA5B9940FA13" box="[725,887,1481,1503]" metricMagnitude="3" metricUnit="m" metricValue="1.6149999999999998" metricValueMax="1.95" metricValueMin="1.28" pageId="6" pageNumber="975" unit="m" value="1615.0" valueMax="1950.0" valueMin="1280.0">
<date id="FF991068FFC0FFE7FD77FA5B996DFA13" box="[725,858,1481,1503]" pageId="6" pageNumber="975" value="1280" valueMax="1950">1280 to 1950</date>
m
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>