319 lines
32 KiB
XML
319 lines
32 KiB
XML
<document ID-DOI="10.37520/aemnp.2022.012" ID-ISSN="1804-6487" ID-Zenodo-Dep="7161302" ID-ZooBank="D3830DDC-6589-4B0C-8DEE-015053395254" approvalRequired="11" approvalRequired_for_textStreams="10" approvalRequired_for_treatments="1" checkinTime="1665202473490" checkinUser="felipe" docAuthor="Švec, Zdeněk" docDate="2022" docId="8C011006FFCDFFF5FC5BFF32FAE8E115" docLanguage="en" docName="ActaEntMusNatPra.62.1.155-164.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2022.012" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.6:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="6" docTitle="Agathidium (Agathidium) pilulum Švec 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="158" masterDocId="7038687EFFCFFFF6FFDAFFC5FFE0E700" masterDocTitle="Leiodidae (Coleoptera) of the Hainan Island with new faunistic records from China and with notes on the unique body modifications in the genus Agathidium" masterLastPageNumber="164" masterPageNumber="155" pageNumber="157" updateTime="1665202904673" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Leiodidae (Coleoptera) of the Hainan Island with new faunistic records from China and with notes on the unique body modifications in the genus Agathidium</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Švec, Zdeněk</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Acta Entomologica Musei Nationalis Pragae</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2022</mods:date>
|
||
<mods:detail type="series">
|
||
<mods:title>Acta. Ent. Mus. Natl. Pragae</mods:title>
|
||
</mods:detail>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2022-08-28</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>62</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>155</mods:start>
|
||
<mods:end>164</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.37520/aemnp.2022.012</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.37520/aemnp.2022.012</mods:identifier>
|
||
<mods:identifier type="ISSN">1804-6487</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">7161302</mods:identifier>
|
||
<mods:identifier type="ZooBank">D3830DDC-6589-4B0C-8DEE-015053395254</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7161314" ID-Zenodo-Dep="7161314" LSID="urn:lsid:plazi:treatment:8C011006FFCDFFF5FC5BFF32FAE8E115" httpUri="http://treatment.plazi.org/id/8C011006FFCDFFF5FC5BFF32FAE8E115" lastPageId="3" lastPageNumber="158" pageId="2" pageNumber="157">
|
||
<subSubSection box="[897,1368,247,271]" pageId="2" pageNumber="157" type="nomenclature">
|
||
<paragraph blockId="2.[897,1368,247,271]" box="[897,1368,247,271]" pageId="2" pageNumber="157">
|
||
<heading box="[897,1368,247,271]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="157" reason="3">
|
||
<emphasis bold="true" box="[897,1368,247,271]" pageId="2" pageNumber="157">
|
||
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[897,1275,247,271]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="species" species="pilulum" status="sp. nov." subGenus="Agathidium">
|
||
<emphasis bold="true" box="[897,1027,247,271]" italics="true" pageId="2" pageNumber="157">Agathidium</emphasis>
|
||
(
|
||
<emphasis bold="true" box="[1043,1173,247,271]" italics="true" pageId="2" pageNumber="157">Agathidium</emphasis>
|
||
)
|
||
<emphasis bold="true" box="[1189,1275,247,271]" italics="true" pageId="2" pageNumber="157">pilulum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1282,1368,248,271]" pageId="2" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[1072,1193,278,297]" pageId="2" pageNumber="157" type="description">
|
||
<paragraph blockId="2.[828,1438,278,2070]" box="[1072,1193,278,297]" pageId="2" pageNumber="157">
|
||
(
|
||
<figureCitation box="[1078,1155,278,297]" captionStart="Figs 1–8. 1, 2, 6, 7" captionStartId="3.[151,188,1016,1035]" captionTargetBox="[188,1406,191,997]" captionTargetId="figure-669@3.[158,1431,157,997]" captionTargetPageId="3" captionText="Figs 1–8. 1, 2, 6, 7 – aedeagus laterally and tip of aedeagus dorsally. 3, 5 – lateral view of body. 4, 8 – spermatheca. 1–4 – Agathidium (Agathidium) pilulum sp. nov.; 5–8 – Agathidium (Macroceble) cornigerum sp. nov. Scale bars in Figs 1, 2, 4, 6, 7, 8 = 0.1 mm; 3, 5 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7161304" httpUri="https://zenodo.org/record/7161304/files/figure.png" pageId="2" pageNumber="157">Figs 1–4</figureCitation>
|
||
,
|
||
<figureCitation box="[1164,1186,278,297]" captionStart="Figs 13–16. 13, 15, 16" captionStartId="5.[149,186,1096,1115]" captionTargetBox="[150,1435,189,1072]" captionTargetId="figure-486@5.[150,1435,189,1072]" captionTargetPageId="5" captionText="Figs 13–16. 13, 15, 16 – head dorsally; 14 – head laterally. 13 – Agathidium (Agathidium) pilulum sp. nov.; 14, 16 – Agathidium (Macroceble) cornigerum sp. nov.; 15 – Agathidium (Macroceble) apiforme sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7161308" httpUri="https://zenodo.org/record/7161308/files/figure.png" pageId="2" pageNumber="157">13</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="157" type="materials_examined">
|
||
<paragraph blockId="2.[828,1438,278,2070]" pageId="2" pageNumber="157">
|
||
<emphasis bold="true" box="[828,946,313,332]" pageId="2" pageNumber="157">
|
||
<typeStatus box="[828,873,313,332]" pageId="2" pageNumber="157">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<collectingCountry box="[959,1012,313,332]" name="China" pageId="2" pageNumber="157">China</collectingCountry>
|
||
,
|
||
<collectingRegion box="[1023,1085,313,332]" country="China" name="Hainan" pageId="2" pageNumber="157">Hainan</collectingRegion>
|
||
Island, Limushan Mts, 19°10.5–10.9′N, 109°44–45′E,
|
||
<quantity box="[950,1049,339,358]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="9.0" metricValueMin="6.5" pageId="2" pageNumber="157" unit="m" value="775.0" valueMax="900.0" valueMin="650.0">
|
||
650–
|
||
<specimenCount box="[993,1049,339,358]" pageId="2" pageNumber="157" type="generic">900 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="2.[828,1438,278,2070]" pageId="2" pageNumber="157">
|
||
<emphasis bold="true" box="[828,954,373,392]" pageId="2" pageNumber="157">Type material</emphasis>
|
||
. Hඈඅඈඍඒඉൾ: J, “
|
||
<collectingCountry box="[1094,1166,373,392]" name="China" pageId="2" pageNumber="157">CHINA</collectingCountry>
|
||
:
|
||
<collectingRegion box="[1173,1234,373,392]" country="China" name="Hainan" pageId="2" pageNumber="157">Hainan</collectingRegion>
|
||
isl. [MF 19] / Limushan Mts, mountains / above first admin.centre / 19°10.5–10.9′N 109°44–45′E/
|
||
<quantity box="[828,923,427,446]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="9.0" metricValueMin="6.5" pageId="2" pageNumber="157" unit="m" value="775.0" valueMax="900.0" valueMin="650.0">650-900 m</quantity>
|
||
,
|
||
<date box="[931,1006,427,446]" pageId="2" pageNumber="157" value="2011-05-06">6.v.2011</date>
|
||
, Fikáček // sifting small accumula- / tions of moist leaf litter along / and on the trail in secondary / forest partly with
|
||
<taxonomicName box="[1366,1437,453,472]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="2" pageNumber="157" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[1366,1437,453,472]" italics="true" pageId="2" pageNumber="157">Cyathea</emphasis>
|
||
</taxonomicName>
|
||
and / bamboo” (
|
||
<collectionCode box="[963,1028,480,499]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="157">NMPC</collectionCode>
|
||
). Pൺඋൺඍඒඉൾඌ: 5 JJ
|
||
<specimenCount box="[1193,1239,474,499]" pageId="2" pageNumber="157" type="female">2 ♀♀</specimenCount>
|
||
, same data as
|
||
<typeStatus box="[1362,1437,480,499]" pageId="2" pageNumber="157">holotype</typeStatus>
|
||
(
|
||
<collectionCode box="[834,898,507,526]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="157">NMPC</collectionCode>
|
||
,
|
||
<collectionCode box="[906,961,507,526]" pageId="2" pageNumber="157">ZSPC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="3" lastPageNumber="158" pageId="2" pageNumber="157" type="description">
|
||
<paragraph blockId="2.[828,1438,278,2070]" pageId="2" pageNumber="157">
|
||
<emphasis bold="true" box="[828,967,543,566]" pageId="2" pageNumber="157">Description.</emphasis>
|
||
Length of body in
|
||
<typeStatus box="[1184,1277,543,566]" pageId="2" pageNumber="157">holotype</typeStatus>
|
||
<quantity box="[1287,1373,543,566]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="157" unit="mm" value="1.9">1.9 mm</quantity>
|
||
, head
|
||
<quantity box="[828,915,575,598]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="157" unit="mm" value="0.3">0.3 mm</quantity>
|
||
, pronotum
|
||
<quantity box="[1041,1128,575,598]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="157" unit="mm" value="0.7">0.7 mm</quantity>
|
||
, elytra
|
||
<quantity box="[1211,1298,575,598]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="157" unit="mm" value="0.9">0.9 mm</quantity>
|
||
, antenna
|
||
<quantity metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="157" unit="mm" value="0.6">0.6 mm</quantity>
|
||
, aedeagus
|
||
<quantity box="[983,1080,607,630]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="2" pageNumber="157" unit="mm" value="0.74">0.74 mm</quantity>
|
||
. Maximum width of head
|
||
<quantity box="[1351,1433,607,630]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="157" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, pronotum
|
||
<quantity box="[938,1022,639,662]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="157" unit="mm" value="1.2">1.2 mm</quantity>
|
||
, elytra
|
||
<quantity box="[1101,1186,639,662]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="157" unit="mm" value="1.2">1.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="2.[828,1438,278,2070]" pageId="2" pageNumber="157">Body short oval. Pronotum chestnut, elytra dark chestnut with feebly expressed reddish narrow strip along suture, head lighter than pronotum. Legs and antennae yellow-red with A9 and A10 a little darker. Ventral surface yellow-brown. Dorsum punctured, without microsculpture. Sutural striae absent; femoral lines developed, supraocular carina raised anteriorly.</paragraph>
|
||
<paragraph blockId="2.[828,1438,278,2070]" pageId="2" pageNumber="157">
|
||
Head. Maximum width of head far before posterior margin of eyes. Eyes parabola slice-shaped. Supraocular carina low posteriorly, raised anteriorly. Subocular line not developed. Clypeus straight, feebly emarginate, clypeal line lacking (
|
||
<figureCitation box="[975,1058,1023,1046]" captionStart="Figs 13–16. 13, 15, 16" captionStartId="5.[149,186,1096,1115]" captionTargetBox="[150,1435,189,1072]" captionTargetId="figure-486@5.[150,1435,189,1072]" captionTargetPageId="5" captionText="Figs 13–16. 13, 15, 16 – head dorsally; 14 – head laterally. 13 – Agathidium (Agathidium) pilulum sp. nov.; 14, 16 – Agathidium (Macroceble) cornigerum sp. nov.; 15 – Agathidium (Macroceble) apiforme sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7161308" httpUri="https://zenodo.org/record/7161308/files/figure.png" pageId="2" pageNumber="157">Fig. 13</figureCitation>
|
||
). Antennomere 3 distinctly longer than A2 (A3/A2 = 1.5). Ratio of lengths of A2–A11 (A2 = 1.0): 1.0-1.5-0.8-0.6-0.5-0.5-0.6-1.3-1.1-2.0. Ratio of widths of A2–A11 (A2 = 1.0): 1.0-0.5-0.7-0.8-1.0-1.2- 1.3-1.7-1.8-1.8. Ratio of W/L of A2–A11: 0.8-0.3-0.7- 1.0-1.5-1.8-1.6-1.0-1.2-0.7. Surface of head smooth with very fine very sparse punctures separated by more than 10 times their own diameters.
|
||
</paragraph>
|
||
<paragraph blockId="2.[828,1438,278,2070]" pageId="2" pageNumber="157">
|
||
Pronotum. Lateral outline with shortly rounded angle in lateral view (as in
|
||
<figureCitation box="[1048,1111,1311,1334]" captionStart="Figs 1–8. 1, 2, 6, 7" captionStartId="3.[151,188,1016,1035]" captionTargetBox="[188,1406,191,997]" captionTargetId="figure-669@3.[158,1431,157,997]" captionTargetPageId="3" captionText="Figs 1–8. 1, 2, 6, 7 – aedeagus laterally and tip of aedeagus dorsally. 3, 5 – lateral view of body. 4, 8 – spermatheca. 1–4 – Agathidium (Agathidium) pilulum sp. nov.; 5–8 – Agathidium (Macroceble) cornigerum sp. nov. Scale bars in Figs 1, 2, 4, 6, 7, 8 = 0.1 mm; 3, 5 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7161304" httpUri="https://zenodo.org/record/7161304/files/figure.png" pageId="2" pageNumber="157">Fig. 3</figureCitation>
|
||
). Puncturation sparse and fine, similar to that on head with punctures separated by about 10 or even more times their own diameter.
|
||
</paragraph>
|
||
<paragraph blockId="2.[828,1438,278,2070]" pageId="2" pageNumber="157">
|
||
Elytra widest at lateral angles, then evenly roundly narrowed posteriorly. Elytral margin straight behind lateral angles in lateral view (
|
||
<figureCitation box="[1076,1142,1471,1494]" captionStart="Figs 1–8. 1, 2, 6, 7" captionStartId="3.[151,188,1016,1035]" captionTargetBox="[188,1406,191,997]" captionTargetId="figure-669@3.[158,1431,157,997]" captionTargetPageId="3" captionText="Figs 1–8. 1, 2, 6, 7 – aedeagus laterally and tip of aedeagus dorsally. 3, 5 – lateral view of body. 4, 8 – spermatheca. 1–4 – Agathidium (Agathidium) pilulum sp. nov.; 5–8 – Agathidium (Macroceble) cornigerum sp. nov. Scale bars in Figs 1, 2, 4, 6, 7, 8 = 0.1 mm; 3, 5 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7161304" httpUri="https://zenodo.org/record/7161304/files/figure.png" pageId="2" pageNumber="157">Fig. 3</figureCitation>
|
||
). Lateral elytral channel as narrow behind lateral angles as in basal part. Puncturation similar to that on head but punctures a little stronger, separated by about 6–10 or more times their own diameter. Sutural striae not developed.
|
||
</paragraph>
|
||
<paragraph blockId="2.[828,1438,278,2070]" pageId="2" pageNumber="157">
|
||
Legs. Tarsi slim, all tibiae slim, metafemora simple, without specific characters. Tarsal formula: 5-5-
|
||
<quantity box="[1332,1373,1663,1686]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="157" unit="in" value="4.0">4 in</quantity>
|
||
male.
|
||
</paragraph>
|
||
<paragraph blockId="2.[828,1438,278,2070]" pageId="2" pageNumber="157">Mesoventrite. Posterior part subconcave. Anterior part raised with longitudinal carina. Lateral lines not developed.</paragraph>
|
||
<paragraph blockId="2.[828,1438,278,2070]" pageId="2" pageNumber="157">Metaventrite moderately developed. Femoral lines developed, shortened for half of their possible length. Fovea located in middle of posterior margin of metaventrite equipped with bush of erect setae.</paragraph>
|
||
<paragraph blockId="2.[828,1438,278,2070]" box="[858,1193,1887,1910]" pageId="2" pageNumber="157">Membranous wings developed.</paragraph>
|
||
<paragraph blockId="2.[828,1438,278,2070]" lastBlockId="3.[150,759,1150,2069]" lastPageId="3" lastPageNumber="158" pageId="2" pageNumber="157">
|
||
Genitalia. Basal part of median lobe narrow ring-shaped of
|
||
<typeStatus box="[855,901,1951,1974]" pageId="2" pageNumber="157">type</typeStatus>
|
||
C. Shape of median lobe apex resembles skittle pin in dorsal view. Apical part of median lobe first contracted, emarginated laterally, then feebly widened. Operculum egg-shaped. Parameres narrowed apically; each with two short and fine setae (aedeagus as in
|
||
<figureCitation box="[532,627,1150,1173]" captionStart="Figs 1–8. 1, 2, 6, 7" captionStartId="3.[151,188,1016,1035]" captionTargetBox="[188,1406,191,997]" captionTargetId="figure-669@3.[158,1431,157,997]" captionTargetPageId="3" captionText="Figs 1–8. 1, 2, 6, 7 – aedeagus laterally and tip of aedeagus dorsally. 3, 5 – lateral view of body. 4, 8 – spermatheca. 1–4 – Agathidium (Agathidium) pilulum sp. nov.; 5–8 – Agathidium (Macroceble) cornigerum sp. nov. Scale bars in Figs 1, 2, 4, 6, 7, 8 = 0.1 mm; 3, 5 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7161304" httpUri="https://zenodo.org/record/7161304/files/figure.png" pageId="3" pageNumber="158">Figs 1, 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7161304" ID-Zenodo-Dep="7161304" httpUri="https://zenodo.org/record/7161304/files/figure.png" pageId="3" pageNumber="158" startId="3.[151,188,1016,1035]" targetBox="[188,1406,191,997]" targetPageId="3">
|
||
<paragraph blockId="3.[151,1437,1016,1061]" pageId="3" pageNumber="158">
|
||
Figs 1–8. 1, 2, 6, 7 – aedeagus laterally and tip of aedeagus dorsally. 3, 5 – lateral view of body. 4, 8 – spermatheca. 1–4 –
|
||
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="pilulum" status="sp. nov." subGenus="Agathidium">
|
||
<emphasis box="[1218,1318,1016,1035]" italics="true" pageId="3" pageNumber="158">Agathidium</emphasis>
|
||
(
|
||
<emphasis box="[1330,1430,1016,1035]" italics="true" pageId="3" pageNumber="158">Agathidium</emphasis>
|
||
)
|
||
<emphasis box="[151,216,1042,1061]" italics="true" pageId="3" pageNumber="158">pilulum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[222,291,1042,1061]" pageId="3" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 5–8 –
|
||
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[352,677,1042,1061]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="cornigerum" status="sp. nov." subGenus="Macroceble">
|
||
<emphasis box="[352,452,1042,1061]" italics="true" pageId="3" pageNumber="158">Agathidium</emphasis>
|
||
(
|
||
<emphasis box="[464,565,1042,1061]" italics="true" pageId="3" pageNumber="158">Macroceble</emphasis>
|
||
)
|
||
<emphasis box="[577,677,1042,1061]" italics="true" pageId="3" pageNumber="158">cornigerum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[682,749,1042,1061]" pageId="3" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Scale bars in Figs 1, 2, 4, 6, 7, 8 = 0.1 mm; 3, 5 = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="3.[150,759,1150,2069]" lastBlockId="3.[828,1437,1150,1557]" pageId="3" pageNumber="158">
|
||
<emphasis bold="true" box="[150,550,1182,1205]" pageId="3" pageNumber="158">Variation and sexual dimorphism.</emphasis>
|
||
Length of body is 1.8–2.0 mm in the
|
||
<typeStatus box="[356,402,1214,1237]" pageId="3" pageNumber="158">type</typeStatus>
|
||
series. The ratio of A3/A2 varies between 1.3–1.6. Tarsal formula 5-4-
|
||
<quantity box="[559,604,1246,1269]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="158" unit="in" value="4.0">4 in</quantity>
|
||
female. Spermatheca with pear-shaped basal part and simply bowed distal thin part (
|
||
<figureCitation box="[319,385,1310,1333]" captionStart="Figs 1–8. 1, 2, 6, 7" captionStartId="3.[151,188,1016,1035]" captionTargetBox="[188,1406,191,997]" captionTargetId="figure-669@3.[158,1431,157,997]" captionTargetPageId="3" captionText="Figs 1–8. 1, 2, 6, 7 – aedeagus laterally and tip of aedeagus dorsally. 3, 5 – lateral view of body. 4, 8 – spermatheca. 1–4 – Agathidium (Agathidium) pilulum sp. nov.; 5–8 – Agathidium (Macroceble) cornigerum sp. nov. Scale bars in Figs 1, 2, 4, 6, 7, 8 = 0.1 mm; 3, 5 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7161304" httpUri="https://zenodo.org/record/7161304/files/figure.png" pageId="3" pageNumber="158">Fig. 4</figureCitation>
|
||
). Length of spermatheca
|
||
<quantity box="[656,754,1310,1333]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="158" unit="mm" value="0.13">0.13 mm</quantity>
|
||
.
|
||
<emphasis bold="true" box="[150,397,1342,1365]" pageId="3" pageNumber="158">Differential diagnosis.</emphasis>
|
||
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[401,759,1342,1365]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="pilulum" status="sp. nov." subGenus="Agathidium">
|
||
<emphasis box="[401,525,1342,1365]" italics="true" pageId="3" pageNumber="158">Agathidium</emphasis>
|
||
(
|
||
<emphasis box="[539,663,1342,1365]" italics="true" pageId="3" pageNumber="158">Agathidium</emphasis>
|
||
)
|
||
<emphasis box="[678,759,1342,1365]" italics="true" pageId="3" pageNumber="158">pilulum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[150,233,1374,1397]" pageId="3" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
as well as the species it is compared with bellow habitually belong to the
|
||
<taxonomicName authorityName="Portevin" authorityYear="1937" box="[402,546,1406,1429]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="madurense">
|
||
<emphasis box="[402,546,1406,1429]" italics="true" pageId="3" pageNumber="158">A. madurense</emphasis>
|
||
</taxonomicName>
|
||
species group sensu AඇGൾඅංඇං (1993) due to raised anterior part of the supraocular carina. The new species stands closest to
|
||
<taxonomicName authority="Angelini & De Marzo, 1998" authorityName="Angelini & De Marzo" authorityYear="1998" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="occultum" subGenus="Agathidium">
|
||
<emphasis box="[642,658,1470,1493]" italics="true" pageId="3" pageNumber="158">A</emphasis>
|
||
. (
|
||
<emphasis italics="true" pageId="3" pageNumber="158">Agathidium</emphasis>
|
||
)
|
||
<emphasis box="[219,317,1502,1525]" italics="true" pageId="3" pageNumber="158">occultum</emphasis>
|
||
Angelini & De Marzo, 1998
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry box="[689,755,1502,1525]" name="China" pageId="3" pageNumber="158">China</collectingCountry>
|
||
,
|
||
<collectingRegion box="[150,231,1534,1557]" country="China" name="Yunnan" pageId="3" pageNumber="158">Yunnan</collectingRegion>
|
||
(AඇGൾඅංඇං & Dൾ Mൺඋඓඈ 1998b) as both species are similar in the shape of body that is broadly oval, in the shape of straight, feebly emarginated clypeus, in the eyes that are parabola slice-shaped and in the raised anterolateral bead of head. Colour of the antenna is yellow-red with A9 and A10 a little darker in both species. Both compared species possess A3 distinctly longer than A2 and also slim tibiae. Sutural stria is missing in both species.
|
||
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="pilulum" status="sp. nov.">
|
||
<emphasis italics="true" pageId="3" pageNumber="158">Agathidium pilulum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[240,325,1790,1813]" pageId="3" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName authorityName="Angelini & De Marzo" authorityYear="1998" box="[469,599,1790,1813]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="occultum">
|
||
<emphasis box="[469,599,1790,1813]" italics="true" pageId="3" pageNumber="158">A. occultum</emphasis>
|
||
</taxonomicName>
|
||
in the specific shape of its median lobe that is simply cylindrical with broadly rounded tip in
|
||
<taxonomicName authorityName="Angelini & De Marzo" authorityYear="1998" box="[399,531,1854,1877]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="occultum">
|
||
A.
|
||
<emphasis box="[433,531,1854,1877]" italics="true" pageId="3" pageNumber="158">occultum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[545,758,1854,1877]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="pilulum" status="sp. nov.">
|
||
<emphasis box="[545,758,1854,1877]" italics="true" pageId="3" pageNumber="158">Agathidium pilulum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[150,234,1886,1909]" pageId="3" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be also compared to
|
||
<taxonomicName authority="Angelini & De Marzo, 1998" authorityName="Angelini & De Marzo" authorityYear="1998" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="alesi" subGenus="Agathidium">
|
||
<emphasis box="[505,528,1886,1909]" italics="true" pageId="3" pageNumber="158">A.</emphasis>
|
||
(
|
||
<emphasis box="[543,566,1886,1909]" italics="true" pageId="3" pageNumber="158">A.</emphasis>
|
||
)
|
||
<emphasis box="[581,631,1886,1909]" italics="true" pageId="3" pageNumber="158">alesi</emphasis>
|
||
Angelini & De Marzo, 1998
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry box="[399,477,1918,1941]" name="Taiwan" pageId="3" pageNumber="158">Taiwan</collectingCountry>
|
||
(AඇGൾඅංඇං & Dൾ Mൺඋඓඈ 1998a), that also belongs to the
|
||
<taxonomicName authorityName="Portevin" authorityYear="1937" box="[518,670,1950,1973]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="madurense">
|
||
<emphasis box="[518,670,1950,1973]" italics="true" pageId="3" pageNumber="158">A. madurense</emphasis>
|
||
</taxonomicName>
|
||
species group, having similar skittle pin-shaped apical part of the median lobe. The new species differs from
|
||
<taxonomicName authorityName="Angelini & De Marzo" authorityYear="1998" box="[651,731,2014,2037]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="alesi">
|
||
<emphasis box="[651,731,2014,2037]" italics="true" pageId="3" pageNumber="158">A. alesi</emphasis>
|
||
</taxonomicName>
|
||
in distinctly smaller body (1.8–2.0 mm), while the length of body is 3.0–
|
||
<quantity box="[958,1037,1150,1173]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="158" unit="mm" value="4.1">4.1 mm</quantity>
|
||
in
|
||
<emphasis box="[1070,1157,1150,1173]" italics="true" pageId="3" pageNumber="158">
|
||
<taxonomicName authorityName="Angelini & De Marzo" authorityYear="1998" box="[1070,1153,1150,1173]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="alesi">A. alesi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Antenna is bicolored in
|
||
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="pilulum" status="sp. nov.">
|
||
<emphasis italics="true" pageId="3" pageNumber="158">A. pilulum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[916,1001,1182,1205]" pageId="3" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, antenna in
|
||
<taxonomicName authorityName="Angelini & De Marzo" authorityYear="1998" box="[1129,1209,1182,1205]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="alesi">
|
||
<emphasis box="[1129,1209,1182,1205]" italics="true" pageId="3" pageNumber="158">A. alesi</emphasis>
|
||
</taxonomicName>
|
||
is unicolorous, light. Posterior femora are straight on their hind margin, while the same are toothed in
|
||
<taxonomicName authorityName="Angelini & De Marzo" authorityYear="1998" box="[1075,1154,1246,1269]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="alesi">
|
||
<emphasis box="[1075,1154,1246,1269]" italics="true" pageId="3" pageNumber="158">A. alesi</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, the apex of the tegmen is distinctly bent upwards and paramera is shorter in
|
||
<taxonomicName authorityName="Angelini & De Marzo" authorityYear="1998" box="[853,931,1310,1333]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="alesi">
|
||
<emphasis box="[853,931,1310,1333]" italics="true" pageId="3" pageNumber="158">A. alesi</emphasis>
|
||
</taxonomicName>
|
||
compared with
|
||
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[1099,1208,1310,1333]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="pilulum" status="sp. nov.">
|
||
<emphasis box="[1099,1208,1310,1333]" italics="true" pageId="3" pageNumber="158">A. pilulum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1213,1294,1310,1333]" pageId="3" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Spermatheca is similar in its shape in all the compared species; basal part of spermatheca is pear-shaped in
|
||
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[1235,1346,1374,1397]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="pilulum" status="sp. nov.">
|
||
<emphasis box="[1235,1346,1374,1397]" italics="true" pageId="3" pageNumber="158">A. pilulum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1354,1437,1374,1397]" pageId="3" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
while the same is subcylindrical in
|
||
<taxonomicName authorityName="Angelini & De Marzo" authorityYear="1998" box="[1184,1307,1406,1429]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="occultum">
|
||
<emphasis box="[1184,1307,1406,1429]" italics="true" pageId="3" pageNumber="158">A. occultum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis box="[1353,1437,1406,1429]" italics="true" pageId="3" pageNumber="158">
|
||
<taxonomicName authorityName="Angelini & De Marzo" authorityYear="1998" box="[1353,1433,1406,1429]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="alesi">A. alesi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<emphasis bold="true" box="[828,956,1438,1461]" pageId="3" pageNumber="158">Etymology.</emphasis>
|
||
The name of the new species draws the attention to the globular shape of the apex of the median lobe. Therefore the name is derived from the Latin word
|
||
<emphasis box="[1374,1436,1502,1525]" italics="true" pageId="3" pageNumber="158">pilula</emphasis>
|
||
(= globule in English). Noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |