319 lines
33 KiB
XML
319 lines
33 KiB
XML
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<mods:title id="B473CDB5862E5C8A970015AE1571587B">Eight new species and additional records of the Pselaphodes complex from Laos and Vietnam, with a key to known species (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="4FC51A6CBDF6CB27D4E6DA2C819390B6">Yin, Zi-Wei</mods:namePart>
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<mods:namePart id="FB4EA90921DD370F556E4B781034B0A3">Li, Ning</mods:namePart>
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<mods:affiliation id="80EF234B35ADB9B1B3E03EE77B828B00">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China.</mods:affiliation>
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<emphasis id="B90C5669E80551397F44FD8A3D3DFD92" bold="true" box="[274,636,606,629]" italics="true" pageId="5" pageNumber="40">
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Linan fortunatus
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<bibRefCitation id="EFE9F78AE80551397F83FD8A3D3DFD92" author="Yin & Li" box="[469,636,606,629]" firstAuthor="Yin" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="5" pageNumber="40" pagination="141 - 153" part="53" refId="ref13570" refString="YIN Z. - W. & LI L. - Z. 2013 a: Five new species of the genera Labomimus and Linan from Guangxi, South China (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae 53 (1): 141 - 153." title="Five new species of the genera Labomimus and Linan from Guangxi, South China (Coleoptera: Staphylinidae: Pselaphinae)" type="journal article" year="2013">Yin & Li, 2013</bibRefCitation>
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,
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(
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16.V.
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, sift 06,
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lgt’ (
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,
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;
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, ‘
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13.V.
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|
||
<elevation id="00556D48E80551397B24FF0A3C0FFEEB" pageId="5" pageNumber="40">1128 m</elevation>
|
||
, sift 03,
|
||
<collectorName id="268DEFADE80551397DC4FF2D3B56FEEB" box="[914,1047,249,268]" pageId="5" pageNumber="40">V.Grebennikov</collectorName>
|
||
lgt’ (
|
||
<collectionCode id="ED6912BEE80551397A16FF2D3BC5FEEB" box="[1088,1156,249,268]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="40" type="Museum">MHNG</collectionCode>
|
||
,
|
||
<collectionCode id="ED6912BEE80551397ADDFF2D3B86FEEB" box="[1163,1223,249,268]" collectionName="China, Shanghai, Shanghai Normal University, Department of Biology, insect Collection" pageId="5" pageNumber="40">SNUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B108026E80551397A83FF2D3CBFFEA6" ID-GBIF-Occurrence="3898373306" collectingDate="2012-05-17" collectionCode="MHNG" collectorName="V. Grebennikov" country="Vietnam" elevation="0" latitude="21.462223" location="Tam Dao" longLatPrecision="21" longitude="105.648605" pageId="5" pageNumber="40" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Grebennikov Lgt'">
|
||
<specimenCount id="9D7E41F2E80551397A83FF2D3BB2FEEA" box="[1237,1267,244,269]" pageId="5" pageNumber="40" type="male">1 ♂</specimenCount>
|
||
<specimenCount id="9D7E41F2E80551397AA1FF2D3A62FEEC" box="[1271,1315,242,268]" pageId="5" pageNumber="40" type="female">4♀♀</specimenCount>
|
||
, ‘
|
||
<collectingCountry id="F36FCAEBE80551397B67FF2D3AD8FEEB" box="[1329,1433,249,268]" name="Vietnam" pageId="5" pageNumber="40" value="Vietnam">VIETNAM</collectingCountry>
|
||
,
|
||
<location id="8EA7DCA0E80551397D6AFEC03CCAFEC0" LSID="urn:lsid:plazi:treatment:03D13B6DE805513A7F44FD8A3DB9FED2:8EA7DCA0E80551397D6AFEC03CCAFEC0" box="[828,907,276,295]" country="Vietnam" latitude="21.462223" longLatPrecision="21" longitude="105.648605" name="Tam Dao" pageId="5" pageNumber="40" stateProvince="Grebennikov Lgt'">Tam Dao</location>
|
||
,
|
||
<geoCoordinate id="EE4CECBCE80551397DC4FEC03CBBFEC0" box="[914,1018,276,295]" degrees="21" direction="north" minutes="27" orientation="latitude" pageId="5" pageNumber="40" precision="15" seconds="44" value="21.462223">N21°27′44″</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4CECBCE80551397A57FEC03B30FEC0" box="[1025,1137,276,295]" degrees="105" direction="east" minutes="38" orientation="longitude" pageId="5" pageNumber="40" precision="15" seconds="55" value="105.648605">E105°38′55″</geoCoordinate>
|
||
,
|
||
<date id="FFC6ACBBE80551397A2FFEC03B93FEC0" box="[1145,1234,276,295]" pageId="5" pageNumber="40" value="2012-05-17">
|
||
<collectingDate id="EF825553E80551397A2FFEC03B93FEC0" box="[1145,1234,276,295]" pageId="5" pageNumber="40" value="2012-05-17">
|
||
17.V.
|
||
<quantity id="4C80279EE80551397AF5FEC03B93FEC0" box="[1187,1234,276,295]" metricMagnitude="3" metricUnit="m" metricValue="2.0121185" pageId="5" pageNumber="40" unit="m" value="2012.1185">2012</quantity>
|
||
</collectingDate>
|
||
</date>
|
||
,
|
||
<elevation id="00556D48E80551397A8FFEC03A42FEC0" box="[1241,1283,276,295]" pageId="5" pageNumber="40">1185</elevation>
|
||
m, sift 07,
|
||
<collectorName id="268DEFADE80551397B09FEC03CCBFEA6" pageId="5" pageNumber="40">V. Grebennikov</collectorName>
|
||
lgt’ (
|
||
<collectionCode id="ED6912BEE80551397DE5FEFA3CB9FEA6" box="[947,1016,302,321]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="40" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B108026E80551397A53FEFA3AC7FEBB" ID-GBIF-Occurrence="3898371318" collectingDate="2012-05-12" collectionCode="MHNG" collectorName="V. Grebennikov'" country="Vietnam" elevation="0" latitude="21.465" location="Tam Dao" longLatPrecision="21" longitude="105.64555" municipality="Tam" pageId="5" pageNumber="40" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Grebennikov Lgt'">
|
||
<specimenCount id="9D7E41F2E80551397A53FEFA3B73FEA5" box="[1029,1074,297,322]" pageId="5" pageNumber="40" type="male">2 ♂♂</specimenCount>
|
||
<specimenCount id="9D7E41F2E80551397A60FEFA3B23FEA6" box="[1078,1122,296,321]" pageId="5" pageNumber="40" type="female">2♀♀</specimenCount>
|
||
, ‘
|
||
<collectingCountry id="F36FCAEBE80551397A26FEFA3B96FEA6" box="[1136,1239,302,321]" name="Vietnam" pageId="5" pageNumber="40" value="Vietnam">VIETNAM</collectingCountry>
|
||
,
|
||
<location id="8EA7DCA0E80551397A88FEFA3A6DFEA6" LSID="urn:lsid:plazi:treatment:03D13B6DE805513A7F44FD8A3DB9FED2:8EA7DCA0E80551397A88FEFA3A6DFEA6" box="[1246,1324,302,321]" country="Vietnam" latitude="21.465" longLatPrecision="21" longitude="105.64555" municipality="Tam" name="Tam Dao" pageId="5" pageNumber="40" stateProvince="Grebennikov Lgt'">Tam Dao</location>
|
||
,
|
||
<geoCoordinate id="EE4CECBCE80551397B65FEFA3AD8FEA6" box="[1331,1433,302,321]" degrees="21" direction="north" minutes="27" orientation="latitude" pageId="5" pageNumber="40" precision="15" seconds="54" value="21.465">N21°27′54″</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4CECBCE80551397D6AFE9D3CEDFEBB" box="[828,940,329,348]" degrees="105" direction="east" minutes="38" orientation="longitude" pageId="5" pageNumber="40" precision="15" seconds="44" value="105.64555">E105°38′44″</geoCoordinate>
|
||
,
|
||
<date id="FFC6ACBBE80551397DE3FE9D3B4FFEBB" box="[949,1038,329,348]" pageId="5" pageNumber="40" value="2012-05-12">
|
||
<collectingDate id="EF825553E80551397DE3FE9D3B4FFEBB" box="[949,1038,329,348]" pageId="5" pageNumber="40" value="2012-05-12">
|
||
12.V.
|
||
<quantity id="4C80279EE80551397D89FE9D3B4FFEBB" box="[991,1038,329,348]" metricMagnitude="3" metricUnit="m" metricValue="2.0121218" pageId="5" pageNumber="40" unit="m" value="2012.1218">2012</quantity>
|
||
</collectingDate>
|
||
</date>
|
||
,
|
||
<elevation id="00556D48E80551397A41FE9D3B1BFEBB" box="[1047,1114,329,348]" pageId="5" pageNumber="40">1218 m</elevation>
|
||
, sift 02,
|
||
<collectorName id="268DEFADE80551397AF2FE9D3A70FEBB" box="[1188,1329,329,348]" pageId="5" pageNumber="40">V. Grebennikov’</collectorName>
|
||
(
|
||
<collectionCode id="ED6912BEE80551397B6AFE9D3AC1FEBB" box="[1340,1408,329,348]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="40" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B108026E80551397BC7FE9D3BFEFE76" ID-GBIF-Occurrence="3898371322" collectingDate="2012-05-14" collectionCode="MHNG" collectorName="V. Grebennikov" country="Vietnam" elevation="0" latitude="21.465" location="Tam Dao" longLatPrecision="21" longitude="105.64889" pageId="5" pageNumber="40" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Grebennikov Lgt'">
|
||
<specimenCount id="9D7E41F2E80551397BC7FE9D3C1AFE90" pageId="5" pageNumber="40" type="male">3 ♂♂</specimenCount>
|
||
<specimenCount id="9D7E41F2E80551397D09FEB03CCBFE91" box="[863,906,349,375]" pageId="5" pageNumber="40" type="female">1♀♀</specimenCount>
|
||
, ‘
|
||
<collectingCountry id="F36FCAEBE80551397DCEFEB03CBEFE90" box="[920,1023,356,375]" name="Vietnam" pageId="5" pageNumber="40" value="Vietnam">VIETNAM</collectingCountry>
|
||
,
|
||
<location id="8EA7DCA0E80551397A53FEB03B12FE90" LSID="urn:lsid:plazi:treatment:03D13B6DE805513A7F44FD8A3DB9FED2:8EA7DCA0E80551397A53FEB03B12FE90" box="[1029,1107,356,375]" country="Vietnam" latitude="21.465" longLatPrecision="21" longitude="105.64889" name="Tam Dao" pageId="5" pageNumber="40" stateProvince="Grebennikov Lgt'">Tam Dao</location>
|
||
,
|
||
<geoCoordinate id="EE4CECBCE80551397A0CFEB03B82FE90" box="[1114,1219,356,375]" degrees="21" direction="north" minutes="27" orientation="latitude" pageId="5" pageNumber="40" precision="15" seconds="54" value="21.465">N 21°27′54″</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4CECBCE80551397A9CFEB03A7AFE90" box="[1226,1339,356,375]" degrees="105" direction="east" minutes="38" orientation="longitude" pageId="5" pageNumber="40" precision="15" seconds="56" value="105.64889">E 105°38′56″</geoCoordinate>
|
||
,
|
||
<date id="FFC6ACBBE80551397B14FEB03AD8FE90" box="[1346,1433,356,375]" pageId="5" pageNumber="40" value="2012-05-14">
|
||
<collectingDate id="EF825553E80551397B14FEB03AD8FE90" box="[1346,1433,356,375]" pageId="5" pageNumber="40" value="2012-05-14">
|
||
14.V.
|
||
<quantity id="4C80279EE80551397B3DFEB03AD8FE90" box="[1387,1433,356,375]" metricMagnitude="3" metricUnit="m" metricValue="2.0121237" pageId="5" pageNumber="40" unit="m" value="2012.1237">2012</quantity>
|
||
</collectingDate>
|
||
</date>
|
||
,
|
||
<elevation id="00556D48E80551397D6AFEAA3C3CFE76" box="[828,893,382,401]" pageId="5" pageNumber="40">1237 m</elevation>
|
||
, sift 04,
|
||
<collectorName id="268DEFADE80551397D95FEAA3B09FE76" box="[963,1096,382,401]" pageId="5" pageNumber="40">V. Grebennikov</collectorName>
|
||
lgt’ (
|
||
<collectionCode id="ED6912BEE80551397A24FEAA3BF9FE76" box="[1138,1208,382,401]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="40" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B108026E80551397A90FEAA3A33FE4B" ID-GBIF-Occurrence="3898318326" collectingDate="2003-05-13" collectionCode="MHNG" collectorName="S Nomura" country="Iceland" location="Mt. Tam Dao" pageId="5" pageNumber="40" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phu Prov">
|
||
<specimenCount id="9D7E41F2E80551397A90FEAA3BA4FE75" box="[1222,1253,377,402]" pageId="5" pageNumber="40" type="male">1 ♂</specimenCount>
|
||
, ‘
|
||
<location id="8EA7DCA0E80551397AA4FEAA3A24FE76" LSID="urn:lsid:plazi:treatment:03D13B6DE805513A7F44FD8A3DB9FED2:8EA7DCA0E80551397AA4FEAA3A24FE76" box="[1266,1381,382,401]" country="Iceland" name="Mt. Tam Dao" pageId="5" pageNumber="40" stateProvince="Vinh Phu Prov">Mt. Tam Dao</location>
|
||
(950),
|
||
<collectingRegion id="49BC4499E80551397D6AFE4D3CF4FE4B" box="[828,949,409,428]" pageId="5" pageNumber="40">Vinh Phu Prov</collectingRegion>
|
||
C N-VIETNAM],
|
||
<date id="FFC6ACBBE80551397A1EFE4D3BDFFE4B" box="[1096,1182,409,428]" pageId="5" pageNumber="40" value="2003-05-13">
|
||
<collectingDate id="EF825553E80551397A1EFE4D3BDFFE4B" box="[1096,1182,409,428]" pageId="5" pageNumber="40" value="2003-05-13">13.v.2003</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="268DEFADE80551397AF2FE4D3BB8FE4B" box="[1188,1273,409,428]" pageId="5" pageNumber="40">S Nomura</collectorName>
|
||
leg’ (
|
||
<collectionCode id="ED6912BEE80551397B70FE4D3A2DFE4B" box="[1318,1388,409,428]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="40" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B108026E80551397B2FFE4D3A73FE20" ID-GBIF-Occurrence="3898318351" collectingDate="2003-05-21" collectionCode="MHNG" country="Iceland" pageId="5" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Vinh Phu Prov">
|
||
<specimenCount id="9D7E41F2E80551397B2FFE4D3AD6FE4C" box="[1401,1431,402,428]" pageId="5" pageNumber="40" type="female">1 ♀</specimenCount>
|
||
, ‘same locality and collector, except ‘
|
||
<date id="FFC6ACBBE80551397A25FE603B96FE20" box="[1139,1239,436,455]" pageId="5" pageNumber="40" value="2003-05-21">
|
||
<collectingDate id="EF825553E80551397A25FE603B96FE20" box="[1139,1239,436,455]" pageId="5" pageNumber="40" value="2003-05-21">21. v. 2003</collectingDate>
|
||
</date>
|
||
’ (
|
||
<collectionCode id="ED6912BEE80551397AB3FE603A6AFE20" box="[1253,1323,436,455]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="40" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C362D9F0E805513A7D6AFE0B3E3AFF12" lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40" type="discussion">
|
||
<paragraph id="8BC78A7BE805513A7D6AFE0B3E3AFF12" blockId="5.[828,1437,479,790]" lastBlockId="6.[151,760,190,309]" lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40">
|
||
<emphasis id="B90C5669E80551397D6AFE0B3CFFFE11" bold="true" box="[828,958,479,502]" pageId="5" pageNumber="40">Comments.</emphasis>
|
||
The male of
|
||
<taxonomicName id="4C78F1F8E80551397A18FE0B3B99FE11" authorityName="Yin & Li" authorityYear="2013" box="[1102,1240,479,502]" class="Insecta" family="Staphylinidae" genus="Linan" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="fortunatus">
|
||
<emphasis id="B90C5669E80551397A18FE0B3B1CFE11" box="[1102,1117,479,502]" italics="true" pageId="5" pageNumber="40">L</emphasis>
|
||
.
|
||
<emphasis id="B90C5669E80551397A3CFE0B3B99FE11" box="[1130,1240,479,502]" italics="true" pageId="5" pageNumber="40">fortunatus</emphasis>
|
||
</taxonomicName>
|
||
can be readily separated from all congeners by denser setae at the posterolateral margins of pronotum and the anterolateral margins of elytra, as well as by apically broadened antennomere 9, which is massively impressed on the lateral surface. The female, which was previously unknown, lacks these dense setae on pronotum and elytra. This species was originally described from Guangxi, southern China, and is here reported from northern Vietnam. The populations from China and Vietnam differ only slightly in the shape of the apex of the median lobe and the position of the endophallic sclerites of aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF07DAF3E80551397EC1F7DB3B34F7C5" ID-DOI="http://doi.org/10.5281/zenodo.5037476" ID-Zenodo-Dep="5037476" box="[151,1141,2063,2082]" httpUri="https://zenodo.org/record/5037476/files/figure.png" pageId="5" pageNumber="40" startId="5.[151,182,2063,2082]" targetBox="[156,1431,843,2025]" targetPageId="5">
|
||
<paragraph id="8BC78A7BE80551397EC1F7DB3B34F7C5" blockId="5.[151,1141,2063,2082]" box="[151,1141,2063,2082]" pageId="5" pageNumber="40">
|
||
Fig. 4. Dorsal habitus of
|
||
<taxonomicName id="4C78F1F8E80551397F3AF7DB3E99F7C5" authorityName="Westwood" authorityYear="1870" box="[364,472,2063,2082]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C5669E80551397F3AF7DB3E99F7C5" box="[364,472,2063,2082]" italics="true" pageId="5" pageNumber="40">Pselaphodes</emphasis>
|
||
</taxonomicName>
|
||
males. A –
|
||
<taxonomicName id="4C78F1F8E80551397C6AF7DB3DEDF7C5" authority="Yin & Li, 2021" authorityName="Yin & Li" authorityYear="2021" box="[572,684,2063,2082]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="banphabat" status="sp. nov.">
|
||
<emphasis id="B90C5669E80551397C6AF7DB3DEDF7C5" box="[572,684,2063,2082]" italics="true" pageId="5" pageNumber="40">P. banphabat</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23FEB12E80551397CE7F7DB3DB6F7C5" box="[689,759,2063,2082]" pageId="5" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; B –
|
||
<taxonomicName id="4C78F1F8E80551397D74F7DB3CCFF7C5" authority="Yin & Li, 2021" authorityName="Yin & Li" authorityYear="2021" box="[802,910,2063,2082]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="dalatensis" status="sp. nov.">
|
||
<emphasis id="B90C5669E80551397D74F7DB3CCFF7C5" box="[802,910,2063,2082]" italics="true" pageId="5" pageNumber="40">P. dalatensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23FEB12E80551397DC5F7DB3C94F7C5" box="[915,981,2063,2082]" pageId="5" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C362D9F0E806513A7EC1FF2A3DB9FED2" pageId="6" pageNumber="41" type="distribution">
|
||
<paragraph id="8BC78A7BE806513A7EC1FF2A3DB9FED2" blockId="6.[151,760,190,309]" pageId="6" pageNumber="41">
|
||
<emphasis id="B90C5669E806513A7EC1FF2A3E64FEF2" bold="true" box="[151,293,254,277]" pageId="6" pageNumber="41">Distribution.</emphasis>
|
||
China: Guangxi; Vietnam: Vĩnh Phúc (Y I N & L
|
||
<date id="FFC6ACBBE806513A7EF6FECA3FADFED2" box="[160,236,286,309]" pageId="6" pageNumber="41" value="2013-01">I 2013</date>
|
||
a; this paper).
|
||
<emphasis id="B90C5669E806513A7F28FECA3DB9FED2" bold="true" box="[382,760,286,309]" pageId="6" pageNumber="41">First country record for Vietnam.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |