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<document id="1FD15E8BAC9CA0F922F6339051B8F92E" ID-DOI="10.5281/zenodo.13245290" ID-ISSN="2345-7600" ID-Zenodo-Dep="13245290" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722970560201" checkinUser="felipe" docAuthor="Lambkin, L. A. Ballantyne C." docDate="2006" docId="039D87D4FFDFBA57CAEBBF05FC0AFCB6" docLanguage="en" docName="RafflesBZool.54.1.21-48.pdf" docOrigin="Raffles Bulletin of Zoology 54 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Pygoluciola kinabalua" docType="treatment" docVersion="3" lastPageNumber="29" masterDocId="FFA4FFACFFD7BA5FC950BE65FFF6FFC4" masterDocTitle="A Phylogenetic Reassessment Of The Rare S. E. Asian Firefly Genus Pygoluciola Wittmer (Coleoptera: Lampyridae: Luciolinae)" masterLastPageNumber="48" masterPageNumber="21" pageNumber="29" updateTime="1722977453427" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="434C5DFD9EE4695DA1F7B4E5D482DD9F">A Phylogenetic Reassessment Of The Rare S. E. Asian Firefly Genus Pygoluciola Wittmer (Coleoptera: Lampyridae: Luciolinae)</mods:title>
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<mods:namePart id="DA0BC56810A30DB7B8FD1BE96874176A">Lambkin, L. A. Ballantyne C.</mods:namePart>
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<mods:title id="4C604B3E5D2FE10ECD217330B9D46965">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="5A9E6E8450B2191FA9D24480B3ADB6CD">2006</mods:date>
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<paragraph id="8B8B36C2FFDFBA57CAEBBF05FABFFEBC" blockId="8.[955,1353,352,408]" box="[955,1353,352,376]" pageId="8" pageNumber="29">
<heading id="D0C381AEFFDFBA57CAEBBF05FABFFEBC" bold="true" box="[955,1353,352,376]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="29" reason="2">
<taxonomicName id="4C344D41FFDFBA57CAEBBF05FABFFEBC" ID-CoL="6X2QB" authority="(Ballantyne)" baseAuthorityName="Ballantyne &amp; Lambkin" baseAuthorityYear="2001" box="[955,1353,352,376]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="kinabalua">
<emphasis id="B940EAD0FFDFBA57CAEBBF05FABFFEBC" bold="true" box="[955,1353,352,376]" pageId="8" pageNumber="29">
<emphasis id="B940EAD0FFDFBA57CAEBBF05FB43FEBC" bold="true" box="[955,1205,352,376]" italics="true" pageId="8" pageNumber="29">Pygoluciola kinabalua</emphasis>
(Ballantyne)
</emphasis>
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<paragraph id="8B8B36C2FFDFBA57CD70BFE5FB13FE5C" blockId="8.[955,1353,352,408]" box="[1056,1253,384,408]" pageId="8" pageNumber="29">
(
<figureCitation id="130F2A47FFDFBA57CD78BFE5FB67FE5C" box="[1064,1169,384,408]" captionStart="Figs" captionStartId="4.[136,180,1949,1971]" captionTargetBox="[202,1389,195,1875]" captionTargetId="figure-8@4.[184,1426,191,1909]" captionTargetPageId="4" captionText="Figs. 1-17. Features of male Pygoluciola. 1, 6, 13-17: P. guigliae; 2: Mt Missim male; 3, 7: P. kinabalua holotype male; 4, 10-12: P. stylifer holotype male; 5, 8-9: P. wittmeri. 1-5: dorsal, male pronota (left side shows colour pattern, right side sculpturing; solid line indicates extent of irregular tubercles, dotted lines retraction of fat body): 6,7: dorsal, whole body; aedeagi (8; 10 ventral; 9, 12 dorsal; 11, left lateral); 13: maxillary palp, ventral surface; 14: labial palp, ventral surface; 15-17: aedeagal sheath, dorsal, ventral, and lateral. These figures share scale lines: 1-5; 6-7; 8-14; 15-17. Figure legend: A, anus; BP, basal piece; EO, ejaculatory orifice; LL, lateral lobe; ML, median lobe; TS, tergite aedeagal sheath; VS, ventrite aedeagal sheath." figureDoi="http://doi.org/10.5281/zenodo.13245292" httpUri="https://zenodo.org/record/13245292/files/figure.png" pageId="8" pageNumber="29">Figs. 3, 7</figureCitation>
,
<figureCitation id="130F2A47FFDFBA57CDCEBFE5FB2BFE5C" box="[1182,1245,384,408]" captionStart-0="Figure 38" captionStart-1="Figure 39" captionStart-2="Figure 40" captionStartId-0="8.[136,199,2035,2057]" captionStartId-1="9.[112,175,2064,2086]" captionStartId-2="10.[136,199,2036,2058]" captionTargetBox-0="[141,1473,1000,1981]" captionTargetBox-1="[160,1412,199,2026]" captionTargetBox-2="[211,1412,205,1964]" captionTargetId-0="figure-386@8.[114,1492,968,2002]" captionTargetId-1="figure-45@9.[127,1438,171,2043]" captionTargetId-2="figure-61@10.[176,1439,177,1991]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="Figure 38. (A). The strict consensus of the 2992 MPT for 49 taxa of the Luciolinae found with phylogenetic analysis of 115 characters, excluding 10 colour and mimicry characters. (B). The majority rule consensus of the 2992 MPT for the Luciolinae, showing above the branch the percentage of MPT retaining that node. 153 of the MPT were compatible with the majority rule consensus." captionText-1="Figure 39. Semistrict consensus of 195 successively weighted trees. Percentage of successively weighted trees with this resolution shown above the branch. None of the MPT was compatible with the semistrict consensus successively weighted trees." captionText-2="Figure 40. Preferred MPT from 153 filtered MPT compatible with the majority rule consensus of the 2992 MPT, Part 1, without Atyphella clade. Unambiguous character and state changes were mapped using MacClade. Colour and mimicry characters, not used in the phylogenetic analysis, are enclosed in boxes. Bremer support values shown above nodes." figureDoi-0="http://doi.org/10.5281/zenodo.13245298" figureDoi-1="http://doi.org/10.5281/zenodo.13245302" figureDoi-2="http://doi.org/10.5281/zenodo.13245304" httpUri-0="https://zenodo.org/record/13245298/files/figure.png" httpUri-1="https://zenodo.org/record/13245302/files/figure.png" httpUri-2="https://zenodo.org/record/13245304/files/figure.png" pageId="8" pageNumber="29">38-40</figureCitation>
)
</paragraph>
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<subSubSection id="C32E6549FFDFBA57CA13BFA5FA68FE36" pageId="8" pageNumber="29" type="reference_group">
<paragraph id="8B8B36C2FFDFBA57CA13BFA5FA68FE36" blockId="8.[835,1474,448,499]" pageId="8" pageNumber="29">
<treatmentCitationGroup id="AB2411ECFFDFBA57CA13BFA5FA68FE36" pageId="8" pageNumber="29">
<treatmentCitation id="0A9510D3FFDFBA57CA13BFA5FA34FE11" author="Lambkin, C" box="[835,1474,448,470]" page="371" pageId="8" pageNumber="29" year="2001">
<taxonomicName id="4C344D41FFDFBA57CA13BFA5FA34FE11" ID-CoL="3WCSR" authority="Ballantyne &amp; Lambkin, 2001: 371" authorityName="Ballantyne &amp; Lambkin" authorityPageNumber="371" authorityYear="2001" box="[835,1474,448,470]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="kinabalua" subGenus="Pygoluciola">
<emphasis id="B940EAD0FFDFBA57CA13BFA5FB8FFE11" box="[835,1145,448,469]" italics="true" pageId="8" pageNumber="29">Luciola (Pygoluciola) kinabalua</emphasis>
<materialsCitation id="3B5C3C9FFFDFBA57CD2EBFA5FA34FE11" box="[1150,1474,448,470]" collectionCode="BMNH" country="Malaysia" location="Mt Kinabalu" pageId="8" pageNumber="29" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
Ballantyne &amp;
<bibRefCitation id="EFA54B33FFDFBA57CC52BFA5FA34FE11" author="Lambkin, C" box="[1282,1474,448,470]" pageId="8" pageNumber="29" refId="ref20972" refString="Lambkin, C. L., 2001. Systematics and evolution of Australian exoprosopine flies (Diptera: Bombyliidae: Anthracinae) based on molecules, morphology, and colour patterns. Ph. D. Thesis, University of Queensland." type="book" year="2001">Lambkin, 2001: 371</bibRefCitation>
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[
<typeStatus id="548F8860FFDFBA57CA3EBFB8FC3CFE37" box="[878,970,477,499]" pageId="8" pageNumber="29" type="holotype">Holotype</typeStatus>
male
<collectingCountry id="F3237652FFDFBA57CD57BFB8FB97FE37" box="[1031,1121,477,499]" name="Malaysia" pageId="8" pageNumber="29">Malaysia</collectingCountry>
,
<collectingRegion id="49F0F820FFDFBA57CD3CBFB8FB5CFE37" box="[1132,1194,477,499]" country="Malaysia" name="Sabah" pageId="8" pageNumber="29">Sabah</collectingRegion>
:
<location id="8EEB6019FFDFBA57CDE5BFB8FAC6FE37" LSID="urn:lsid:plazi:treatment:039D87D4FFDFBA57CAEBBF05FC0AFCB6:8EEB6019FFDFBA57CDE5BFB8FAC6FE37" box="[1205,1328,477,499]" country="Malaysia" name="Mt Kinabalu" pageId="8" pageNumber="29" stateProvince="Sabah">Mt Kinabalu</location>
(BMNH)].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32E6549FFDFBA57CA13BC7FFC0AFCB6" pageId="8" pageNumber="29" type="diagnosis">
<paragraph id="8B8B36C2FFDFBA57CA13BC7FFC0AFCB6" blockId="8.[835,1474,538,882]" pageId="8" pageNumber="29">
<emphasis id="B940EAD0FFDFBA57CA13BC7FFC3FFDF5" bold="true" box="[835,969,538,562]" italics="true" pageId="8" pageNumber="29">Diagnosis. </emphasis>
Males elongate slender (
<figureCitation id="130F2A47FFDFBA57CD85BC7FFACFFDF6" box="[1237,1337,538,562]" captionStart="Figs" captionStartId="4.[136,180,1949,1971]" captionTargetBox="[202,1389,195,1875]" captionTargetId="figure-8@4.[184,1426,191,1909]" captionTargetPageId="4" captionText="Figs. 1-17. Features of male Pygoluciola. 1, 6, 13-17: P. guigliae; 2: Mt Missim male; 3, 7: P. kinabalua holotype male; 4, 10-12: P. stylifer holotype male; 5, 8-9: P. wittmeri. 1-5: dorsal, male pronota (left side shows colour pattern, right side sculpturing; solid line indicates extent of irregular tubercles, dotted lines retraction of fat body): 6,7: dorsal, whole body; aedeagi (8; 10 ventral; 9, 12 dorsal; 11, left lateral); 13: maxillary palp, ventral surface; 14: labial palp, ventral surface; 15-17: aedeagal sheath, dorsal, ventral, and lateral. These figures share scale lines: 1-5; 6-7; 8-14; 15-17. Figure legend: A, anus; BP, basal piece; EO, ejaculatory orifice; LL, lateral lobe; ML, median lobe; TS, tergite aedeagal sheath; VS, ventrite aedeagal sheath." figureDoi="http://doi.org/10.5281/zenodo.13245292" httpUri="https://zenodo.org/record/13245292/files/figure.png" pageId="8" pageNumber="29">Figs. 3, 7</figureCitation>
), about three times as long as wide, with elytra subparallel sided; one of two species of
<taxonomicName id="4C344D41FFDFBA57CAB3BC3FFB92FDB6" box="[995,1124,602,626]" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAD0FFDFBA57CAB3BC3FFB92FDB6" box="[995,1124,602,626]" italics="true" pageId="8" pageNumber="29">Pygoluciola</emphasis>
</taxonomicName>
with the apex of the MPP of V7 emarginate, gently so in this species; distinguished from
<taxonomicName id="4C344D41FFDFBA57CCFBBC1EFC6AFD75" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" class="Insecta" family="Lampyridae" genus="Pygoluciola" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="B940EAD0FFDFBA57CCFBBC1EFC6AFD75" italics="true" pageId="8" pageNumber="29">P. wittmeri</emphasis>
</taxonomicName>
, which has a deeply emarginate apex; female with posterior margin of V7 deeply and broadly emarginate, with a small transverse ridge just anterior to this emargination, lateral projections moderately broad and apically rounded; tergite 7 with strongly sclerotised median mound; lateral margins of anterior prolongation of V8 not well sclerotised nor well defined.
</paragraph>
</subSubSection>
</treatment>
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