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<document ID-DOI="http://dx.doi.org/10.3897/dez.66.31754" ID-GBIF-Dataset="22c3ecd6-0d0a-443a-ba47-023ac8504bcc" ID-GBIF-Taxon="182223586" ID-Pensoft-Pub="1860-1324-1-1" ID-Pensoft-UUID="8A92B765B02355A2A9D1168A93C881F2" ID-Zenodo-Dep="2551893" ID-ZooBank="11B6BFA2262745108C7C5D38945AA4F1" LSID="urn:lsid:zoobank.org:act:7C58EE0D-541B-4C5A-9937-CCDE8AAF01C5" ModsDocID="1860-1324-66-1-1" checkinTime="1621372408696" checkinUser="pensoft" docAuthor="Lohaj, Roman &amp; Delic, Teo" docDate="2019" docId="327918A99275A12B6CD2518E41C7612C" docLanguage="en" docName="DeutEntomolZeit 66(1): 1-15" docOrigin="Deutsche Entomologische Zeitschrift 66 (1)" docPubDate="2019-01-21" docSource="http://dx.doi.org/10.3897/dez.66.31754" docTitle="Adriaphaenops petrimaris Lohaj &amp; Delić 2019, sp. n." docType="treatment" docUuid="7C58EE0D-541B-4C5A-9937-CCDE8AAF01C5" docUuidSource="ZooBank" docVersion="3" id="8A92B765B02355A2A9D1168A93C881F2" lastPageNumber="1" masterDocId="8A92B765B02355A2A9D1168A93C881F2" masterDocTitle="Playing hard to get: two new species of subterranean Trechini beetles (Coleoptera, Carabidae, Trechinae) from the Dinaric Karst" masterLastPageNumber="15" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1643457464602" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Playing hard to get: two new species of subterranean Trechini beetles (Coleoptera, Carabidae, Trechinae) from the Dinaric Karst</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lohaj, Roman</mods:namePart>
<mods:affiliation>Zahradna 61 A, SK- 900 91 Limbach, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Delic, Teo</mods:namePart>
<mods:affiliation>SubBioLab, Department of Biology, Biotechnical Faculty, University of Ljubljana, Jamnikarjeva 101, SI- 1000 Ljubljana, Slovenia</mods:affiliation>
<mods:nameIdentifier type="email">tejc86@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-01-21</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>66</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>15</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.66.31754</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.66.31754</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-1-1</mods:identifier>
<mods:identifier type="ZooBank">11B6BFA2262745108C7C5D38945AA4F1</mods:identifier>
<mods:identifier type="Pensoft-UUID">8A92B765B02355A2A9D1168A93C881F2</mods:identifier>
<mods:identifier type="Zenodo-Dep">2551893</mods:identifier>
</mods:mods>
<subSection pageId="0" pageNumber="1" type="results">
<treatment ID-GBIF-Taxon="182223586" LSID="urn:lsid:zoobank.org:act:7C58EE0D-541B-4C5A-9937-CCDE8AAF01C5" httpUri="http://treatment.plazi.org/id/327918A99275A12B6CD2518E41C7612C" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/7C58EE0D-541B-4C5A-9937-CCDE8AAF01C5" authority="Lohaj &amp; Delić, 2019" authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops petrimaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="petrimaris" status="sp. n.">Adriaphaenops petrimaris</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geographical distribution of the Dinaric aphaenopsoid trechines of the genera Derossiella and Adriaphenops. Source of data: http: // subbio. net / db /" figureDoi="10.3897/dez.66.31754.figure2" httpUri="https://binary.pensoft.net/fig/258009" pageId="0" pageNumber="1">Figs 2</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Adriaphaenops petrimaris sp. n. in its natural habitat in Pistet 4 cave, Velji Pistet, Kameno more, Risan." figureDoi="10.3897/dez.66.31754.figure6" httpUri="https://binary.pensoft.net/fig/258013" pageId="0" pageNumber="1">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of morphologically similar Adriaphaenops species. Adriaphaenops petrimaris sp. n. (a), Adriaphaenops rumijaensis (b) and Adriaphenops kevser (c). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure7" httpUri="https://binary.pensoft.net/fig/258014" pageId="0" pageNumber="1">, 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male genitalia of Adriaphaenops petrimaris sp. n. (a) and similarly looking A. kevser (b) (illustrations by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure8" httpUri="https://binary.pensoft.net/fig/258015" pageId="0" pageNumber="1">, 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia; gonocoxite 1 and 2 (basal and apical segments of gonostylus) of Adriaphaenops petrimaris sp. n. (a), A. rumijaensis (b), and A. kevser (c) (illustrations by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure9" httpUri="https://binary.pensoft.net/fig/258016" pageId="0" pageNumber="1">, 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type series">
<paragraph pageId="0" pageNumber="1">Type series.</paragraph>
<paragraph pageId="0" pageNumber="1">
Holotype male labelled: &quot;MONTENEGRO, Risan, Velji
<normalizedToken originalValue="Pištet">Pistet</normalizedToken>
, Kameno more, cave
<normalizedToken originalValue="Pištet">Pistet</normalizedToken>
4 (PT4), (
<geoCoordinate degrees="42.55183" direction="north" orientation="latitude" precision="1" value="42.55183">42.55183°N</geoCoordinate>
,
<geoCoordinate degrees="18.73864" direction="east" orientation="longitude" precision="1" value="18.73864">18.73864°E</geoCoordinate>
), - 40 m, 3.5.2018, T.
<normalizedToken originalValue="Delić">Delic</normalizedToken>
lgt.&quot; (white label, printed) / &quot;DNA extraction XA475&quot; (white label, printed) / &quot;HOLOTYPUS
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops petrimaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="petrimaris">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops petrimaris</emphasis>
</taxonomicName>
sp. n. Lohaj &amp;
<normalizedToken originalValue="Delić">Delic</normalizedToken>
des. 2018&quot; (red label, printed) (PMSL, voucher code
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
-11519). Paratype one female (left antenna missing), labelled: MONTENEGRO, Risan, Velji
<normalizedToken originalValue="Pištet">Pistet</normalizedToken>
, Kameno more, cave
<normalizedToken originalValue="Pištet">Pistet</normalizedToken>
4 (PT4), (
<geoCoordinate degrees="42.55183" direction="north" orientation="latitude" precision="1" value="42.55183">42.55183°N</geoCoordinate>
,
<geoCoordinate degrees="18.73864" direction="east" orientation="longitude" precision="1" value="18.73864">18.73864°E</geoCoordinate>
), - 100 m, 2.5.2018, T.
<normalizedToken originalValue="Delić">Delic</normalizedToken>
lgt.&quot; (white label, printed) / &quot;PARATYPUS
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops petrimaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="petrimaris">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops petrimaris</emphasis>
</taxonomicName>
sp. n. Lohaj &amp;
<normalizedToken originalValue="Delić">Delic</normalizedToken>
des. 2018&quot; (red label, printed) (CRL).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A medium-sized trechine beetle with aphaenopsoid features: head and pronotum elongate; elytra ovoid, strongly narrowed at the base, obviously wider than head and pronotum; body depigmented, strongly flattened, covered with sparse pubescence (Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Adriaphaenops petrimaris sp. n. in its natural habitat in Pistet 4 cave, Velji Pistet, Kameno more, Risan." figureDoi="10.3897/dez.66.31754.figure6" httpUri="https://binary.pensoft.net/fig/258013" pageId="0" pageNumber="1">6</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of morphologically similar Adriaphaenops species. Adriaphaenops petrimaris sp. n. (a), Adriaphaenops rumijaensis (b) and Adriaphenops kevser (c). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure7" httpUri="https://binary.pensoft.net/fig/258014" pageId="0" pageNumber="1">7</figureCitation>
). This new species is mainly characterized by the presence of four pairs of dorsal setae (three discal and one preapical) on elytra. This character is so far present only in presumably closely related species,
<taxonomicName genus="A." lsidName="A. kevser" pageId="0" pageNumber="1" rank="species" species="kevser">
<emphasis italics="true" pageId="0" pageNumber="1">A. kevser</emphasis>
</taxonomicName>
and
<taxonomicName genus="A." lsidName="A. rumijaensis" pageId="0" pageNumber="1" rank="species" species="rumijaensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. rumijaensis</emphasis>
</taxonomicName>
; see identification key below.
</paragraph>
<caption ID-Zenodo-Dep="2551943" doi="10.3897/dez.66.31754.figure6" httpUri="https://binary.pensoft.net/fig/258013" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops petrimaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="petrimaris">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops petrimaris</emphasis>
</taxonomicName>
sp. n. in its natural habitat in
<normalizedToken originalValue="Pištet">Pistet</normalizedToken>
4 cave, Velji
<normalizedToken originalValue="Pištet">Pistet</normalizedToken>
, Kameno more, Risan.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="2551945" doi="10.3897/dez.66.31754.figure7" httpUri="https://binary.pensoft.net/fig/258014" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
Habitus of morphologically similar
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
species.
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops petrimaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="petrimaris">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops petrimaris</emphasis>
</taxonomicName>
sp. n.
<emphasis bold="true" pageId="0" pageNumber="1">(a)</emphasis>
,
<taxonomicName class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops rumijaensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rumijaensis">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops rumijaensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">(b)</emphasis>
and
<taxonomicName genus="Adriaphenops" lsidName="Adriaphenops kevser" pageId="0" pageNumber="1" rank="species" species="kevser">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphenops kevser</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">(c)</emphasis>
. Chetotaxy is presented as white points (photo courtesy of
<normalizedToken originalValue="Dušan">Dusan</normalizedToken>
<normalizedToken originalValue="Beňo">Beno</normalizedToken>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">L: 5.0 mm (HT)-5.4 mm (PT), TL: 4.7 mm (HT)-5.1 mm (PT). Head relatively large, rounded, with maximum width behind middle, distinctly longer than wide (index HL/HW 1.23 (HT)-1.33 (PT), slightly wider than pronotum, sparsely pubescent. Frontal furrows weakly impressed, short, ending in the front half of head. Two pairs of long supraorbital setae present; neck constriction distinct; genae gently convex. Clypeus and labrum with three pairs of setae, outer pairs longer. Antennae length 3.3 mm (HT)-3.4 mm (PT), scape as long as pedicel, almost as long as terminal antennomere.</paragraph>
<paragraph pageId="0" pageNumber="1">Pronotum elongate, slightly longer and wider than head, with maximum width in middle, only very slightly narrowed anteriorly, basal part distinctly narrower than anterior (index PL/PW 1.66 (HT)-1.57 (PT)), sparsely pubescent, setae long, suberect; median furrow weakly marked, visible in the middle part of pronotum. Propleura visible from dorsal aspect only in basal half. Anterior angles of pronotum distinctly protruding, obtuse, posterior angles obtuse. Lateral furrows developed, deep, with one pair of anterolateral setae, situated in apical fourth of pronotal length.</paragraph>
<paragraph pageId="0" pageNumber="1">Elytra subovate elongate, distinctly longer than wide (index EL/EW 1.74 (HT)-1.78 (PT)), with maximum width in middle; elytral surface covered with very sparse, long and erect pubescence; striae absent. Site of stria 3 with four (three discal and one preapical) setae, humeral group of umbilicate pores not aggregated, first anterior pore of humeral group isolated and situated before the level of the first discal seta. Pore 5 located nearer to pore 6 than to pore 4.</paragraph>
<paragraph pageId="0" pageNumber="1">Legs long, slender, densely pubescent. First two tarsomeres of male protarsi distinctly dilated and protracted at their internal margins. Tarsal claws very long and slender, without traces of denticulation on their internal sides.</paragraph>
<paragraph pageId="0" pageNumber="1">
Male genitalia (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male genitalia of Adriaphaenops petrimaris sp. n. (a) and similarly looking A. kevser (b) (illustrations by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure8" httpUri="https://binary.pensoft.net/fig/258015" pageId="0" pageNumber="1">8</figureCitation>
): aedeagus very long and slender, gradually narrowed towards apex in lateral aspect, sagittal aileron large, endophallus without distinct copulatory piece. Parameres long and slender, with two long setae at apex. Female genitalia as in Figure
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia; gonocoxite 1 and 2 (basal and apical segments of gonostylus) of Adriaphaenops petrimaris sp. n. (a), A. rumijaensis (b), and A. kevser (c) (illustrations by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure9" httpUri="https://binary.pensoft.net/fig/258016" pageId="0" pageNumber="1">9</figureCitation>
.
</paragraph>
<caption ID-Zenodo-Dep="2551947" doi="10.3897/dez.66.31754.figure8" httpUri="https://binary.pensoft.net/fig/258015" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
Male genitalia of
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops petrimaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="petrimaris">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops petrimaris</emphasis>
</taxonomicName>
sp. n.
<emphasis bold="true" pageId="0" pageNumber="1">(a)</emphasis>
and similarly looking
<taxonomicName genus="A." lsidName="A. kevser" pageId="0" pageNumber="1" rank="species" species="kevser">
<emphasis italics="true" pageId="0" pageNumber="1">A. kevser</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">(b)</emphasis>
(illustrations by Fedor
<normalizedToken originalValue="Čiampor">Ciampor</normalizedToken>
).
</paragraph>
</caption>
<caption ID-Zenodo-Dep="2551949" doi="10.3897/dez.66.31754.figure9" httpUri="https://binary.pensoft.net/fig/258016" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
Female genitalia; gonocoxite 1 and 2 (basal and apical segments of gonostylus) of
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops petrimaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="petrimaris">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops petrimaris</emphasis>
</taxonomicName>
sp. n.
<emphasis bold="true" pageId="0" pageNumber="1">(a)</emphasis>
,
<taxonomicName genus="A." lsidName="A. rumijaensis" pageId="0" pageNumber="1" rank="species" species="rumijaensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. rumijaensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">(b)</emphasis>
, and
<taxonomicName genus="A." lsidName="A. kevser" pageId="0" pageNumber="1" rank="species" species="kevser">
<emphasis italics="true" pageId="0" pageNumber="1">A. kevser</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">(c)</emphasis>
(illustrations by Fedor
<normalizedToken originalValue="Čiampor">Ciampor</normalizedToken>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Topotypic, referring to the toponym where the
<normalizedToken originalValue="Pišet">Piset</normalizedToken>
4 cave is situated, Kameno more (in English, Sea of stone and in Latin, Mare petris).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Genus
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
currently comprises 13 described species, including
<taxonomicName genus="A." lsidName="A. petrimaris" pageId="0" pageNumber="1" rank="species" species="petrimaris">
<emphasis italics="true" pageId="0" pageNumber="1">A. petrimaris</emphasis>
</taxonomicName>
sp. n. This species,
<taxonomicName genus="A." lsidName="A. kevser" pageId="0" pageNumber="1" rank="species" species="kevser">
<emphasis italics="true" pageId="0" pageNumber="1">A. kevser</emphasis>
</taxonomicName>
, and
<taxonomicName genus="A." lsidName="A. rumijaensis" pageId="0" pageNumber="1" rank="species" species="rumijaensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. rumijaensis</emphasis>
</taxonomicName>
form a group of species with four discal setae (three dorsal and one preapical) on elytra, with elytral pubescence in all three species sparser in comparison to the other species (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of morphologically similar Adriaphaenops species. Adriaphaenops petrimaris sp. n. (a), Adriaphaenops rumijaensis (b) and Adriaphenops kevser (c). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure7" httpUri="https://binary.pensoft.net/fig/258014" pageId="0" pageNumber="1">7</figureCitation>
).
<taxonomicName genus="A." lsidName="A. petrimaris" pageId="0" pageNumber="1" rank="species" species="petrimaris">
<emphasis italics="true" pageId="0" pageNumber="1">A. petrimaris</emphasis>
</taxonomicName>
morphologically resembles
<taxonomicName genus="A." lsidName="A. kevser" pageId="0" pageNumber="1" rank="species" species="kevser">
<emphasis italics="true" pageId="0" pageNumber="1">A. kevser</emphasis>
</taxonomicName>
, described from Mt
<normalizedToken originalValue="Lebršnik">Lebrsnik</normalizedToken>
, Bosnia &amp; Hercegovina. These two species can be easily recognized by the shape of head, pronotum, and aedeagus (Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of morphologically similar Adriaphaenops species. Adriaphaenops petrimaris sp. n. (a), Adriaphaenops rumijaensis (b) and Adriaphenops kevser (c). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure7" httpUri="https://binary.pensoft.net/fig/258014" pageId="0" pageNumber="1">7</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male genitalia of Adriaphaenops petrimaris sp. n. (a) and similarly looking A. kevser (b) (illustrations by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure8" httpUri="https://binary.pensoft.net/fig/258015" pageId="0" pageNumber="1">8</figureCitation>
) (see identification key below).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
So far this species is known only from the type locality,
<normalizedToken originalValue="Pištet">Pistet</normalizedToken>
4 cave (synonym = PT4), Kameno more, Risan, Montenegro. Both specimens, HT and PT, were found walking on the wet and damp vertical cave walls at the depth of 40 and 100 m.
</paragraph>
<paragraph pageId="0" pageNumber="1">Associated subterranean fauna observed in the pit:</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="? arthropoda" rank="family">Carabidae</taxonomicName>
:
<taxonomicName authorityName="Bonelli" authorityYear="1810" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Trechinae">Trechinae</taxonomicName>
:
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="L. Schaufuss" baseAuthorityYear="1864" class="Insecta" family="Carabidae" genus="Neotrechus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Neotrechus suturalis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis italics="true" pageId="0" pageNumber="1">Neotrechus suturalis</emphasis>
</taxonomicName>
ssp. (Schaufuss, 1864)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Ganglbauer" baseAuthorityYear="1896" class="Insecta" family="Carabidae" genus="Neotrechus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Neotrechus paganettii" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="paganettii">
<emphasis italics="true" pageId="0" pageNumber="1">Neotrechus paganettii</emphasis>
</taxonomicName>
ssp. (Ganglbauer, 1896)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fleming" authorityYear="1821" class="Insecta" family="Leiodidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
:
<taxonomicName authorityName="Kirby" authorityYear="1837" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Cholevinae">Cholevinae</taxonomicName>
:
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Reitter" authorityYear="1910" class="Insecta" family="Leiodidae" genus="Blattochaeta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Blattochaeta" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Blattochaeta</emphasis>
</taxonomicName>
sp.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Reitter" authorityYear="1889" class="Insecta" family="Leiodidae" genus="Anthroherpon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Anthroherpon" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Anthroherpon</emphasis>
</taxonomicName>
sp.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Zariquiey" authorityYear="1927" class="Insecta" family="Leiodidae" genus="Hadesia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadesia weiratheri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="weiratheri">Hadesia cf. weiratheri</taxonomicName>
Zariquiey, 1927
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="identification key to the species of the genus adriaphaenops noesske (for the drawings and details, see lohaj et al. 2016)">
<paragraph pageId="0" pageNumber="1">
Identification key to the species of the genus
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
Noesske (for the drawings and details, see
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4205.6.1" author="Lohaj, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="501 - 531" refId="B32" refString="Lohaj, R, Lakota, J, Queinnec, E, Pavicevic, D, Ceplik, D, 2016. Study on Adriaphaenops Noesske with the description of five new species from the Dinarides (Coleoptera: Carabidae: Trechini). Zootaxa 4205 (6): 501 - 531, DOI: https://doi.org/10.11646/zootaxa.4205.6.1" title="Study on Adriaphaenops Noesske with the description of five new species from the Dinarides (Coleoptera: Carabidae: Trechini)." url="https://doi.org/10.11646/zootaxa.4205.6.1" volume="4205" year="2016">Lohaj et al. 2016</bibRefCitation>
)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<table inLine="true" pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1(6)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Head almost rounded, wider</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2(5)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Head with 2 pairs of supraorbital setae</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3(4)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Clypeus with 4 pairs of setae, pronotum wider, index PL/PW 1.21, base of pronotum as wide as anterior part L: 4.65-5 mm. BiH, Hercegovina, Popovo polje</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
9.
<taxonomicName genus="A." lsidName="A. pretneri" pageId="0" pageNumber="1" rank="species" species="pretneri">
<emphasis italics="true" pageId="0" pageNumber="1">A. pretneri</emphasis>
</taxonomicName>
Scheibel, 1935
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4(3)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Clypeus with 3 pairs of setae, pronotum narrower, index PL/PW 1.42, base of pronotum narrower than anterior part. L: 5.5 mm. BiH, Hercegovina, Nevesinje</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
3.
<taxonomicName genus="A." lsidName="A. jasminkoi" pageId="0" pageNumber="1" rank="species" species="jasminkoi">
<emphasis italics="true" pageId="0" pageNumber="1">A. jasminkoi</emphasis>
</taxonomicName>
Lohaj et al., 2016
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5(2)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
Head without supraorbital setae or these setae are indistinguishable from head pubescence L: 4.6 mm. BiH, Hercegovina, Turica, Mt
<normalizedToken originalValue="Bjelašnica">Bjelasnica</normalizedToken>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
7.
<taxonomicName genus="A." lsidName="A. perreaui" pageId="0" pageNumber="1" rank="species" species="perreaui">
<emphasis italics="true" pageId="0" pageNumber="1">A. perreaui</emphasis>
</taxonomicName>
<normalizedToken originalValue="Quéinnec">Queinnec</normalizedToken>
&amp;
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
, 2008
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6(1)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Head elongate or parallel-sided, distinctly narrower</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7(12)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Elytra with 4 pairs of discal setae (3 dorsal and 1 preapical), only very sparsely pubescent</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8(11)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Pronotum widest in anterior third/middle. Ultimate segment of maxillar palpi distinctly shorter than penultimate</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">9</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">9(10)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
Pronotum widest in anterior third, distinctly narrowed anteriorly, head narrower. Aedeagus wider in lateral view, apex widely rounded (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male genitalia of Adriaphaenops petrimaris sp. n. (a) and similarly looking A. kevser (b) (illustrations by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure8" httpUri="https://binary.pensoft.net/fig/258015" pageId="0" pageNumber="1">8</figureCitation>
) L: 4.8-5.7 mm. BiH, Hercegovina, Mt
<normalizedToken originalValue="Lebršnik">Lebrsnik</normalizedToken>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
4.
<taxonomicName genus="A." lsidName="A. kevser" pageId="0" pageNumber="1" rank="species" species="kevser">
<emphasis italics="true" pageId="0" pageNumber="1">A. kevser</emphasis>
</taxonomicName>
<normalizedToken originalValue="Quéinnec">Queinnec</normalizedToken>
,
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
&amp; Ollivier, 2008
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">10(9)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
Pronotum widest in middle, only very slightly narrowed anteriorly, head more rounded. Aedeagus narrower in lateral view, apex pointed (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of morphologically similar Adriaphaenops species. Adriaphaenops petrimaris sp. n. (a), Adriaphaenops rumijaensis (b) and Adriaphenops kevser (c). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure7" httpUri="https://binary.pensoft.net/fig/258014" pageId="0" pageNumber="1">7</figureCitation>
). L: 5.0-5.4 mm. Montenegro, Risan, Kameno more
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
8.
<taxonomicName genus="A." lsidName="A. petrimaris" pageId="0" pageNumber="1" rank="species" species="petrimaris">
<emphasis italics="true" pageId="0" pageNumber="1">A. petrimaris</emphasis>
</taxonomicName>
, sp. n.
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">11(8)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Pronotum widest in anterior fourth. Ultimate segment of maxillar palpi as long as penultimate. L: 5.05-5.4 mm. Montenegro, Mt Rumija</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
10.
<taxonomicName genus="A." lsidName="A. rumijaensis" pageId="0" pageNumber="1" rank="species" species="rumijaensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. rumijaensis</emphasis>
</taxonomicName>
Lohaj et al., 2016
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">12(7)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Elytra with 3 pairs of discal setae (two dorsal and one preapical), densely pubescent</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">13</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">13(16)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Head without supraorbital setae or these setae are very short, indistinguishable from head pubescence</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">14</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">14(15)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Clypeus with 4 pairs of setae. Smaller species, L: 3.8 mm. Montenegro, Virpazar, Trnovo</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
11.
<taxonomicName genus="A." lsidName="A. staudacheri" pageId="0" pageNumber="1" rank="species" species="staudacheri">
<emphasis italics="true" pageId="0" pageNumber="1">A. staudacheri</emphasis>
</taxonomicName>
Scheibel, 1939
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15(14)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Clypeus with three pairs of setae. Larger species L: 4.9 mm. Montenegro, Cetinje</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
6.
<taxonomicName genus="A." lsidName="A. njegosiensis" pageId="0" pageNumber="1" rank="species" species="njegosiensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. njegosiensis</emphasis>
</taxonomicName>
Lohaj et al., 2016
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">16(13)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Head with one or two pairs of long supraorbital setae</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">17</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">17(20)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Head with only posterior pair of supraorbital setae, anterior pair absent</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">18</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">18(19)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
Clypeus with four pairs of setae, frontal furrows longer, exceeding half length of head, anterior angles of pronotum rounded. L: 4.7-5.75 mm. Montenegro,
<normalizedToken originalValue="Kucˇke">Kucˇke</normalizedToken>
planine Mts
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
5.
<taxonomicName genus="A." lsidName="A. mlejneki" pageId="0" pageNumber="1" rank="species" species="mlejneki">
<emphasis italics="true" pageId="0" pageNumber="1">A. mlejneki</emphasis>
</taxonomicName>
Lohaj et al., 2016
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">19(18)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
Clypeus with three pairs of setae, frontal furrows distinctly shorter than half length of head, anterior angles of pronotum pointed. L: 4.65 mm. Albania,
<normalizedToken originalValue="Shkodër">Shkoder</normalizedToken>
, Boga
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
1.
<taxonomicName genus="A." lsidName="A. albanicus" pageId="0" pageNumber="1" rank="species" species="albanicus">
<emphasis italics="true" pageId="0" pageNumber="1">A. albanicus</emphasis>
</taxonomicName>
Lohaj et al., 2016
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">20(17)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Head with both anterior and posterior pairs of supraorbital setae</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">21</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">21(22)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Clypeus with 3 pairs of setae, head wider, index HL/HW 1.10-1.15, posterior angles of pronotum protruding, acute. L: 3.5-5.1 mm. Montenegro, Mt Durmitor</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
13.
<taxonomicName genus="A." lsidName="A. zupcense" pageId="0" pageNumber="1" rank="species" species="zupcense">
<emphasis italics="true" pageId="0" pageNumber="1">A. zupcense</emphasis>
</taxonomicName>
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
, 1990
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">22(21)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Clypeus with 4 pairs of setae, head narrower, index HL/HW 1.33-1.36, posterior angles of pronotum not protruding, obtuse</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">23</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">23(24)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
Head parallel-sided, pronotum narrower, index PL/PW 1.5, with maximum width in anterior fourth. L:4.7-4.85 mm. BiH, Hercegovina, Gacko, Mt
<normalizedToken originalValue="Bjelašnica">Bjelasnica</normalizedToken>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
2.
<taxonomicName genus="A." lsidName="A. antroherponomimus" pageId="0" pageNumber="1" rank="species" species="antroherponomimus">
<emphasis italics="true" pageId="0" pageNumber="1">A. antroherponomimus</emphasis>
</taxonomicName>
(Noesske, 1928)
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">24(23)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
Head slightly rounded, pronotum wider, index PL/PW 1.35, with maximum width in anterior third. L: 5-5.35 mm. Montenegro,
<normalizedToken originalValue="Nikšic´">Niksic'</normalizedToken>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
12.
<taxonomicName genus="A." lsidName="A. stirni" pageId="0" pageNumber="1" rank="species" species="stirni">
<emphasis italics="true" pageId="0" pageNumber="1">A. stirni</emphasis>
</taxonomicName>
(Pretner, 1959)
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="annotated catalogue of the genus adriaphaenops noesske">
<paragraph pageId="0" pageNumber="1">
Annotated catalogue of the genus
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
Noesske
</paragraph>
<paragraph pageId="0" pageNumber="1">
1
<taxonomicName class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops albanicus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="albanicus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Adriaphaenops albanicus</emphasis>
</taxonomicName>
Lohaj, Lakota,
<normalizedToken originalValue="Quéinnec">Queinnec</normalizedToken>
,
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
&amp;
<normalizedToken originalValue="Čeplík">Ceplik</normalizedToken>
, 2016: 518 (
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
). Type locality: Albania, District
<normalizedToken originalValue="Shkodër">Shkoder</normalizedToken>
, V. Boga, Mts
<normalizedToken originalValue="Thatë">Thate</normalizedToken>
, Grotte No. 25.
</paragraph>
<paragraph pageId="0" pageNumber="1">Distribution: Albania, Prokletije Mts.</paragraph>
<paragraph pageId="0" pageNumber="1">
2
<taxonomicName baseAuthorityName="Noesske" baseAuthorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops antroherponomimus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="antroherponomimus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Adriaphaenops antroherponomimus</emphasis>
</taxonomicName>
Noesske, 1928: 7 (
<taxonomicName class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Trechus</emphasis>
</taxonomicName>
). Type locality:
<normalizedToken originalValue="Čatol">Catol</normalizedToken>
jama des Bjelasica-Gebirges (weitere Umgebung von Gacko) im nordostherzegowinischen Karstlande [=
<normalizedToken originalValue="Sniježnica">Snijeznica</normalizedToken>
,
<normalizedToken originalValue="Tišov">Tisov</normalizedToken>
<normalizedToken originalValue="krš">krs</normalizedToken>
]). Distribution: Bosnia and Hercegovina, Mt
<normalizedToken originalValue="Bjelašnica">Bjelasnica</normalizedToken>
near Gacko.
</paragraph>
<paragraph pageId="0" pageNumber="1">
3
<taxonomicName class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops jasminkoi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="jasminkoi">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Adriaphaenops jasminkoi</emphasis>
</taxonomicName>
Lohaj, Lakota,
<normalizedToken originalValue="Quéinnec">Queinnec</normalizedToken>
,
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
&amp;
<normalizedToken originalValue="Čeplík">Ceplik</normalizedToken>
, 2016: 520 (
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
). Type locality: Bosnia and Hercegovina, Nevesinje,
<normalizedToken originalValue="Bišina">Bisina</normalizedToken>
village,
<normalizedToken originalValue="Novakuša">Novakusa</normalizedToken>
(Novakova)
<normalizedToken originalValue="pećina">pecina</normalizedToken>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">Distribution: Bosnia and Hercegovina, Nevesinje.</paragraph>
<paragraph pageId="0" pageNumber="1">
4
<taxonomicName genus="Adriaphenops" lsidName="Adriaphenops kevser" pageId="0" pageNumber="1" rank="species" species="kevser">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Adriaphenops kevser</emphasis>
</taxonomicName>
<normalizedToken originalValue="Quéinnec">Queinnec</normalizedToken>
,
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
&amp; Ollivier, 2008: 154 (
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
). Type locality: Vilina
<normalizedToken originalValue="pećina">pecina</normalizedToken>
, alt. 1840 m a.s.l.,
<normalizedToken originalValue="Lebršnik">Lebrsnik</normalizedToken>
planina, eastern Hercegovina, Bosnia and Hercegovina)
</paragraph>
<paragraph pageId="0" pageNumber="1">
Distribution: Bosnia &amp; Hercegovina, Mt
<normalizedToken originalValue="Lebršnik">Lebrsnik</normalizedToken>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
5
<taxonomicName class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops mlejneki" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mlejneki">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Adriaphaenops mlejneki</emphasis>
</taxonomicName>
Lohaj, Lakota,
<normalizedToken originalValue="Quéinnec">Queinnec</normalizedToken>
,
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
&amp;
<normalizedToken originalValue="Čeplík">Ceplik</normalizedToken>
, 2016: 524 (
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
). Type locality: Montenegro,
<normalizedToken originalValue="Žijovo">Zijovo</normalizedToken>
Mts,
<normalizedToken originalValue="Šila">Sila</normalizedToken>
Mt env., Gornje
<normalizedToken originalValue="Stravče">Stravce</normalizedToken>
, Katun Guzovalja, 1690 m a.s.l., Prometheus abyss (-130 m). Other localities: Montenegro,
<normalizedToken originalValue="Žijovo">Zijovo</normalizedToken>
, Borova jama 1, Borova jama 2,
<normalizedToken originalValue="Snježna">Snjezna</normalizedToken>
jama,
<normalizedToken originalValue="Milići-Miliči">Milici-Milici</normalizedToken>
<normalizedToken originalValue="snježnica">snjeznica</normalizedToken>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Distribution: Montenegro,
<normalizedToken originalValue="Kučke">Kucke</normalizedToken>
planine Mts (=
<normalizedToken originalValue="Žijovo">Zijovo</normalizedToken>
Mts).
</paragraph>
<paragraph pageId="0" pageNumber="1">
6
<taxonomicName class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops njegosiensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="njegosiensis">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Adriaphaenops njegosiensis</emphasis>
</taxonomicName>
Lohaj, Lakota,
<normalizedToken originalValue="Quéinnec">Queinnec</normalizedToken>
,
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
&amp;
<normalizedToken originalValue="Čeplík">Ceplik</normalizedToken>
, 2016: 526 (
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
). Type locality: Cetinjska
<normalizedToken originalValue="pećina">pecina</normalizedToken>
, Cetinje, Montenegro. Other localities:
<normalizedToken originalValue="Lovćen">Lovcen</normalizedToken>
Mts,
<normalizedToken originalValue="Štirovnik">Stirovnik</normalizedToken>
, Dvogrla jama (synonym =
<normalizedToken originalValue="Kétlyukú-barlang">Ketlyuku-barlang</normalizedToken>
), -130 m (T.
<normalizedToken originalValue="Delić">Delic</normalizedToken>
lgt., new locality).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Distribution: Montenegro, Mt
<normalizedToken originalValue="Lovćen">Lovcen</normalizedToken>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
7
<taxonomicName authorityName="Queinnec &amp; Pavicevic" authorityYear="2008" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops perreaui" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="perreaui">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Adriaphaenops perreaui</emphasis>
</taxonomicName>
<normalizedToken originalValue="Quéinnec">Queinnec</normalizedToken>
&amp;
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
, 2008: 144 (
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
) Type locality:
<normalizedToken originalValue="Pećina">Pecina</normalizedToken>
u Mravinjac, alt. 1000 m a.s.l., Turica, Motka,
<normalizedToken originalValue="Bjelašnica">Bjelasnica</normalizedToken>
planina, Bosnia and Hercegovina.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Distribution: Bosnia and Hercegovina, Mt
<normalizedToken originalValue="Bjelašnica">Bjelasnica</normalizedToken>
near Turica, above Popovo polje.
</paragraph>
<paragraph pageId="0" pageNumber="1">
8
<taxonomicName authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops petrimaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="petrimaris">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Adriaphaenops petrimaris</emphasis>
</taxonomicName>
sp. n. (
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
). Type locality: Montenegro, Risan, Kameno more, Velji
<normalizedToken originalValue="Pištet">Pistet</normalizedToken>
,
<normalizedToken originalValue="Pištet">Pistet</normalizedToken>
4 cave (synonym = PT4) (42.55183°N, 18.73864°E).
</paragraph>
<paragraph pageId="0" pageNumber="1">Distribution: Montenegro, Risan, Kameno more.</paragraph>
<paragraph pageId="0" pageNumber="1">
9
<taxonomicName authorityName="Scheibel" authorityYear="1935" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops pretneri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pretneri">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Adriaphaenops pretneri</emphasis>
</taxonomicName>
Scheibel, 1935b: 35 (
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
). Type locality: Windhohle bei Zavala, Herzegowina [= Vjetrenica
<normalizedToken originalValue="pećina">pecina</normalizedToken>
]. Another locality: Popovo polje,
<normalizedToken originalValue="Turkovići">Turkovici</normalizedToken>
,
<normalizedToken originalValue="Žira">Zira</normalizedToken>
jama (Lohaj et al. 2017).
</paragraph>
<paragraph pageId="0" pageNumber="1">Distribution: Bosnia and Hercegovina, Popovo polje.</paragraph>
<paragraph pageId="0" pageNumber="1">
10
<taxonomicName class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops rumijaensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rumijaensis">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Adriaphaenops rumijaensis</emphasis>
</taxonomicName>
Lohaj, Lakota,
<normalizedToken originalValue="Quéinnec">Queinnec</normalizedToken>
,
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
&amp;
<normalizedToken originalValue="Čeplík">Ceplik</normalizedToken>
, 2016: 522 (
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
). Type locality: Montenegro, Virpazar, Mt Rumija, ca 1100 m a.s.l. Phoenix (cave) (-70 m).
</paragraph>
<paragraph pageId="0" pageNumber="1">Distribution: Montenegro, Mt Rumija.</paragraph>
<paragraph pageId="0" pageNumber="1">
11
<taxonomicName authorityName="Scheibel" authorityYear="1939" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops staudacheri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="staudacheri">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Adriaphaenops staudacheri</emphasis>
</taxonomicName>
Scheibel, 1939: 372 (
<taxonomicName authorityName="Noesske" authorityYear="1928" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Adriaphaenops</emphasis>
</taxonomicName>
). Type locality: in der
<normalizedToken originalValue="“Grbovica“">&quot;Grbovica&quot;</normalizedToken>
, etwa 500 Meter langen
<normalizedToken originalValue="Höhle">Hoehle</normalizedToken>
am Rande des Polje von Trnovo, bei Virpazar in Montenegro [=
<normalizedToken originalValue="Grbočica">Grbocica</normalizedToken>
<normalizedToken originalValue="pećina">pecina</normalizedToken>
]
</paragraph>
<paragraph pageId="0" pageNumber="1">Distribution: Montenegro, Virpazar.</paragraph>
<paragraph pageId="0" pageNumber="1">
12
<taxonomicName baseAuthorityName="Pretner" baseAuthorityYear="1959" class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops stirni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stirni">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Adriaphaenops stirni</emphasis>
</taxonomicName>
Pretner, 1959: 83 (
<taxonomicName authorityName="G. Muller" authorityYear="1913" class="Insecta" family="Carabidae" genus="Aphaenopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphaenopsis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Aphaenopsis</emphasis>
</taxonomicName>
). Type locality: Velja
<normalizedToken originalValue="Peć">Pec</normalizedToken>
appelatur apud Carev most in margine meridiano regionis
<normalizedToken originalValue="Nikšićko">Niksicko</normalizedToken>
polje (Respublica Montenegro)
</paragraph>
<paragraph pageId="0" pageNumber="1">
Distribution: Montenegro,
<normalizedToken originalValue="Nikšić">Niksic</normalizedToken>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
13
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Aphaenopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphaenopsis zupcense" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zupcense">Aphaenopsis zupcense</taxonomicName>
<taxonomicName class="Insecta" family="Carabidae" genus="Aphaenopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphaenopsis zupcense" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zupcense">Aphaenopsis zupcense</taxonomicName>
</emphasis>
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
, 1990: 365 (
<taxonomicName authorityName="G. Muller" authorityYear="1913" class="Insecta" family="Carabidae" genus="Aphaenopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphaenopsis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Aphaenopsis</emphasis>
</taxonomicName>
). Type locality: Durmitor:
<normalizedToken originalValue="pećina">pecina</normalizedToken>
u Zupcima (Sedleni do, 1900-2000 m a.s.l.)
</paragraph>
<paragraph pageId="0" pageNumber="1">
14
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Aphaenopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphaenopsis zupcense" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zupcense">Aphaenopsis zupcense</taxonomicName>
<taxonomicName class="Insecta" family="Carabidae" genus="Adriaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adriaphaenops tartariensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tartariensis">Adriaphaenops tartariensis</taxonomicName>
</emphasis>
<normalizedToken originalValue="Pavićević">Pavicevic</normalizedToken>
, 2001: 35 (
<taxonomicName authority="de Bonvouloir, 1862" authorityName="de Bonvouloir" authorityYear="1862" class="Insecta" family="Carabidae" genus="Aphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aphaenops" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Aphaenops</emphasis>
</taxonomicName>
). Type locality: Montenegro, Mt Durmitor, Jama na Vjetrenim Brdima (&quot;Pit on the Windy Hills&quot;), 2196 m a.s.l. (entrance).
</paragraph>
<paragraph pageId="0" pageNumber="1">Distribution: Montenegro, Mt Durmitor.</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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