treatments-xml/data/DA/0E/44/DA0E444FA33FBE099C1C8BFEB71240A6.xml

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<mods:title id="CDE28A13A2FB857077C7B4323896C238">A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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<mods:namePart id="40C69395BEC4D5688E43A4B4CC296C76">Caterino, Michael S.</mods:namePart>
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<treatment id="DA0E444FA33FBE099C1C8BFEB71240A6" ID-GBIF-Taxon="152040869" LSID="urn:lsid:plazi:treatment:DA0E444FA33FBE099C1C8BFEB71240A6" httpUri="http://treatment.plazi.org/id/DA0E444FA33FBE099C1C8BFEB71240A6" lastPageId="69" lastPageNumber="70" pageId="65" pageNumber="66" scope_family="Histeridae" scope_order="Coleoptera" scope_tribe="Exosternini">
<subSubSection id="C206C899E8E856F7A4437DD0FFEEF8F6" pageId="65" pageNumber="66" type="nomenclature">
<paragraph id="0D355C793699C7E8913CED01EC0971D7" pageId="65" pageNumber="66">
<taxonomicName id="E1584AE7E6710A3BA8208349D02BB3F9" ID-CoL="74NLW" LSID="http://species-id.net/wiki/Operclipygus_panamensis" authority="Wenzel &amp; Dybas, 1941" authorityName="Wenzel &amp; Dybas" authorityYear="1941" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis (Wenzel &amp; Dybas, 1941)</taxonomicName>
Figs 17 
<normalizedToken id="526CFD1A66C6E181E1C84FF716AFE6AA" originalValue="A–B18A–D">A-B18A-D</normalizedToken>
, FMap 6
</paragraph>
</subSubSection>
<subSubSection id="5B53D39BC9E73DB268006B68F8EFEAA0" pageId="65" pageNumber="66" type="reference_group">
<paragraph id="DF69FD227D3FE361AD596249F8075725" pageId="65" pageNumber="66">
<taxonomicName id="751D17403159D354149CEA509FC700BE" genus="Phelisteroides" lsidName="Phelisteroides panamensis" pageId="65" pageNumber="66" rank="species" species="panamensis">Phelisteroides panamensis</taxonomicName>
Wenzel &amp; Dybas, 1941: 450; 
<taxonomicName id="C2A3638197B61AE20C34970798C4D6BB" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
: 
<bibRefCitation id="E3D8B577D2FEE3C88D4B3610C8C20F5D" author="Wenzel, RL" journalOrPublisher="Resultats Scientifiques" pageId="377" pageNumber="378" pagination="235 - 266" title="XXVII - Histeridae from Ecuador. Mission zoologique Belge aux Iles Galapagos et en Ecuador (N. et J. Leleup 1964 - 1965)." volume="3" year="1976">Wenzel 1976</bibRefCitation>
: 259.
</paragraph>
</subSubSection>
<subSubSection id="6F00E8B5E5755762BA4C9671FC88E700" pageId="65" pageNumber="66" type="type locality">
<paragraph id="6A4018A5E7610D035088F831E422B8D2" pageId="65" pageNumber="66">Type locality.</paragraph>
<paragraph id="C6C0B734EA6CF0562222E14AE697FE27" pageId="65" pageNumber="66">
PANAMA: 
<normalizedToken id="936B4F143601E68122E6DB49AA9B9D7A" originalValue="Panamá">Panama</normalizedToken>
: Barro Colorado Island [
<geoCoordinate id="9171CEA0208EE17C1BC7A1DDD47C034A" direction="north" orientation="latitude" precision="925" value="9.183333">9°11'N</geoCoordinate>
, 
<geoCoordinate id="39D126E669BED15A989766838C3ED514" direction="west" orientation="longitude" precision="925" value="-79.85">79°51'W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection id="721E801F904DEB7624B0AA3EAD985148" pageId="65" pageNumber="66" type="type material">
<paragraph id="42AD3E5E4CFE6F0801B57EF730EA4256" pageId="65" pageNumber="66">Type material.</paragraph>
<paragraph id="E61B281D8FEE5D316F6C83FBA83C0A19" pageId="65" pageNumber="66">
Holotype male (FMNH): &quot;Barro Clrdo. Is. 
<normalizedToken id="84F12979894F7B5FC3D6D32161505AE0" originalValue="Panamá">Panama</normalizedToken>
, C.Z., 9-14-38 1, 211&quot; / &quot;Coll. by E.C. Williams 
<normalizedToken id="EA4CF00A04E30F58D8C5264AF4C33DB3" originalValue="”/”">&quot;/&quot;</normalizedToken>
pres. by R.L. Wenzel&quot;; examined 2006.
</paragraph>
</subSubSection>
<subSubSection id="A1A698AB5F848012E9581FC2C81E4283" lastPageId="69" lastPageNumber="70" pageId="65" pageNumber="66" type="other material">
<paragraph id="9847E930D7BB8E2D9F59244E36EAE1AE" pageId="65" pageNumber="66">Other material.</paragraph>
<paragraph id="39C9CB25D4314556F5001569D0A7F337" lastPageId="69" lastPageNumber="70" pageId="65" pageNumber="66">
COLOMBIA: 
<normalizedToken id="1A6C293BEA55DEA7F790F0CE301AFEA0" originalValue="Vaupés">Vaupes</normalizedToken>
: 9: Parque Nac. Mosiro-Itajura (
<normalizedToken id="C391E54E059965650261035FA4522734" originalValue="Caparú">Caparu</normalizedToken>
), Centro Ambiental, 
<geoCoordinate id="7D92E78610F4344C93645B932A288008" direction="south" orientation="latitude" precision="925" value="-1.0666667">1°04'S</geoCoordinate>
, 
<geoCoordinate id="B0C6C656BE14CD9C4FC9578F603FDD81" direction="west" orientation="longitude" precision="925" value="-69.51667">69°31'W</geoCoordinate>
, 60m, 20-30.i.2003, FIT, D. Arias &amp; M. Sharkey (IAVH, FMNH, MSCC, AKTC). COSTA RICA: Alajuela: 22: Eladio's, 
<normalizedToken id="77431378685D2D1069B5FDDCC7C3D141" originalValue="Peñas">Penas</normalizedToken>
Blancas Trail, 875m, 19.v.1989, FIT, J. Ashe, R. Brooks, R. Leschen (SEMC); same data except as noted: 25: 800m, 19.v.1989, FIT (SEMC), 1: 1190m, 20.v.1989, FIT (SEMC), 1: 970m, 20.v.1989, FIT (SEMC). Cartago: 1: Parque Nac. Barbilla, R.F. 
<pageBreakToken id="6C08AE497A45B9FA522105265A085297" pageId="66" pageNumber="67" start="start">Rio</pageBreakToken>
Pacuare. Turrialba, Send. Rio Danto, 500-600m, 28. 
<normalizedToken id="7A61B0618E33CE353D48264BCBA31A24" originalValue="viii–">viii-</normalizedToken>
5.ix.2001, FIT, W. Arana (INBIO); same data except as noted: 1:800m despues del R. Dantas, 400m, 19. 
<normalizedToken id="135F084FCDA242B1BC5D5CE839C149F1" originalValue="x–">x-</normalizedToken>
2.xi.2000, FIT, W. Arana (INBIO). Guanacaste: 1: Parque Nac. Guanacaste, 9km S Santa Cecilia, Est. Pitilla, 700m, ii.1990, C. Moraga &amp; R. Blanco, (INBIO), 2: ix.1991, (INBIO), 2: Est. Pitilla, 
<geoCoordinate id="81AA1EE4F59312099D968237CF03DBC8" direction="north" orientation="latitude" precision="15" value="10.989445">10°59'22&quot;N</geoCoordinate>
, 
<geoCoordinate id="3501912DE77857BE07C5825919F0973C" direction="west" orientation="longitude" precision="15" value="-85.42583">85°25'33&quot;W</geoCoordinate>
, 610m, 13-15.vii.2000, FIT, J. Ashe, R. Brooks, Z. Falin (SEMC); 1: Rio San Lorenzo, Tierras Morenas, Z.P. Tenorio, 1050m, i.1993, G. Rodriguez, (INBIO). Heredia: 6: La Selva Biol. Stn., 
<geoCoordinate id="F1A6178D17F34EC5B1FA004103D05654" direction="north" orientation="latitude" precision="925" value="10.433333">10°26'N</geoCoordinate>
, 
<geoCoordinate id="FD7F9217D5467ADC81B258DF1A819DDA" direction="west" orientation="longitude" precision="925" value="-84.01667">84°01'W</geoCoordinate>
, 19.vi.1998, FIT, C.E. Carlton &amp; A.K. Tishechkin (LSAM); same data except as noted: 2: 20.vi.1998, FIT (LSAM), 1: 21.vi.1998, FIT (LSAM), 10: 22.vi.1998, FIT (LSAM), 2: 23.vi.1998, FIT (LSAM), 2: 24.vi.1998, FIT (LSAM), 1: 26.vi.1998, FIT (LSAM), 2: 29.vi.1998, FIT (LSAM), 1: 50-150m, 5-8.iii.2001, FIT, E.G. Riley (TAMU), 2: 6.vii.2005, FIT, M. Ferro (LSAM), 2: 23.vii.1976, H.A. Hespenheide, (INBIO), 1: 100m, 11.iii.1992, FIT, W. Bell (SEMC), 1: 10.vi.2012, OTS Beetle Course, DNA Extract MSC-2306 (AKTC). 
<normalizedToken id="FA03D016FB1119969455525EA1800D75" originalValue="Limón">Limon</normalizedToken>
: 2: Area Cons. Tortuguero, Sector Cerro Cocori, Fca. de E. Rojas, 150m, i.1993, E. Rojas, (INBIO), 4: ix.1991, E. Rojas, (INBIO), 3: viii.1991, E. Rojas, (INBIO), 2: Parque Nac. Tortuguero, Est. Cuatro Esquinas, 0m, v.1990, J. Solano, (INBIO), 2: vi.1990, J. Solano, (INBIO), 1: v.1990, (INBIO); 1: Res. Biol. Hitoy Cerere, send. Plano Espavel, 180m, 24.iii.2002, Red de Golpe, W. Arana (INBIO); 1: Sardinas, Barra del Colorado, 15m, 28.i-12.ii.1995, F. Araya, (INBIO). Puntarenas: 1: Est. Biol. Las Cruces, 
<geoCoordinate id="602610BC3AA105572FA896321A280BB7" direction="north" orientation="latitude" precision="9" value="8.785666">8°47.14'N</geoCoordinate>
, 
<geoCoordinate id="C0453949E4FA141EB666AA14CCE4C560" direction="west" orientation="longitude" precision="9" value="-82.95966">82°57.58'W</geoCoordinate>
, 1330m, 28-30.v.2004, FIT, J.S. Ashe, Z. Falin, I. Hinojosa (SEMC); 1: 
<pageBreakToken id="EE7A4D3195EBBBD1A7368E118848A66C" pageId="67" pageNumber="68" start="start">Wilson</pageBreakToken>
Botanical Garden, 1200m, 27.v.1993, FIT, J.S. &amp; A.K. Ashe (SEMC); 1: Est. Esquinas, Peninsula de Osa, 0m, i.1993, F. Quesada, (INBIO), 1: iv.1993, M. Segura, (INBIO), 1: 200m, iv.1993, J.F. Quesada, (INBIO); 1: Parque Nac. Corcovado, Est. Sirena, 0-100m, i.1990, G. Fonseca, (INBIO), 1: x.1990, C. Saborio, (INBIO); 1: Corcovado Trail, 
<geoCoordinate id="90A66638BEC912B8D31228536BE586C3" direction="north" orientation="latitude" precision="15" value="8.485278">8°29'7&quot;N</geoCoordinate>
, 
<geoCoordinate id="E1F75B5387C89A6C8D6EFD20C7049EDA" direction="west" orientation="longitude" precision="15" value="-83.5775">83°34'39&quot;W</geoCoordinate>
, 150m, 28.vi-1.vii.2000, FIT, Z. Falin (SEMC); 2: upper Ollas Trail, 
<geoCoordinate id="6E45AD2EFD9FE67BFC07DCD1B7CB66C8" direction="north" orientation="latitude" precision="15" value="8.485278">8°29'7&quot;N</geoCoordinate>
, 
<geoCoordinate id="B3253873CA810163B99D69FBA4EC008E" direction="west" orientation="longitude" precision="15" value="-83.5775">83°34'39&quot;W</geoCoordinate>
, 140m, 24-28.vi.2000, FIT, Z. Falin (SEMC); 1: Rancho Quemado, Peninsula de Osa, 200m, v.1991, F. Quesada, (INBIO), 3: v.1991, J.C. Saborio, (INBIO), 2: v.1992, F. Quesada y G. Varela, (INBIO); 1: Res. Biol. Carara, Est. Quebrada Bonita, 100m, i.1995, R. Guzman, (INBIO); 2: Laguna Meandrica, 100m, vi.1990, R. Zuniga, (INBIO); 1: Reserva Forestal Golfo Dulce, Est. Agujas, Golfito, 250-350m, 26.v.2000, FIT, A. Azofeifa (INBIO); 1: Rincon de Osa, 
<geoCoordinate id="2F9546069EC91844C0BF4ED61B346A53" direction="north" orientation="latitude" precision="1" value="8.685683">8°41.141'N</geoCoordinate>
, 
<geoCoordinate id="1A9138F61A7CA14FA295C706C0EF333D" direction="west" orientation="longitude" precision="1" value="-83.518616">83°31.117'W</geoCoordinate>
, 150m, 23-26.vi.2001, FIT, S. &amp; J. Peck (SEMC), 2: 40m, 23-26.vi.2001, FIT, stream side, S. &amp; J. Peck (SEMC). ECUADOR: Orellana: 1:Parque Nac. 
<normalizedToken id="8E9A3F6D2CBED064E0B381A36CDCC800" originalValue="Yasuní">Yasuni</normalizedToken>
, Via Maxus at Puente 
<normalizedToken id="47DC9633C8A8CDD1ADADCD7036186B9A" originalValue="Piraña">Pirana</normalizedToken>
, 
<geoCoordinate id="275DE3456BCC42494DE856A328BB5EE5" direction="south" orientation="latitude" precision="92" value="-0.65833336">0°39.5'S</geoCoordinate>
, 
<geoCoordinate id="2C7EDE8AC783A429DB52ABA848D74456" direction="west" orientation="longitude" precision="925" value="-76.433334">76°26'W</geoCoordinate>
, 14-20.vii.2008, FIT, A.K. Tishechkin (AKTC), 1: 24.vii.2008, FIT, A.K. Tishechkin (AKTC); 2: Tiputini Biodiversity Station, 
<geoCoordinate id="C26C855E2071A0C413AEE6A99BE175AB" direction="south" orientation="latitude" precision="5" value="-0.6376">0.6376°S</geoCoordinate>
, 
<geoCoordinate id="2EF76AA1C2BC778EA5B2C7E5300AFD35" direction="west" orientation="longitude" precision="5" value="-76.1499">76.1499°W</geoCoordinate>
, 2-9.vi.2011, FIT, M.S. Caterino &amp; A.K. Tishechkin (FMNH, USFQ). Pastaza: 1: Cushuime, on Rio Cushuime, 150km SE Puyo, 320m, 19.v.1971, polypore fungus, B. Malkin (FMNH). FRENCH GUIANA: 1: Mont tabulaire, 
<normalizedToken id="09BB3E73E139307D24A688FD83A095CD" originalValue="Itoupé">Itoupe</normalizedToken>
, 
<geoCoordinate id="325C6B94B834F6733FC47DA729BD7D3D" direction="north" orientation="latitude" precision="9" value="3.023">3°1.38'N</geoCoordinate>
, 
<geoCoordinate id="8B17B9C51A91C3BC030010FA7EB8FF5A" direction="west" orientation="longitude" precision="9" value="-53.0955">53°5.73'W</geoCoordinate>
, 570m, 24.iii.2010, FIT, SEAG (CHND); 1: Montagne des Chevaux, 
<geoCoordinate id="455F79D0B8AA36100470ADBDEEA08055" direction="north" orientation="latitude" precision="925" value="4.7166667">4°43'N</geoCoordinate>
, 
<geoCoordinate id="2B59D3F3B39D6786D17D42F57E0285A7" direction="west" orientation="longitude" precision="925" value="-52.4">52°24'W</geoCoordinate>
, 18.iv.2009, FIT, SEAG (CHND). HONDURAS: 
<normalizedToken id="37FDF02CC573FA10D3A0CA3A7CD99C13" originalValue="Cortés">Cortes</normalizedToken>
: 1: Lago Yojoa, 650m, 23-28.viii.1994, FIT, tropical, evergreen forest, S. &amp; J. Peck (CMNC); 3: Parque Nac. Cerro Azul-Meambar, Los Pinos, 
<geoCoordinate id="EB07937AF8EE70C92C92388DF8C74988" direction="north" orientation="latitude" precision="92" value="14.873333">14°52.4'N</geoCoordinate>
, 
<geoCoordinate id="C6DDB1F7BE95F95576ED375141254A94" direction="west" orientation="longitude" precision="92" value="-87.91167">87°54.7'W</geoCoordinate>
, 800m, 10-16.v.2001, FIT, S. Peck (SEMC); 4: Yojoa Lake, Deer Island, 
<geoCoordinate id="E6146B56660989BF7CB140516FFB4C02" direction="north" orientation="latitude" precision="925" value="14.916667">14°55'N</geoCoordinate>
, 
<geoCoordinate id="99B1B7C26B324D1D981E908D0BA5136F" direction="west" orientation="longitude" precision="925" value="-87.96667">87°58'W</geoCoordinate>
, 670m, 19-21.vi.1994, FIT, J. Ashe, R. Brooks (SEMC), 13: 22-26.vi.1994, FIT, J. Ashe, R. Brooks (SEMC); Santa Barbara: 1: La Fe, Finca La Roca 5.3km S. Pena Blanca, 
<geoCoordinate id="2B3A485C445D2E7861050DF7DA43713F" direction="north" orientation="latitude" precision="925" value="14.95">14°57'N</geoCoordinate>
, 
<geoCoordinate id="0A2263C958197F595D723ACB14766091" direction="west" orientation="longitude" precision="925" value="-88.03333">88°02'W</geoCoordinate>
, 740m, 19-21.vi.1994, FIT, J.S. Ashe &amp; R. Brooks (SEMC). MEXICO: Veracruz: 1: Cordoba, 4.viii.1969, tropical evergreen forest Berlese, S. Peck (FMNH). NICARAGUA: Granada: 3: Res. Nat. Volcan Mombacho, 
<geoCoordinate id="6CEC3F926E0E9463687FEC3AFD8C4BC6" direction="north" orientation="latitude" precision="925" value="11.833333">11°50'N</geoCoordinate>
, 
<geoCoordinate id="BAFEB35C3FC70507C46B41662626C29D" direction="west" orientation="longitude" precision="92" value="-85.98">85°58.8'W</geoCoordinate>
, 1150m, 2-5.vi.2002, FIT, elfin montane forest, S. Peck (CMNC); 2: Volcan Mombacho, El Progreso #2, 30.vi.1998, malaise trap, J.M. Maes (MEL); Rio San Juan: 1: Refugio Bartola, 60km SE San Carlos, 
<geoCoordinate id="542862E90E6E7B490CD597BADE502EE3" direction="north" orientation="latitude" precision="9" value="10.973333">10°58.40'N</geoCoordinate>
, 
<geoCoordinate id="8AE3BC00627C409C7E35798C9C0854B5" direction="west" orientation="longitude" precision="9" value="-84.33833">84°20.30'W</geoCoordinate>
, 100m, 28-30.v.2002, FIT, Z. Falin, S. Chatzimanolis (SEMC), 3: 30m, 23-31.v.2002, FIT, S. Peck (SEMC). PANAMA: 
<normalizedToken id="33C5FFBF0C40FB8ECBC3E6AB28EB2445" originalValue="Chiriquí">Chiriqui</normalizedToken>
: 1: La Fortuna, &quot;Cont. Divide Trail&quot;, 
<geoCoordinate id="E63E65125A64F4CD4788C41A5B43AB5D" direction="north" orientation="latitude" precision="925" value="8.766666">8°46'N</geoCoordinate>
, 
<geoCoordinate id="0087F9CA2D241C06DB2C81DA3C5CDBB8" direction="west" orientation="longitude" precision="925" value="-82.2">82°12'W</geoCoordinate>
, 1150m, 23. 
<normalizedToken id="F91D1C42FB44AE5E773231695939084C" originalValue="v–">v-</normalizedToken>
9.vi.1995, FIT, J. Ashe, R. Brooks (SEMC); 1: La Fortuna, &quot;Hydro. Trail&quot;, 
<geoCoordinate id="979D325BC5D970E373F33619E5FDF138" direction="north" orientation="latitude" precision="925" value="8.7">8°42'N</geoCoordinate>
, 
<geoCoordinate id="A402D2B89D3F279F436B96F56F7B0A78" direction="west" orientation="longitude" precision="925" value="-82.23333">82°14'W</geoCoordinate>
, 1150m, 21-23.v.1995, FIT, J. &amp; A. Ashe (SEMC); 1: Reserva La Fortuna, 
<geoCoordinate id="8B99FE8A36228F4BFE55F2A8D8A39666" direction="north" orientation="latitude" precision="15" value="8.721666">8°43'18&quot;N</geoCoordinate>
, 
<geoCoordinate id="C519CF0DE1CDBE2ABC55BD2453FF6AFA" direction="west" orientation="longitude" precision="15" value="-82.23805">82°14'17&quot;W</geoCoordinate>
, 3900ft, 4-10.viii.1999, FIT, Schaffner &amp; Woolley (TAMU); 
<normalizedToken id="E22D2CC16A00745AB10015E0C56B127A" originalValue="Colón">Colon</normalizedToken>
: 40: San Lorenzo Forest, 
<geoCoordinate id="813947B4B164360B5FD5030AE88F4289" direction="north" orientation="latitude" precision="925" value="9.283334">9°17'N</geoCoordinate>
, 
<geoCoordinate id="2E8806A9B938ADCEFB1BBEEB2FF16CA8" direction="west" orientation="longitude" precision="925" value="-79.96667">79°58'W</geoCoordinate>
, 11.v.2004, 12.v.2004, FIT, A.K. Tishechkin (LSAM), 18: 12-13.v.2004 (LSAM), 10: 13-14.v.2004 (GBFM, LSAM), 10: 14-15.v.2004 (LSAM), 2: 15-17.v.2004 (LSAM), 6: 18-19.v.2004 (GBFM, LSAM), 5: 19-20.v.2004 (LSAM), 1: 20-21.v.2004 (LSAM), 1: 28-30.ix.2003 (LSAM), 2: 21-24.v.2004 (LSAM), 1: 24-25.v.2004 (LSAM), 2: 25-26.v.2004 (LSAM), 1: 26-29.v.2004 (LSAM), 4: 29.v.2004 (LSAM), 1: 12-13.v.2004 (LSAM), 5: 12-23. 
<pageBreakToken id="29681A164317EB067EC205A7B3479C73" pageId="68" pageNumber="69" start="start">ix</pageBreakToken>
.2004, M. Rapp (LSAM), 1: 12-19.v.2004, R. Didham (LSAM), 5: 14-26.vii.2004, M. Rapp (LSAM), 10: 15-25.v.2004, R. Didham (LSAM), 2: 23. 
<normalizedToken id="AACC3D4E735FEE5601747A65538FB0D9" originalValue="x–">x-</normalizedToken>
2.xi.2004, M. Rapp (LSAM), 5: 24.vi-4.vii.2004, M. Rapp (GBFM, LSAM), 3: 25. 
<normalizedToken id="C38EF4CD0499C44E6F513079987E5094" originalValue="v–">v-</normalizedToken>
4.vi.2004, R. Didham (LSAM), 2: 25. 
<normalizedToken id="E8C5F033D1B9E300D58E81B79DAA27E1" originalValue="v–">v-</normalizedToken>
5.vi.2004, R. Didham (LSAM), 2: 25.vi-5.vii.2004, M. Rapp (LSAM), 1: 3-13.viii.2004, M. Rapp (LSAM), 3: 3-13.x.2004, M. Rapp (LSAM), 2: 4-17.vi.2004, M. Rapp (LSAM), 1: 4-14.viii.2004, M. Rapp (LSAM), 4: 5-18.vi.2004, M. Rapp (LSAM), 1: 1.3m, 25. 
<normalizedToken id="611AA732D4866A95663102840ED5C77C" originalValue="v–">v-</normalizedToken>
5.vi.2004, R. Didham (LSAM), 1: 14m, 25.iv-5.v.2004, M. 
<pageBreakToken id="1D5595E9AF4B4FEC2C84A95BB4BDA846" pageId="69" pageNumber="70" start="start">Gonzalez</pageBreakToken>
(LSAM), 1: 7m, 12-23.ix.2004, M. Rapp (LSAM); 
<normalizedToken id="C258F41F2C1FD582C4FD25197F93FC23" originalValue="Darién">Darien</normalizedToken>
: 1: Cana Biological Station, 
<geoCoordinate id="ED014E1A0B086CE8E7934DE93600BFA6" direction="north" orientation="latitude" precision="15" value="7.755">7°45'18&quot;N</geoCoordinate>
, 
<geoCoordinate id="464B43620C3FE60E844C80A2BEEC1794" direction="west" orientation="longitude" precision="15" value="-77.685">77°41'6&quot;W</geoCoordinate>
, 600m, 7-9.vi.1996, FIT, J. Ashe, R. Brooks (SEMC); 
<normalizedToken id="610A995987069C67F3283D4D5DCAEF89" originalValue="Panamá">Panama</normalizedToken>
: 3: Barro Colorado Isl., 
<geoCoordinate id="BFB781000D1D7DE2D0AC057478BDEFE6" direction="north" orientation="latitude" precision="925" value="9.183333">9°11'N</geoCoordinate>
, 
<geoCoordinate id="1AE8586245DDAAC458592913DD50E4A9" direction="west" orientation="longitude" precision="925" value="-79.85">79°51'W</geoCoordinate>
, 1.viii.1994, FIT, D. Banks, 1: 16.viii.1994, FIT, D. Banks, 1: 21-22.vii.2000, FIT, S. Chatzimanolis (SEMC), 1: 24.viii.1994, D. Banks, 1: 29.vii.1994, D. Banks, 2: 6.viii.1994, D. Banks, 1: 7.vii.1994, D. Banks, 3: 8.vii.1994, D. Banks, 1: Cano Saddle Gatun Lake, Close's Pltn, 12.v.1923, RC Shannon (USNM); 1: Cerro Azul, ca., 2000ft, 21.ii.1976, flood debris, A.F. Newton (FMNH); 1: Pipeline Rd., 17-22.vi.1993, FIT, S. Lingafelter (SEMC); PERU: Cusco: 1: Consuelo, Manu rd. km 165, 6-7.x.1982, FIT, L. Watrous &amp; G. Mazurek (FMNH); 1: 6.x.1982, rotten palm, L. Watrous &amp; G. Mazurek (FMNH); 1: 7-8.x.1982, FIT, L. Watrous &amp; G. Mazurek (FMNH); 
<normalizedToken id="05BE03DF5B7F00938537CF0BB67CD6F7" originalValue="Junín">Junin</normalizedToken>
: 2: 11km NE Puerto Ocopa, Los Olivos, 
<geoCoordinate id="5A3460899EC5DAAC254B2A75C9C42848" direction="south" orientation="latitude" precision="9" value="-11.116667">11°7.00'S</geoCoordinate>
, 
<geoCoordinate id="DDB92E7CE835CC1CAD249F1110623AEB" direction="west" orientation="longitude" precision="9" value="-74.25867">74°15.52'W</geoCoordinate>
, 1200m, 26-28.iii.2009, Window trap, A.V. Petrov (AKTC, MUSM); Loreto: 4: 1.5km N Teniente Lopez, 
<geoCoordinate id="2E275491679F4727EEABA46F89F17063" direction="south" orientation="latitude" precision="9" value="-2.5943334">2°35.66'S</geoCoordinate>
, 
<geoCoordinate id="4A38058BDE7C638A3487DC136B8111B1" direction="west" orientation="longitude" precision="9" value="-76.11533">76°06.92'W</geoCoordinate>
, 210-240m, 18.vii.1993, FIT, R. Leschen (SEMC), 5: 20.vii.1993, FIT, R. Leschen (SEMC); 1: 68km SW Iquitos to Nauta, Rio Itaya, 
<geoCoordinate id="8BE5F8887030C8DB32276EB5E5AB271B" direction="south" orientation="latitude" precision="925" value="-4.1833334">4°11'S</geoCoordinate>
, 
<geoCoordinate id="7A476AC80BE395CA6FE193D0524537DB" direction="west" orientation="longitude" precision="925" value="-73.433334">73°26'W</geoCoordinate>
, 110m, 18.ii.2008, A.V. Petrov (AKTC); 2: km 63, rd. Iquitos-Nauta, Rio Itaya, 
<geoCoordinate id="1D305C483F4FDB753B425A8675898C5C" direction="south" orientation="latitude" precision="1" value="-4.2534165">4°15.205'S</geoCoordinate>
, 
<geoCoordinate id="A77964279781D9D5909AB0D497B1C00F" direction="west" orientation="longitude" precision="1" value="-73.43457">73°26.074'W</geoCoordinate>
, 140m, 10-14.ii.2010, A.V. Petrov (AKTC), 1: 9-13.i.2011, A.V. Petrov (MUSM); Madre de Dios: 1: Manu National Park, Pantiacolla Lodge, Alto Madre de Dios River, 
<geoCoordinate id="8D01B9F2F48D6A3AB2B1E6237C03AFD4" direction="south" orientation="latitude" precision="92" value="-12.655">12°39.3'S</geoCoordinate>
, 
<geoCoordinate id="596050AF753A44DD65B526207A5A11F3" direction="west" orientation="longitude" precision="92" value="-71.23167">71°13.9'W</geoCoordinate>
, 420m, 14-19.xi.2007, FIT, D. Brzoska (SEMC).
</paragraph>
</subSubSection>
<subSubSection id="A9B63809942A93B0D476F74EC098C14F" pageId="69" pageNumber="70" type="diagnosis">
<paragraph id="E229C9DCF4B98665EC1D8AC63703C7E9" pageId="69" pageNumber="70">Diagnosis.</paragraph>
<paragraph id="03E42786F4DFEAA0816C85028E72F15C" pageId="69" pageNumber="70">
Length: 2.18-2.84 mm, width: 1.75-2.37 mm; body rufo-brunneus; frons with median portion of frontal stria well impressed; supraorbital stria briefly interrupted at vertex; pronotum with anterior submarginal stria detached from lateral portion, slightly recurved posterad at apices, more or less parallel to outwardly arcuate anterior emargination, anterior marginal stria interrupted for short distance at middle; elytra generally lacking any fragments of inner subhumeral stria, sutural stria nearly or fully complete, frequently with the 4th stria arched over at the base nearly meeting it; pygidium with punctation mainly fine, dense, with larger punctures interspersed only near the base, marginal sulcus complete, deep, crenulate. Male genitalia (Figs 18 
<normalizedToken id="39C50C219F5A5E842580EEBCBE3404CC" originalValue="A–D">A-D</normalizedToken>
, F): T8 with unique secondary sclerotization along edge of apical emargination; S9 narrow, evenly expanded to apex, with midline sclerotization distinct; tegmen abruptly narrowed toward apex, strongly narrowed and slightly bent near base in lateral aspect.
</paragraph>
</subSubSection>
<subSubSection id="456298345CADA5A13EAFE7486314332A" pageId="69" pageNumber="70" type="remarks">
<paragraph id="D111A956E1E77405B229A4052886684A" pageId="69" pageNumber="70">Remarks.</paragraph>
<paragraph id="716CB7BA5B56F99926E8C42E5B7937E7" pageId="69" pageNumber="70">
The best characters for separating 
<taxonomicName id="B3C33077A20BB24896CEEAF6ABD26BB6" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
and 
<taxonomicName id="96C026AC858C31136AC774DD2434C2D7" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
are the details of the anterior pronotal margin and submarginal stria, with the central portion of the pronotal margin outwardly arcuate in 
<taxonomicName id="19991B1978065A29FE37AF0894088E5B" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
(Fig. 17A), and the anterior submarginal stria always detached from the lateral submarginal. In 
<taxonomicName id="40F91286D76685091AA2E3FAC0EFA44A" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
, the anterior margin is not outwardly arcuate (Fig. 17C), and the lateral and anterior portions of the submarginal pronotal stria are usually connected behind the median pronotal gland openings. The body color and pygidium also fairly readily distinguish the species, with 
<taxonomicName id="669B57FCF86CB3BCE07D3A9824B4764F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
consistently darker (Fig. 17C), and with a more coarsely punctate pygidium (Fig. 17D).
</paragraph>
<caption id="E89C87DE58AE4C25D23AA4843CA558E8" pageId="69" pageNumber="70">
<paragraph id="8D80C0C59C950060F68658CBAF6A4839" pageId="69" pageNumber="70">
Figure 17. 
<taxonomicName id="31E717403E46D33B46BC97EACE4BB768" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
group. A Dorsal habitus 
<taxonomicName id="94FCA95C128665BDDE8CB62436EC3791" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
(holotype) B Pygidia of 
<taxonomicName id="5894EF070890FAC5A1D2A92C9A7D034D" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
C Dorsal habitus 
<taxonomicName id="8A032445C9DDDDBB4DA89622EF5ABBD2" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
D Pygidia of 
<taxonomicName id="48A7EA361EDF195729A71CED9226E49D" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
.
</paragraph>
</caption>
<caption id="5F1D7BF73DB8AB7BCE5D5DD81B54EABB" pageId="69" pageNumber="70">
<paragraph id="E439145167D9A320B5F6F694EE29C49D" pageId="69" pageNumber="70">
Map 6. Records of the 
<taxonomicName id="38A5613D7BA2D5578C1F7385C62B7786" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
group.
</paragraph>
</caption>
<caption id="F63F223C5EE7165F7213F262C80C0F8A" pageId="69" pageNumber="70">
<paragraph id="A61D4E86677A23D3ABE2F332DB06D722" pageId="69" pageNumber="70">
Figure 18. Male genitalia of 
<taxonomicName id="5256600D269BACE06BF0948E4FA5DB53" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
group. A T8 of 
<taxonomicName id="4870A04B3C62259FB19EB8491911096D" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
B S8 of 
<taxonomicName id="A18C93EF181E5274702ECD5D1DBF7355" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
C T9 &amp; T10 of 
<taxonomicName id="6541E25CCAA3B88D72DD3C3264A1876F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
D S9 of 
<taxonomicName id="5C2C55F54F94606CB325B23D23C54911" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
E S9 of 
<taxonomicName id="E735D246F299978B1644C9E9F565D693" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
F Aedeagus, dorsal and lateral views, of 
<taxonomicName id="70855CF8B58E9E13C6E9CBDFB870C9EB" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
G Aedeagus, dorsal and lateral views, of 
<taxonomicName id="4B3896A4DCA16DA0A01D869AE18840E2" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>