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<document id="062803E3A6870984B6A745777D6B5F02" ID-DOI="10.3853/j.2201-4349.69.2017.1676" ID-ISSN="2201-4349" ID-Zenodo-Dep="4676991" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617989790948" checkinUser="carolina" docAuthor="Hyman, Isabel T. &amp; Köhler, Frank" docDate="2017" docId="03DD87CF9371FFAECAB29FEA0A2E84D0" docLanguage="en" docName="RecAustMus.69.2.65-72.pdf" docOrigin="Records of the Australian Museum 69 (2)" docSource="http://dx.doi.org/10.3853/j.2201-4349.69.2017.1676" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Attenborougharion rubicundus Hyman &amp; Köhler 2017" docType="treatment" docVersion="3" lastPageNumber="69" masterDocId="FFE4FFB79375FFAAC92E9F7C0F05807C" masterDocTitle="Attenborougharion gen. nov. (Mollusca: Pulmonata: Helicarionidae): A Likely Case of Convergent Evolution in Southeastern Tasmania" masterLastPageNumber="72" masterPageNumber="65" pageNumber="69" updateTime="1698904482992" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="6C2B7E65BDDEB522401EA3E8B2F07612" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="00A74B0973AB5090CDF38EB8D5169FDD">
<mods:title id="33D6A861F5281AE1F2E4AAB65D8ECB52">Attenborougharion gen. nov. (Mollusca: Pulmonata: Helicarionidae): A Likely Case of Convergent Evolution in Southeastern Tasmania</mods:title>
</mods:titleInfo>
<mods:name id="1237C4257EF93A2D25037785BC632167" type="personal">
<mods:role id="73EF6CDD629A6C02AD401349084A4185">
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</mods:role>
<mods:namePart id="3CB73597E17175C3A4C0DF9B421F73F4">Hyman, Isabel T.</mods:namePart>
</mods:name>
<mods:name id="F9C21A836225F40D5189784B8048BE7B" type="personal">
<mods:role id="1FC97D208EFB866C635958996B5B78BB">
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<mods:namePart id="E1E89F96C7593965E5CAF84E0D788640">Köhler, Frank</mods:namePart>
</mods:name>
<mods:typeOfResource id="38F7B8DFF431E45F6FFB3BE35D866932">text</mods:typeOfResource>
<mods:relatedItem id="7CFE897B0F93C8235970706F218E25AD" type="host">
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<mods:title id="E75815BCE9F998C63E98AE4C12533B71">Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part id="86CAE08A15F2715C132121F8A554AE0F">
<mods:date id="550D2BCFD1E5038AF436FD2686BA6EC5">2017</mods:date>
<mods:detail id="01B60AA4AC22A3D6151E66BBA88FEF5D" type="pubDate">
<mods:number id="E89DEF01DBC587EEF21EA7E74216B92E">2017-08-09</mods:number>
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<mods:detail id="DFB9764635CC0312DA3307E8AA273714" type="volume">
<mods:number id="07C18FDCB681EAB036ACBB3FA6E02710">69</mods:number>
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<mods:detail id="8A870D366A70F938E24D67C2554B98C2" type="issue">
<mods:number id="C97590BB0B2CF57CACDAC926A9091238">2</mods:number>
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<mods:start id="32328E7C7819D1698FD4CEC03CE35F72">65</mods:start>
<mods:end id="117507A03A1FCBF34905FCD6EF33CA79">72</mods:end>
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<mods:location id="335ADE5AA42E353A10FA9DA78F998821">
<mods:url id="FC6EC74A8533C9E90AADB92CB8E08FCA">http://dx.doi.org/10.3853/j.2201-4349.69.2017.1676</mods:url>
</mods:location>
<mods:classification id="A2D991E927A1E05BC5566999964278D7">journal article</mods:classification>
<mods:identifier id="1E67A6DB6BFA85CCA19A9601734AA915" type="DOI">10.3853/j.2201-4349.69.2017.1676</mods:identifier>
<mods:identifier id="A002B23346F71BAD9F885B86C0878DFB" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="EC116F6692EFD5A5DF7474F99C48787C" type="Zenodo-Dep">4676991</mods:identifier>
</mods:mods>
<treatment id="03DD87CF9371FFAECAB29FEA0A2E84D0" LSID="urn:lsid:plazi:treatment:03DD87CF9371FFAECAB29FEA0A2E84D0" httpUri="http://treatment.plazi.org/id/03DD87CF9371FFAECAB29FEA0A2E84D0" lastPageNumber="69" pageId="4" pageNumber="69">
<subSubSection id="C36E65529371FFAECAB29FEA0AAC8098" pageId="4" pageNumber="69" type="nomenclature">
<paragraph id="8BCB36D99371FFAECAB29FEA0A6080C9" blockId="4.[924,1381,150,181]" box="[924,1381,150,181]" pageId="4" pageNumber="69">
<heading id="D08381B59371FFAECAB29FEA0A6080C9" box="[924,1381,150,181]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="69" reason="2">
<taxonomicName id="4C744D5A9371FFAECAB29FEA0A6080C9" ID-CoL="JMX5" authorityName="Hyman &amp; Köhler" authorityYear="2017" baseAuthorityName="Dartnall &amp; Kershaw" baseAuthorityYear="1978" box="[924,1381,150,181]" class="Gastropoda" family="Helicarionidae" genus="Attenborougharion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="69" phylum="Mollusca" rank="species" species="rubicundus" status="comb. nov.">
<emphasis id="B900EACB9371FFAECAB29FEA0A6080C9" bold="true" box="[924,1381,150,181]" italics="true" pageId="4" pageNumber="69">Attenborougharion rubicundus</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCB36D99371FFAECA779FB80AAC8098" blockId="4.[857,1449,196,228]" box="[857,1449,196,228]" pageId="4" pageNumber="69">
<heading id="D08381B59371FFAECA779FB80AAC8098" bold="true" box="[857,1449,196,228]" centered="true" fontSize="13" level="2" pageId="4" pageNumber="69" reason="4">
<emphasis id="B900EACB9371FFAECA779FB80AAC8098" bold="true" box="[857,1449,196,228]" pageId="4" pageNumber="69">(Dartnall &amp; Kershaw, 1978) comb. nov.</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36E65529371FFAECD639F810BB08164" box="[1101,1205,253,280]" pageId="4" pageNumber="69" type="description">
<paragraph id="8BCB36D99371FFAECD639F810BB08164" blockId="4.[1101,1205,253,280]" box="[1101,1205,253,280]" pageId="4" pageNumber="69">
<figureCitation id="134F2A5C9371FFAECD639F810BB08164" box="[1101,1205,253,280]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="1.[121,184,1070,1092]" captionStartId-1="2.[346,409,1157,1179]" captionStartId-2="3.[121,184,1233,1255]" captionStartId-3="5.[121,184,1248,1270]" captionTargetBox-0="[121,1421,151,1058]" captionTargetBox-1="[353,1263,160,1128]" captionTargetBox-2="[134,1421,151,1217]" captionTargetBox-3="[129,1421,166,1223]" captionTargetId-0="figure-485@1.[121,1421,151,1058]" captionTargetId-1="figure-386@2.[346,1280,152,1137]" captionTargetId-2="figure-459@3.[121,1421,151,1217]" captionTargetId-3="figure-10@5.[121,1421,151,1233]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="5" captionText-0="Figure 1. Living animal of Attenborougharion rubicundus from Forestier Peninsula (QVM 9:15514). Courtesy of Simon Grove, TMAG." captionText-1="Figure 2. Comparative external morphology. (A) Attenborougharion rubicundus from Eaglehawk Neck (QVM 9:9665). (B) Helicarion cuvieri from Forestier Peninsula (ex-QVM:9:9671). Scale" captionText-2="Figure 3. Shell of Attenborougharion rubicundus from Forestier Peninsula QVM 9:15514). (A) Photograph of shell viewed from top, front, and below. Scale bar = 5 mm. (B) Scanning electron micrograph showing protoconch. (C) Scanning electron micrograph showing micro-sculpture of fine radial wrinkles near suture on protoconch." captionText-3="Figure 4. Reproductive anatomy of Attenborougharion rubicundus from Forestier Peninsula (QVM 9:15514). (A) Reproductive system. (B) Interior of penis. Scale bars = 5 mm.Abbreviations:alb, albumen gland;bc, bursa copulatrix;ca, caecum;ep, epiphallus;fl, flagellum; her, hermaphroditic duct; p, penis; pp, penial pilasters; prm, penial retractor muscle; ut, uterus; vd, vas deferens." figureDoi-0="http://doi.org/10.5281/zenodo.4676994" figureDoi-1="http://doi.org/10.5281/zenodo.4676996" figureDoi-2="http://doi.org/10.5281/zenodo.4676998" figureDoi-3="http://doi.org/10.5281/zenodo.4677000" httpUri-0="https://zenodo.org/record/4676994/files/figure.png" httpUri-1="https://zenodo.org/record/4676996/files/figure.png" httpUri-2="https://zenodo.org/record/4676998/files/figure.png" httpUri-3="https://zenodo.org/record/4677000/files/figure.png" pageId="4" pageNumber="69">Figs 1–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36E65529371FFAECA4B9E500B498127" pageId="4" pageNumber="69" type="reference_group">
<paragraph id="8BCB36D99371FFAECA4B9E500B498127" blockId="4.[869,1435,300,348]" pageId="4" pageNumber="69">
<treatmentCitationGroup id="AB6411F79371FFAECA4B9E500B498127" pageId="4" pageNumber="69">
<taxonomicName id="4C744D5A9371FFAECA4B9E500A92813D" ID-CoL="3K686" authority="Dartnall &amp; Kershaw, 1978: 2" authorityName="Dartnall &amp; Kershaw" authorityPageNumber="2" authorityYear="1978" box="[869,1431,300,322]" class="Gastropoda" family="Helicarionidae" genus="Helicarion" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="69" phylum="Mollusca" rank="species" species="rubicundus">
<emphasis id="B900EACB9371FFAECA4B9E500B57813D" box="[869,1106,300,321]" italics="true" pageId="4" pageNumber="69">Helicarion rubicundus</emphasis>
Dartnall &amp; Kershaw, 1978: 2
</taxonomicName>
;
<treatmentCitation id="0AD510C89371FFAECAAA9E3A0B4D8127" author="Kershaw, R" box="[900,1096,326,348]" page="213" pageId="4" pageNumber="69" year="1980">
<bibRefCitation id="EFE54B289371FFAECAAA9E3A0B4D8127" author="Kershaw, R" box="[900,1096,326,348]" pageId="4" pageNumber="69" pagination="213 - 214" refId="ref4390" refString="Kershaw, R. C. 1980. Notes on Helicarion rubicundus (Pulmonata: Helicarionidae). Journal of the Malacological Society of Australia 4: 213 - 214." type="journal article" year="1980">Kershaw, 1980: 213</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36E65529371FFAECA699E060BC2827B" pageId="4" pageNumber="69" type="materials_examined">
<paragraph id="8BCB36D99371FFAECA699E060BC2827B" blockId="4.[839,1468,378,520]" pageId="4" pageNumber="69">
<emphasis id="B900EACB9371FFAECA699E060B2C81EE" bold="true" box="[839,1065,378,402]" pageId="4" pageNumber="69">Material examined</emphasis>
.
<materialsCitation id="3B1C3C849371FFAECD159E060BC2827B" collectingDate="1975-01-09" collectionCode="TMH" collectorName="Eagle Hawk Neck &amp; Dartnall &amp; Table" location="Eagle Hawk Neck" municipality="Hawkes Hill" pageId="4" pageNumber="69" specimenCode="E9028, QVM 9" specimenCount="2" stateProvince="Tasmania" typeStatus="holotype">
<typeStatus id="54CF887B9371FFAECD159E060BA381EE" box="[1083,1190,378,402]" pageId="4" pageNumber="69" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED65AE1C9371FFAECD9D9E060BEA81ED" box="[1203,1263,378,401]" country="Australia" httpUri="http://grbio.org/cool/qxtm-hypi" name="Tasmanian Museum and Art Gallery" pageId="4" pageNumber="69" type="Museum">TMH</collectionCode>
<specimenCode id="DBD29EA29371FFAECDD49E060A4781EE" box="[1274,1346,378,402]" collectionCode="TMH" country="Australia" httpUri="http://grbio.org/cool/qxtm-hypi" name="Tasmanian Museum and Art Gallery" pageId="4" pageNumber="69" type="Museum">E9028</specimenCode>
(northeast flank of
<collectingMunicipality id="6BAFACA39371FFAECA8D9EEB0B2B81D3" box="[931,1070,407,431]" pageId="4" pageNumber="69">Hawkes Hill</collectingMunicipality>
, near
<location id="8EAB60029371FFAECD5E9EEB0A3C81D3" LSID="urn:lsid:plazi:treatment:03DD87CF9371FFAECAB29FEA0A2E84D0:8EAB60029371FFAECD5E9EEB0A3C81D3" box="[1136,1337,407,431]" municipality="Hawkes Hill" name="Eagle Hawk Neck" pageId="4" pageNumber="69" stateProvince="Tasmania">Eagle Hawk Neck</location>
,
<collectingRegion id="49B0F83B9371FFAECC6A9EEB0AAA81D3" box="[1348,1455,407,431]" country="Australia" name="Tasmania" pageId="4" pageNumber="69">Tasmania</collectingRegion>
);
<typeStatus id="54CF887B9371FFAECA699EC90CA881B1" box="[839,941,437,461]" pageId="4" pageNumber="69" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D72FD509371FFAECA939EC90B4C81B0" box="[957,1097,436,461]" pageId="4" pageNumber="69" type="generic">2 specimens</specimenCount>
):
<specimenCode id="DBD29EA29371FFAECD759EC90BB581B1" box="[1115,1200,437,461]" collectionCode="QVM" country="Australia" httpUri="http://grbio.org/cool/2s8k-2m7w" name="Queen Victoria Museum" pageId="4" pageNumber="69">QVM 9</specimenCode>
:184, previously labelled
<date id="FFCA10199371FFAECA679EAE0CAF8196" box="[841,938,466,490]" pageId="4" pageNumber="69" value="1975-01-09">
<collectingDate id="EF8EE9F19371FFAECA679EAE0CAF8196" box="[841,938,466,490]" pageId="4" pageNumber="69" value="1975-01-09">1975/9/1</collectingDate>
</date>
,
<collectorName id="2681530F9371FFAECA9D9EAE0B708196" box="[947,1141,466,490]" pageId="4" pageNumber="69">Eagle Hawk Neck</collectorName>
,
<collectingRegion id="49B0F83B9371FFAECD509EAE0BE28196" box="[1150,1255,466,490]" country="Australia" name="Tasmania" pageId="4" pageNumber="69">Tasmania</collectingRegion>
, leg.
<collectorName id="2681530F9371FFAECC329EAE0A738196" box="[1308,1398,466,490]" pageId="4" pageNumber="69">Dartnall</collectorName>
, 1974; and non-type material (see
<collectorName id="2681530F9371FFAECD479E930BA1827B" box="[1129,1188,495,519]" pageId="4" pageNumber="69">Table</collectorName>
1).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36E65529371FFAECD199D540A2E84D0" pageId="4" pageNumber="69" type="description">
<paragraph id="8BCB36D99371FFAECD199D540BCE823E" blockId="4.[1079,1227,552,578]" box="[1079,1227,552,578]" pageId="4" pageNumber="69">
<emphasis id="B900EACB9371FFAECD199D540BCE823E" bold="true" box="[1079,1227,552,578]" pageId="4" pageNumber="69">Description</emphasis>
</paragraph>
<paragraph id="8BCB36D99371FFAECA699D2B0A4F8363" blockId="4.[839,1467,599,799]" pageId="4" pageNumber="69">
<emphasis id="B900EACB9371FFAECA699D2B0B368213" box="[839,1075,599,623]" italics="true" pageId="4" pageNumber="69">External appearance</emphasis>
(
<figureCitation id="134F2A5C9371FFAECD6B9D2B0BA88213" box="[1093,1197,599,623]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="1.[121,184,1070,1092]" captionStartId-1="2.[346,409,1157,1179]" captionStartId-2="3.[121,184,1233,1255]" captionTargetBox-0="[121,1421,151,1058]" captionTargetBox-1="[353,1263,160,1128]" captionTargetBox-2="[134,1421,151,1217]" captionTargetId-0="figure-485@1.[121,1421,151,1058]" captionTargetId-1="figure-386@2.[346,1280,152,1137]" captionTargetId-2="figure-459@3.[121,1421,151,1217]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Figure 1. Living animal of Attenborougharion rubicundus from Forestier Peninsula (QVM 9:15514). Courtesy of Simon Grove, TMAG." captionText-1="Figure 2. Comparative external morphology. (A) Attenborougharion rubicundus from Eaglehawk Neck (QVM 9:9665). (B) Helicarion cuvieri from Forestier Peninsula (ex-QVM:9:9671). Scale" captionText-2="Figure 3. Shell of Attenborougharion rubicundus from Forestier Peninsula QVM 9:15514). (A) Photograph of shell viewed from top, front, and below. Scale bar = 5 mm. (B) Scanning electron micrograph showing protoconch. (C) Scanning electron micrograph showing micro-sculpture of fine radial wrinkles near suture on protoconch." figureDoi-0="http://doi.org/10.5281/zenodo.4676994" figureDoi-1="http://doi.org/10.5281/zenodo.4676996" figureDoi-2="http://doi.org/10.5281/zenodo.4676998" httpUri-0="https://zenodo.org/record/4676994/files/figure.png" httpUri-1="https://zenodo.org/record/4676996/files/figure.png" httpUri-2="https://zenodo.org/record/4676998/files/figure.png" pageId="4" pageNumber="69">Figs 1–3</figureCitation>
): Large semislug, body
<quantity id="4C8C9B3C9371FFAECA699D080CD282F0" box="[839,983,628,652]" metricMagnitude="-2" metricUnit="m" metricValue="3.625" metricValueMax="4.5" metricValueMin="2.75" pageId="4" pageNumber="69" unit="mm" value="36.25" valueMax="45.0" valueMin="27.5">27.5–45 mm</quantity>
in alcohol, shell golden, 3.0–3.3 whorls. Head, eyestalks, neck and middle of tail burgundy; shell lappets, mantle lobes, sole and top of tail green; mantle lobes edged with burgundy. Border between burgundy and green colouration along tail visible even in spirit-preserved material (although colours cannot be identified).
</paragraph>
<paragraph id="8BCB36D99371FFAECA699C480A2E84D0" blockId="4.[837,1474,820,1196]" pageId="4" pageNumber="69">
<emphasis id="B900EACB9371FFAECA699C480CAE8330" box="[839,939,820,844]" italics="true" pageId="4" pageNumber="69">Genitalia</emphasis>
(
<figureCitation id="134F2A5C9371FFAECA969C480CFD8331" box="[952,1016,820,845]" captionStart="Figure 4" captionStartId="5.[121,184,1248,1270]" captionTargetBox="[129,1421,166,1223]" captionTargetId="figure-10@5.[121,1421,151,1233]" captionTargetPageId="5" captionText="Figure 4. Reproductive anatomy of Attenborougharion rubicundus from Forestier Peninsula (QVM 9:15514). (A) Reproductive system. (B) Interior of penis. Scale bars = 5 mm.Abbreviations:alb, albumen gland;bc, bursa copulatrix;ca, caecum;ep, epiphallus;fl, flagellum; her, hermaphroditic duct; p, penis; pp, penial pilasters; prm, penial retractor muscle; ut, uterus; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.4677000" httpUri="https://zenodo.org/record/4677000/files/figure.png" pageId="4" pageNumber="69">Fig. 4</figureCitation>
): Penis long, tubular, distal half enclosed in penial tunica. Penis interior with 4–5 longitudinal pilasters, smooth basally, pustulose proximally, connected by irregular wavy transverse ridges. Epiphallus slightly greater than penis length, entering penis apically, with a very short epiphallic caecum at point of attachment of penial retractor muscle, about one third along distance between penis and vas deferens. Epiphallic flagellum present, large, curved back alongside penis, with numerous internal cryptae adjacent to the epiphallus, flagellum tip slender with no cryptae. Spermatophore (from Dartnall &amp; Kershaw, 1978: fig. 2) with one large, multiply branched spine at base of capsule, followed by spines of decreasing complexity.
</paragraph>
</subSubSection>
</treatment>
</document>
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