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<document id="EFE8DA259E567DBD7FA179F1C48BE734" ID-DOI="10.12651/JSR.2014.3.2.127" ID-ISSN="2713-8615" ID-Zenodo-Dep="13143732" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722430635402" checkinUser="felipe" docAuthor="Blakemore, Robert J., Lee, Seunghan &amp; Seo, Hong-Yul" docDate="2014" docId="0390636CFFACFFF9FF3DFB8EFBA28A62" docLanguage="en" docName="JSpeciesRes.3.2.127-166.pdf" docOrigin="Journal of Species Research 3 (2)" docSource="http://dx.doi.org/10.12651/jsr.2014.3.2.127" docStyle="DocumentStyle:4D398BFF45FF8CA9E08F546BB06AE372.2:JSpeciesRes.2012-.journal_article" docStyleId="4D398BFF45FF8CA9E08F546BB06AE372" docStyleName="JSpeciesRes.2012-.journal_article" docStyleVersion="2" docTitle="Drawida nemora Kobayashi 1936" docType="treatment" docVersion="2" lastPageNumber="140" masterDocId="FFA91B14FFA0FFF4FFA0FFE5FFA88C73" masterDocTitle="Reports of Drawida (Oligochaeta: Moniligastridae) from far East Asia" masterLastPageNumber="166" masterPageNumber="127" pageNumber="139" updateTime="1722431195132" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3276728C6D346D6E5DF6813423D868D2">Reports of Drawida (Oligochaeta: Moniligastridae) from far East Asia</mods:title>
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<mods:namePart id="4BF2C27FD2D7D65C7E925B82E9E3BE45">Blakemore, Robert J.</mods:namePart>
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<mods:namePart id="728421D34C5D97C5A7EA07545D6C28FB">Lee, Seunghan</mods:namePart>
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<mods:namePart id="5803BA5B58A74812567CC870534F6C7D">Seo, Hong-Yul</mods:namePart>
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<treatment id="0390636CFFACFFF9FF3DFB8EFBA28A62" LSID="urn:lsid:plazi:treatment:0390636CFFACFFF9FF3DFB8EFBA28A62" httpUri="http://treatment.plazi.org/id/0390636CFFACFFF9FF3DFB8EFBA28A62" lastPageId="13" lastPageNumber="140" pageId="12" pageNumber="139">
<subSubSection id="C32381F1FFACFFF8FF3DFB8EFD9C88F0" box="[157,564,1131,1155]" pageId="12" pageNumber="139" type="nomenclature">
<paragraph id="8B86D27AFFACFFF8FF3DFB8EFD9C88F0" blockId="12.[157,564,1131,1155]" box="[157,564,1131,1155]" pageId="12" pageNumber="139">
<heading id="D0CE6516FFACFFF8FF3DFB8EFD9C88F0" bold="true" box="[157,564,1131,1155]" fontSize="36" level="2" pageId="12" pageNumber="139" reason="3">
<emphasis id="B94D0E68FFACFFF8FF3DFB8EFD9C88F0" bold="true" box="[157,564,1131,1155]" pageId="12" pageNumber="139">
8.
<taxonomicName id="4C39A9F9FFACFFF8FF18FB8EFD9C88F0" authority="Kobayashi, 1936" authorityName="Kobayashi" authorityYear="1936" box="[184,564,1131,1155]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="12" pageNumber="139" phylum="Annelida" rank="species" species="nemora">
<emphasis id="B94D0E68FFACFFF8FF18FB8EFED988F0" bold="true" box="[184,369,1131,1155]" italics="true" pageId="12" pageNumber="139">Drawida nemora</emphasis>
<bibRefCitation id="EFA8AF8BFFACFFF8FED8FB8EFD9C88F0" author="Kobayashi, S." box="[376,564,1131,1155]" pageId="12" pageNumber="139" pagination="139 - 184" refId="ref22870" refString="Kobayashi, S. 1936. Earthworms from Koryo, Korea. Science Report of the Tohoku Imperial University 11 (1): 139 - 184." type="journal article" year="1936">Kobayashi, 1936</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32381F1FFACFFF8FF3DFB69FD4689B7" pageId="12" pageNumber="139" type="reference_group">
<paragraph id="8B86D27AFFACFFF8FF3DFB69FD618910" blockId="12.[157,762,1163,1476]" pageId="12" pageNumber="139">
<treatmentCitationGroup id="AB29F554FFACFFF8FF3DFB69FD618910" pageId="12" pageNumber="139">
<treatmentCitation id="0A98F46BFFACFFF8FF3DFB69FDF888D0" author="Kobayashi, S." box="[157,592,1164,1187]" page="141" pageId="12" pageNumber="139" year="1936">
<taxonomicName id="4C39A9F9FFACFFF8FF3DFB69FDF888D0" authority="Kobayashi, 1936: 141" authorityName="Kobayashi" authorityPageNumber="141" authorityYear="1936" box="[157,592,1164,1187]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="12" pageNumber="139" phylum="Annelida" rank="species" species="nemora">
<emphasis id="B94D0E68FFACFFF8FF3DFB69FEFE88D0" box="[157,342,1164,1187]" italics="true" pageId="12" pageNumber="139">Drawida nemora</emphasis>
<materialsCitation id="3B51D827FFACFFF8FEFFFB69FDF888D0" box="[351,592,1164,1187]" collectorName="From Koryo &amp; A.N. Leirikh &amp; Keiki-do &amp; Keijo" country="South Korea" county="July" elevation="300" location="Mr Kiujiro Susaki" municipality="April" pageId="12" pageNumber="139" specimenCount="1" stateProvince="Seoul">
<bibRefCitation id="EFA8AF8BFFACFFF8FEFFFB69FDF888D0" author="Kobayashi, S." box="[351,592,1164,1187]" pageId="12" pageNumber="139" pagination="139 - 184" refId="ref22870" refString="Kobayashi, S. 1936. Earthworms from Koryo, Korea. Science Report of the Tohoku Imperial University 11 (1): 139 - 184." type="journal article" year="1936">Kobayashi, 1936: 141</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
, text-fig. 1.1-8. [
<collectorName id="26CCB7ACFFACFFF8FF1AFB49FEEE88B0" box="[186,326,1196,1219]" pageId="12" pageNumber="139">From Koryo</collectorName>
“(
<emphasis id="B94D0E68FFACFFF8FEC4FB49FE4888B0" box="[356,480,1196,1219]" italics="true" pageId="12" pageNumber="139">
<collectorName id="26CCB7ACFFACFFF8FEC4FB49FE3B88B0" box="[356,403,1196,1219]" name="A.N. Leirikh" pageId="12" pageNumber="139">N.L.</collectorName>
37 45
</emphasis>
)” {sic},
<collectorName id="26CCB7ACFFACFFF8FDEEFB4EFD1888B0" box="[590,688,1195,1219]" pageId="12" pageNumber="139">Keiki-do</collectorName>
, about
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distant from
<collectorName id="26CCB7ACFFACFFF8FE33FB29FE678890" box="[403,463,1228,1251]" pageId="12" pageNumber="139">Keijo</collectorName>
(=
<collectingRegion id="49FD1C98FFACFFF8FE54FB29FD9D8890" box="[500,565,1228,1251]" country="South Korea" name="Seoul" pageId="12" pageNumber="139">Seoul</collectingRegion>
),
<collectingCountry id="F32E92EAFFACFFF8FDECFB29FD268890" box="[588,654,1228,1251]" name="South Korea" pageId="12" pageNumber="139">Korea</collectingCountry>
collected from woodland about
<quantity id="4CC17F9FFFACFFF8FE0AFB09FE468970" box="[426,494,1260,1283]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="139" unit="m" value="300.0">
<elevation id="00143549FFACFFF8FE0AFB09FE468970" box="[426,494,1260,1283]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="139" unit="m" value="300.0">300 m</elevation>
</quantity>
altitude by “
<emphasis id="B94D0E68FFACFFF8FD22FB09FDF08930" italics="true" pageId="12" pageNumber="139">
<location id="8EE684A1FFACFFF8FD22FB09FF548950" LSID="urn:lsid:plazi:treatment:0390636CFFACFFF9FF3DFB8EFBA28A62:8EE684A1FFACFFF8FD22FB09FF548950" country="South Korea" county="July" municipality="April" name="Mr Kiujiro Susaki" pageId="12" pageNumber="139" stateProvince="Seoul">Mr Kiujiro Susaki</location>
, an assistant expert of the
<location id="8EE684A1FFACFFF8FDBDFAE9FEA88930" LSID="urn:lsid:plazi:treatment:0390636CFFACFFF9FF3DFB8EFBA28A62:8EE684A1FFACFFF8FDBDFAE9FEA88930" country="South Korea" county="July" municipality="April" name="Forest Experimental Station" pageId="12" pageNumber="139" stateProvince="Seoul">Forest Experimental Station</location>
,
<location id="8EE684A1FFACFFF8FEABFAC9FDF08930" LSID="urn:lsid:plazi:treatment:0390636CFFACFFF9FF3DFB8EFBA28A62:8EE684A1FFACFFF8FEABFAC9FDF08930" box="[267,600,1324,1347]" country="South Korea" county="July" municipality="April" name="General Governemnt of Chosen" pageId="12" pageNumber="139" stateProvince="Seoul">General Governemnt of Chosen</location>
</emphasis>
” {sic} in
<collectingMunicipality id="6BE24800FFACFFF8FD63FAC9FD528930" box="[707,762,1324,1347]" pageId="12" pageNumber="139">April</collectingMunicipality>
and
<collectingCounty id="62E7AAF6FFACFFF8FF41FAA9FEB88910" box="[225,272,1356,1379]" pageId="12" pageNumber="139">July</collectingCounty>
, 1934.
<typeStatus id="54826CD8FFACFFF8FEFDFAA9FE368910" box="[349,414,1356,1379]" pageId="12" pageNumber="139">Types</typeStatus>
not known, presumed lost].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B86D27AFFACFFF8FF3DFA88FD4689B7" blockId="12.[157,762,1163,1476]" pageId="12" pageNumber="139">
<treatmentCitationGroup id="AB29F554FFACFFF8FF3DFA88FD4689B7" pageId="12" pageNumber="139">
<taxonomicName id="4C39A9F9FFACFFF8FF3DFA88FEFD89F7" authorityName="Kobayashi" authorityYear="1936" box="[157,341,1389,1412]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="12" pageNumber="139" phylum="Annelida" rank="species" species="nemora">
<emphasis id="B94D0E68FFACFFF8FF3DFA88FEFD89F7" box="[157,341,1389,1412]" italics="true" pageId="12" pageNumber="139">Drawida nemora</emphasis>
</taxonomicName>
:
<treatmentCitation id="0A98F46BFFACFFF8FEC5FA89FDD189F0" author="Kobayashi, S." box="[357,633,1388,1411]" page="99" pageId="12" pageNumber="139" year="1938">
<bibRefCitation id="EFA8AF8BFFACFFF8FEC5FA89FDD189F0" author="Kobayashi, S." box="[357,633,1388,1411]" pageId="12" pageNumber="139" pagination="89 - 170" refId="ref22932" refString="Kobayashi, S. 1938. Earthworms of Korea I. Science Report of the Tohoku Imperial University 13 (2): 89 - 170." type="journal article" year="1938">Kobayashi, 1938: 99-102</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A98F46BFFACFFF8FD26FA89FD5D89F0" author="Kobayashi, S." box="[646,757,1388,1411]" page="272" pageId="12" pageNumber="139" year="1940">
<bibRefCitation id="EFA8AF8BFFACFFF8FD26FA89FD5D89F0" author="Kobayashi, S." box="[646,757,1388,1411]" pageId="12" pageNumber="139" refId="ref22959" refString="Kobayashi, S. 1940. Terrestrial Oligochaeta from Manchoukuo. The Science reports of the Tohoku Imperial University, IV Series 15: 261 - 256." type="book" year="1940">1940: 272</bibRefCitation>
</treatmentCitation>
; 1941b: 263;
<treatmentCitation id="0A98F46BFFACFFF8FE98FA68FE4389D7" author="Easton, E. G." box="[312,491,1421,1444]" page="39" pageId="12" pageNumber="139" year="1981">
<bibRefCitation id="EFA8AF8BFFACFFF8FE98FA68FE4389D7" author="Easton, E. G." box="[312,491,1421,1444]" pageId="12" pageNumber="139" pagination="33 - 65" refId="ref21957" refString="Easton, E. G. 1981. Japanese earthworms: a synopsis of the Megadrile species. ‾ Bulletin of the British Museum (Natural History) Zoology 40 (2): 33 - 65 [Available from: http: // www. archive. org / stream / bulletinofbritis 40 zoollond # pa ge / n 43 / mode / 2 up]." type="journal article" year="1981">Easton, 1981: 39</bibRefCitation>
</treatmentCitation>
, tab. 1;
<bibRefCitation id="EFA8AF8BFFACFFF8FD9FFA68FD5E89D7" author="Blakemore, R. J." box="[575,758,1421,1444]" pageId="12" pageNumber="139" pagination="241 - 244" refId="ref20589" refString="Blakemore, R. J. 2003. Japanese earthworms (Annelida: Oligochaeta): a review and checklist of species. Organisms, Diversity and Evolution 3 (3): 241 - 244. [Available from: http: // www. urbanfischer. de / journals / ode /; www. sencken berg. de / odes / 03 - 11. htm]." type="journal article" year="2003">Blakemore, 2003</bibRefCitation>
, 2010, 2014; Blakemore &amp; Kupriyanova, 2010: tab. 3.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32381F1FFACFFF8FF3DFA08FF598BF5" pageId="12" pageNumber="139" type="distribution">
<paragraph id="8B86D27AFFACFFF8FF3DFA08FF598BF5" blockId="12.[157,762,1517,1990]" pageId="12" pageNumber="139">
<emphasis id="B94D0E68FFACFFF8FF3DFA08FE9C8A76" bold="true" box="[157,308,1517,1541]" pageId="12" pageNumber="139">Distribution.</emphasis>
Central and
<collectingCountry id="F32E92EAFFACFFF8FE6BFA08FDF38A77" box="[459,603,1517,1540]" name="North Korea" pageId="12" pageNumber="139">North Korea</collectingCountry>
, Manchuria,
<collectingCountry id="F32E92EAFFACFFF8FF3DF9E8FF758A57" box="[157,221,1549,1572]" name="China" pageId="12" pageNumber="139">China</collectingCountry>
and Honshu,
<collectingCountry id="F32E92EAFFACFFF8FEDAF9E8FE128A57" box="[378,442,1549,1572]" name="Japan" pageId="12" pageNumber="139">Japan</collectingCountry>
. E.g. from Koryo near Keijo (present day
<collectingRegion id="49FD1C98FFACFFF8FE84F9C8FECB8A37" box="[292,355,1581,1604]" country="South Korea" name="Seoul" pageId="12" pageNumber="139">Seoul</collectingRegion>
),
<collectingCountry id="F32E92EAFFACFFF8FED5F9C8FE1D8A37" box="[373,437,1581,1604]" name="South Korea" pageId="12" pageNumber="139">Korea</collectingCountry>
(
<bibRefCitation id="EFA8AF8BFFACFFF8FE61F9C8FDDF8A37" author="Kobayashi, S." box="[449,631,1581,1604]" pageId="12" pageNumber="139" pagination="139 - 184" refId="ref22870" refString="Kobayashi, S. 1936. Earthworms from Koryo, Korea. Science Report of the Tohoku Imperial University 11 (1): 139 - 184." type="journal article" year="1936">Kobayashi, 1936</bibRefCitation>
); with many other Korean records by
<bibRefCitation id="EFA8AF8BFFACFFF8FE17F9A8FDD18A17" author="Kobayashi, S." box="[439,633,1613,1636]" pageId="12" pageNumber="139" pagination="89 - 170" refId="ref22932" refString="Kobayashi, S. 1938. Earthworms of Korea I. Science Report of the Tohoku Imperial University 13 (2): 89 - 170." type="journal article" year="1938">Kobayashi (1938)</bibRefCitation>
some from Mt Kambo (at
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<geoCoordinate id="EE0DB4BDFFACFFF8FEE0F98BFED48AF6" box="[320,380,1646,1669]" degrees="2,000" direction="north" orientation="latitude" pageId="12" pageNumber="139" precision="55" value="2.0">2,000</geoCoordinate>
<geoCoordinate id="EE0DB4BDFFACFFF8FEDCF988FE698AF6" box="[380,449,1645,1669]" degrees="2,500" direction="west" orientation="longitude" pageId="12" pageNumber="139" precision="55" value="-2.5">-2,500</geoCoordinate>
m
</quantity>
) and Akashima in the far northeast of the peninsula “
<emphasis id="B94D0E68FFACFFF8FE66F96BFEC38AB6" italics="true" pageId="12" pageNumber="139">found within wet roots of alpine plants such as</emphasis>
Pedicularis manshurica
<emphasis id="B94D0E68FFACFFF8FDD3F94BFF7F8B76" italics="true" pageId="12" pageNumber="139">Maximowicz in the former and under decayed leaved on granite on the latter</emphasis>
”; from several central “
<emphasis id="B94D0E68FFACFFF8FE7AF90BFDCD8B76" box="[474,613,1774,1797]" italics="true" pageId="12" pageNumber="139">Manchoukuo</emphasis>
” locations by
<bibRefCitation id="EFA8AF8BFFACFFF8FF3DF8EBFEF58B56" author="Kobayashi, S." box="[157,349,1806,1829]" pageId="12" pageNumber="139" refId="ref22959" refString="Kobayashi, S. 1940. Terrestrial Oligochaeta from Manchoukuo. The Science reports of the Tohoku Imperial University, IV Series 15: 261 - 256." type="book" year="1940">Kobayashi (1940)</bibRefCitation>
and a Chubu-Chiho,
<collectingCountry id="F32E92EAFFACFFF8FDECF8EBFD218B56" box="[588,649,1806,1829]" name="Japan" pageId="12" pageNumber="139">Japan</collectingCountry>
record by Kobayashi (1941: 236), possibly introduced/transported there or, more likely given its wide definition, a different species.
</paragraph>
</subSubSection>
<subSubSection id="C32381F1FFACFFF9FF3DF86BFF4D8AC2" lastPageId="13" lastPageNumber="140" pageId="12" pageNumber="139" type="description">
<paragraph id="8B86D27AFFACFFF8FF3DF86BFC6D8A37" blockId="12.[157,762,1517,1990]" lastBlockId="12.[822,1428,1131,1990]" pageId="12" pageNumber="139">
<emphasis id="B94D0E68FFACFFF8FF3DF86BFE808BD5" bold="true" box="[157,296,1934,1958]" pageId="12" pageNumber="139">Description.</emphasis>
Dorsally a dark blue often with a purplish or yellowish tinge, less distinct ventrally and clitellar segments yellowish grey in life. Size
<quantity id="4CC17F9FFFACFFF8FB12FB8EFA8488F1" box="[1202,1324,1131,1154]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.85" metricValueMin="0.65" pageId="12" pageNumber="139" unit="mm" value="125.0" valueMax="185.0" valueMin="65.0">65-185 mm</quantity>
by up to
<quantity id="4CC17F9FFFACFFF8FC96FB69FC2188D0" box="[822,905,1164,1187]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="12" pageNumber="139" unit="mm" value="6.5">6.5 mm</quantity>
with ca. 162-247 segments Prolobous. Dorsal pores absent. Setae small and closely paired (ab=cd). Nephropores small in d-lines. Clitellum mostly saddle-shaped in 9,10-13,14. Spermathecal pores small in 7/8 (or just anterior to intersegment in 7) in c-lines as transverse slits. Male pore secondarily superficial in 10/11 approximately in mid-bc in transverse slits; primary pores shown by
<bibRefCitation id="EFA8AF8BFFACFFF8FC05FA89FBF589F0" author="Kobayashi, S." box="[933,1117,1388,1411]" pageId="12" pageNumber="139" pagination="139 - 184" refId="ref22870" refString="Kobayashi, S. 1936. Earthworms from Koryo, Korea. Science Report of the Tohoku Imperial University 11 (1): 139 - 184." type="journal article" year="1936">Kobayashi (1936</bibRefCitation>
: fig. 1.4) to be on small conical penes. Female pores paired on anterior of 12 near 11/12 approximately in b lines. Genital papillae variously in 6-13 ventrally detectable either externally or internally, or absent; shown by
<bibRefCitation id="EFA8AF8BFFACFFF8FBB2FA08FB618A77" author="Kobayashi, S." box="[1042,1225,1517,1540]" pageId="12" pageNumber="139" pagination="139 - 184" refId="ref22870" refString="Kobayashi, S. 1936. Earthworms from Koryo, Korea. Science Report of the Tohoku Imperial University 11 (1): 139 - 184." type="journal article" year="1936">Kobayashi (1936</bibRefCitation>
: fig. 1.1) as medial to c-lines in 8 and 9lhs, 11rhs and in regular medial to asetae in 7-12.
</paragraph>
<paragraph id="8B86D27AFFACFFF9FCEDF9A8FF4D8AC2" blockId="12.[822,1428,1131,1990]" lastBlockId="13.[159,764,1305,1971]" lastPageId="13" lastPageNumber="140" pageId="12" pageNumber="139">
Septa 5/6-8/9 thickened; septa 9/10 and 10/11 fused dorsally with 11/12 to enclose ovaries and funnels. Pharyngeal glands in 3-4. Hearts 6-9; dorsal blood vessel single. Nephridia holoic and vesiculate (absent from first and last segments and from segment 12) with bladders elongate. Spermatheca with ampulla ventral on septum 7/8, spherical to convoluted duct that exits to body wall without atrium nor accessory gland in 7/8 (
<bibRefCitation id="EFA8AF8BFFACFFF8FB75F8CBFA278B36" author="Kobayashi, S." box="[1237,1423,1838,1861]" pageId="12" pageNumber="139" pagination="139 - 184" refId="ref22870" refString="Kobayashi, S. 1936. Earthworms from Koryo, Korea. Science Report of the Tohoku Imperial University 11 (1): 139 - 184." type="journal article" year="1936">Kobayashi, 1936</bibRefCitation>
: figs. 1.7-8), or (
<bibRefCitation id="EFA8AF8BFFACFFF8FC7DF8ABFB648B16" author="Kobayashi, S." box="[989,1228,1870,1893]" pageId="12" pageNumber="139" pagination="89 - 170" refId="ref22932" refString="Kobayashi, S. 1938. Earthworms of Korea I. Science Report of the Tohoku Imperial University 13 (2): 89 - 170." type="journal article" year="1938">Kobayashi, 1938: 100</bibRefCitation>
) sometimes papillae just laterally to the spermathecal pores and “
<emphasis id="B94D0E68FFACFFF8FA8CF88AFCD58BD5" italics="true" pageId="12" pageNumber="139">accessory glands</emphasis>
” internally (same species?). Testis sacs spherical, intrasegmentally bulging to both sides of 9/10 with vas deferens slightly coiled and entering body wall under prostate. Prostate described definitively as “
<emphasis id="B94D0E68FFADFFF9FDD5FADCFE528903" italics="true" pageId="13" pageNumber="140">thick oval or circular disc, sessile on parietes</emphasis>
” mostly in 10 that exits to 10/11 via blunt, conical penis (
<bibRefCitation id="EFA8AF8BFFADFFF9FDAEFA9CFD6389E3" author="Kobayashi, S." box="[526,715,1401,1424]" pageId="13" pageNumber="140" pagination="139 - 184" refId="ref22870" refString="Kobayashi, S. 1936. Earthworms from Koryo, Korea. Science Report of the Tohoku Imperial University 11 (1): 139 - 184." type="journal article" year="1936">Kobayashi, 1936</bibRefCitation>
: fig. 1.6). Ovarian chamber in 11 with long ovisacs from fused 10/11/12 as far back as 14. Gizzards usually four in 12- 15, or sometimes five in 12-16 or three in 13-15 (i.e. three to five of in 12,13-15,16 suggesting either high plasticity of too broad definition). Corresponding to the external markings are small, white spherical bodies that are buried in body wall and appear to duct to exterior
<emphasis id="B94D0E68FFADFFF9FD10F9BDFD698A02" bold="true" box="[688,705,1624,1649]" pageId="13" pageNumber="140"></emphasis>
however, some may be parasitic artefacts of gregarine sporozoites.
</paragraph>
</subSubSection>
<caption id="DF4682F2FFADFFF9FF3FFB94FEC588A5" ID-DOI="http://doi.org/10.5281/zenodo.13143740" ID-Zenodo-Dep="13143740" httpUri="https://zenodo.org/record/13143740/files/figure.png" pageId="13" pageNumber="140" startId="13.[159,192,1137,1157]" targetBox="[229,1351,263,1103]" targetPageId="13" targetType="figure">
<paragraph id="8B86D27AFFADFFF9FF3FFB94FEC588A5" blockId="13.[159,1430,1137,1238]" pageId="13" pageNumber="140">
<emphasis id="B94D0E68FFADFFF9FF3FFB94FF7388F6" bold="true" box="[159,219,1137,1157]" pageId="13" pageNumber="140">Fig. 6.</emphasis>
<taxonomicName id="4C39A9F9FFADFFF9FF41FB97FDB688F5" authority="Kobayashi, 1936" authorityName="Kobayashi" authorityYear="1936" box="[225,542,1138,1158]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="13" pageNumber="140" phylum="Annelida" rank="species" species="nemora">
<emphasis id="B94D0E68FFADFFF9FF41FB97FED488F5" box="[225,380,1138,1158]" italics="true" pageId="13" pageNumber="140">Drawida nemora</emphasis>
<bibRefCitation id="EFA8AF8BFFADFFF9FE22FB97FDB688F5" author="Kobayashi, S." box="[386,542,1138,1158]" pageId="13" pageNumber="140" pagination="139 - 184" refId="ref22870" refString="Kobayashi, S. 1936. Earthworms from Koryo, Korea. Science Report of the Tohoku Imperial University 11 (1): 139 - 184." type="journal article" year="1936">Kobayashi, 1936</bibRefCitation>
</taxonomicName>
. After
<bibRefCitation id="EFA8AF8BFFADFFF9FDFCFB97FD5188F5" author="Kobayashi, S." box="[604,761,1138,1158]" pageId="13" pageNumber="140" pagination="139 - 184" refId="ref22870" refString="Kobayashi, S. 1936. Earthworms from Koryo, Korea. Science Report of the Tohoku Imperial University 11 (1): 139 - 184." type="journal article" year="1936">Kobayashi (1936</bibRefCitation>
: fig. 1). 1. Ventral view. 2. Male field with GMs. 3. Spermathecal pore in 7/8. 4. Internal organs. 5. Ventral view of prostate and penis with vas deferens. 6. Section of prostate and penis. 7. Spermatheca (Codes: bw. Body wall, giz. Gizzard, gp. Genital papilla, o. Ovary, och. Ovarian chamber, ovs. Ovisac, p. Penis, pr. Prostate, sp. Spermatheca, ts. Testis sac, vd. Vas deferens).
</paragraph>
</caption>
<subSubSection id="C32381F1FFADFFF9FF3FF95FFBA28A62" pageId="13" pageNumber="140" type="discussion">
<paragraph id="8B86D27AFFADFFF9FF3FF95FFBA28A62" blockId="13.[159,764,1305,1971]" lastBlockId="13.[824,1429,1305,1553]" pageId="13" pageNumber="140">
<emphasis id="B94D0E68FFADFFF9FF3FF95FFEA68AA1" bold="true" box="[159,270,1722,1746]" pageId="13" pageNumber="140">Remarks.</emphasis>
Despite
<bibRefCitation id="EFA8AF8BFFADFFF9FECFF95FFDCD8AA2" author="Kobayashi, S." box="[367,613,1722,1745]" pageId="13" pageNumber="140" pagination="139 - 184" refId="ref22870" refString="Kobayashi, S. 1936. Earthworms from Koryo, Korea. Science Report of the Tohoku Imperial University 11 (1): 139 - 184." type="journal article" year="1936">Kobayashi (1936: 146)</bibRefCitation>
saying it was the most predominant in the region in number,
<taxonomicName id="4C39A9F9FFADFFF9FD2CF93EFD548A81" authorityName="Kobayashi" authorityYear="1936" box="[652,764,1755,1778]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="13" pageNumber="140" phylum="Annelida" rank="species" species="nemora">
<emphasis id="B94D0E68FFADFFF9FD2CF93EFD548A81" box="[652,764,1755,1778]" italics="true" pageId="13" pageNumber="140">D. nemora</emphasis>
</taxonomicName>
has not since been definitely collected around
<collectingRegion id="49FD1C98FFADFFF9FD36F91EFD7B8B61" box="[662,723,1787,1810]" country="South Korea" name="Seoul" pageId="13" pageNumber="140">Seoul</collectingRegion>
(cf.
<taxonomicName id="4C39A9F9FFADFFF9FF3FF8FEFEA18B41" authorityName="Blakemore &amp; Lee &amp; Seo" authorityYear="2014" box="[159,265,1819,1842]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="13" pageNumber="140" phylum="Annelida" rank="species" species="csuzdii" status="sp. nov.">
<emphasis id="B94D0E68FFADFFF9FF3FF8FEFEA18B41" box="[159,265,1819,1842]" italics="true" pageId="13" pageNumber="140">D. csuzdii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EB313FFADFFF9FEB0F8FEFEC08B41" box="[272,360,1819,1842]" pageId="13" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
). Location and selection of a typotypic
<typeStatus id="54826CD8FFADFFF9FF19F8DEFEA58B21" box="[185,269,1851,1874]" pageId="13" pageNumber="140" type="neotype">neotype</typeStatus>
is required, especially since
<taxonomicName id="4C39A9F9FFADFFF9FD9FF8DEFE608B01" authority="Zhong &amp; Sun, 2014" authorityName="Zhong &amp; Sun" authorityYear="2014" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="13" pageNumber="140" phylum="Annelida" rank="species" species="dandongensis">
<emphasis id="B94D0E68FFADFFF9FD9FF8DEFF4A8B01" italics="true" pageId="13" pageNumber="140">Drawida dandongensis</emphasis>
Zhong &amp; Sun, 2014
</taxonomicName>
from the Sino-North
<collectingCountry id="F32E92EAFFADFFF9FD1BF8BEFD548B01" box="[699,764,1883,1906]" name="South Korea" pageId="13" pageNumber="140">Korea</collectingCountry>
border is described as being similar. However, there are inconsistencies in their description of
<taxonomicName id="4C39A9F9FFADFFF9FD9FF879FD5D8BC0" authorityName="Zhong &amp; Sun" authorityYear="2014" box="[575,757,1948,1971]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="13" pageNumber="140" phylum="Annelida" rank="species" species="dandongensis">
<emphasis id="B94D0E68FFADFFF9FD9FF879FD5D8BC0" box="[575,757,1948,1971]" italics="true" pageId="13" pageNumber="140">D. dandongensis</emphasis>
</taxonomicName>
, for example they have “
<emphasis id="B94D0E68FFADFFF9FB94FAFCFA218943" box="[1076,1417,1305,1328]" italics="true" pageId="13" pageNumber="140">
Gizzards
<quantity id="4CC17F9FFFADFFF9FB38FAFCFB688943" box="[1176,1216,1305,1328]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="13" pageNumber="140" unit="in" value="4.0">4 in</quantity>
XII-XVII segments
</emphasis>
” meaning four gizzards in 12-17 (=six segments, so six gizzards!?) so it cannot be meaningfully compared to any other
<taxonomicName id="4C39A9F9FFADFFF9FC04FA9CFBAB89E3" authorityName="F. Siemsseni Michaelsen" authorityYear="1910" box="[932,1027,1401,1424]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="13" pageNumber="140" phylum="Annelida" rank="genus">
<emphasis id="B94D0E68FFADFFF9FC04FA9CFBAB89E3" box="[932,1027,1401,1424]" italics="true" pageId="13" pageNumber="140">Drawida</emphasis>
</taxonomicName>
and thus is here given
<emphasis id="B94D0E68FFADFFF9FB5AFA9CFA3D89E3" box="[1274,1429,1401,1424]" italics="true" pageId="13" pageNumber="140">sp. inquirenda</emphasis>
status. [These latter authors also mistake the location of the spermathecal pore in
<taxonomicName id="4C39A9F9FFADFFF9FBE0FA5FFB1889A2" authorityName="Kobayashi" authorityYear="1936" box="[1088,1200,1466,1489]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="13" pageNumber="140" phylum="Annelida" rank="species" species="nemora">
<emphasis id="B94D0E68FFADFFF9FBE0FA5FFB1889A2" box="[1088,1200,1466,1489]" italics="true" pageId="13" pageNumber="140">D. nemora</emphasis>
</taxonomicName>
saying it is at “
<emphasis id="B94D0E68FFADFFF9FAF5FA5FFC468982" italics="true" pageId="13" pageNumber="140">posterior edge of VIII</emphasis>
”, while it is clearly shown at extreme edge of segment 7].
</paragraph>
</subSubSection>
</treatment>
</document>