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<document id="0F82929EAFAC6C7832FB2F643B833B96" ID-CLB-Dataset="39033" ID-DOI="10.11646/zootaxa.3658.1.1" ID-GBIF-Dataset="0adf09ba-5301-4828-abf6-8dbe89f041ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="246551" ID-ZooBank="C02D1C74-25C0-41DD-B098-62098EB7B62A" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1463647879570" checkinUser="plazi" docAuthor="Dragan P. Chobanov, Beata Grzywacz, Ionuţ Ş. Iorgu, Battal Cιplak, Maya B. Ilieva &amp; Elżbieta Warchałowska-Śliwa" docDate="2013" docId="F26F3128392FFFBAB1B00D24FED19F52" docLanguage="en" docName="zt03658p081.pdf.imf" docOrigin="Zootaxa 3658 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Isophya dobrogensis Kis 1994" docType="treatment" docVersion="12" lastPageNumber="41" masterDocId="0E5649503908FF92B1270A03FFD39875" masterDocTitle="Review of the Balkan Isophya (Orthoptera: Phaneropteridae) with particular emphasis on the Isophya modesta group and remarks on the systematics of the genus based on morphological and acoustic data" masterLastPageNumber="81" masterPageNumber="1" pageNumber="40" updateTime="1698662571276" updateUser="ExternalLinkService">
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<mods:title id="F21E9853CFA97D5E62973F0239AE8C54">Review of the Balkan Isophya (Orthoptera: Phaneropteridae) with particular emphasis on the Isophya modesta group and remarks on the systematics of the genus based on morphological and acoustic data</mods:title>
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<mods:namePart id="FCEA25440532DCF211D5116D6370CC54">Dragan P. Chobanov</mods:namePart>
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<mods:namePart id="A5F2763816D3305C99720E799ED644EA">Beata Grzywacz</mods:namePart>
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<mods:namePart id="B98F4374307C93954D3E559347CDFEAB">Ionuţ Ş. Iorgu</mods:namePart>
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<mods:namePart id="680B601647BFA9F82BB8ADE975AD0CB6">Maya B. Ilieva</mods:namePart>
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<mods:namePart id="893F40E4080C19CCB276454FA18D6650">Elżbieta Warchałowska-Śliwa</mods:namePart>
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<treatment id="F26F3128392FFFBAB1B00D24FED19F52" ID-DOI="http://doi.org/10.5281/zenodo.5617383" ID-GBIF-Taxon="120682324" ID-Zenodo-Dep="5617383" LSID="urn:lsid:plazi:treatment:F26F3128392FFFBAB1B00D24FED19F52" httpUri="http://treatment.plazi.org/id/F26F3128392FFFBAB1B00D24FED19F52" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
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<paragraph id="7A79803E392FFFB5B1B00D24FDDA9F34" blockId="39.[151,644,1831,1890]" box="[151,521,1831,1857]" pageId="39" pageNumber="40">
<heading id="21313752392FFFB5B1B00D24FDDA9F34" bold="true" box="[151,521,1831,1857]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
<taxonomicName id="BDC6FBBD392FFFB5B1B00D24FDDA9F34" authority="Kis, 1994" authorityName="Kis" authorityYear="1994" box="[151,521,1831,1857]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="dobrogensis">
<emphasis id="48B25C2C392FFFB5B1B00D24FDDA9F34" bold="true" box="[151,521,1831,1857]" pageId="39" pageNumber="40">
<emphasis id="48B25C2C392FFFB5B1B00D24FE5C9F34" bold="true" box="[151,399,1831,1857]" italics="true" pageId="39" pageNumber="40">Isophya dobrogensis</emphasis>
Kis, 1994
</emphasis>
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</heading>
</paragraph>
<paragraph id="7A79803E392FFFB5B1B00D49FD579F17" blockId="39.[151,644,1831,1890]" box="[151,644,1866,1890]" pageId="39" pageNumber="40">
(
<figureCitation id="E2FD9CBB392FFFB5B1B80D49FECC9F17" box="[159,287,1866,1890]" captionStart="FIGURES 19 30" captionStartId="6.[151,264,1632,1655]" captionTargetBox="[154,1433,187,1606]" captionTargetId="figure@6.[151,1436,183,1611]" captionTargetPageId="6" captionText="FIGURES 19 30. General appearance of some Balkan Isophya; 19 — I. modestior (male); 20, 21 — I. dobrogensis (20 — male, 21 — female); 22, 23 — I. zubowskii (22 — male, 23 — female); 24, 25 — I. camptoxypha (24 — male, 25 — female); 26, 27 — I. gulae (26 — male, 27 — female); 28, 29 — I. obtusa (dark— 28, and green— 29 colour forms; males); 30 — I. speciosa (malebelow, and female—above)." httpUri="https://zenodo.org/record/246553/files/figure.png" pageId="39" pageNumber="40">Figs 20, 21</figureCitation>
,
<figureCitation id="E2FD9CBB392FFFB5B00C0D49FE999F17" box="[299,330,1866,1890]" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="39" pageNumber="40">49</figureCitation>
,
<figureCitation id="E2FD9CBB392FFFB5B0720D49FEA09F17" box="[341,371,1866,1890]" captionStart="FIGURES 56 79" captionStartId="13.[151,264,1769,1792]" captionTargetBox="[178,1421,662,1756]" captionTargetId="figure@13.[178,1422,662,1756]" captionTargetPageId="13" captionText="FIGURES 56 79. Morphology of female head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55 unless specifically given): 56 — I. hospodar; 57 — I. rectipennis; 58 — I. pavelii (paratype of I. rammei Peshev); 59 — I. thracica; 60 — I. bureschi; 61 — I. yaraligozi; 62 — I. tosevski; 63 — I. andreevae (paratype); 64 — I. clara; 65 — I. miksici (paratype); 66 — I. plevnensis (paratype); 67 — I. longicaudata adamovici (paratype, BG: Sliven, 1000 m, NMNHS); 68 — I. l. longicaudata; 69 — I. rhodopensis leonorae (BG: “ Alibotush, 1500 m ”, 13.07. 1959, NMNHS); 70 — I. rh. rhodopensis (as in Fig. 46 but 17.07.1956); 71 — I. rh. petkovi; 72 — I. modestior; 73 — I. dobrogensis; 74 — I. zubowskii; 75 — I. aff. camptoxypha (MK: Bistra Mt., 2.08. 2004, CC); 76 — I. gulae (paratype, BG: Dolna Topchiya, 10.07. 1974, NMNHS); 77 — I. obtusa (BG: Vezhen Peak, 15.07. 1969, NMNHS); 78 — I. amplipennis (TR: “ Turkey | 1960 | Sureya Bey ”, NHM); 79 — I. speciosa (BG: Sakar Mt., 14.06. 1952, NMNHS). Scale (if present; black line right of the specimen) = 10 mm." httpUri="https://zenodo.org/record/246554/files/figure.png" pageId="39" pageNumber="40">73</figureCitation>
,
<figureCitation id="E2FD9CBB392FFFB5B0580D49FE4E9F17" box="[383,413,1866,1890]" captionStart="FIGURES 80 104" captionStartId="14.[151,264,1835,1858]" captionTargetBox="[226,1360,681,1815]" captionTargetId="figure@14.[226,1362,681,1816]" captionTargetPageId="14" captionText="FIGURES 80 104. Male abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55): 80 — I. hospodar; 81 — I. rectipennis; 82 — I. pavelii; 83 — I. thracica; 84 — I. bureschi; 85 — I. yaraligozi; 86 — I. tosevski; 87 — I. andreevae; 88 — I. clara; 89 — I. miksici; 90 — I. plevnensis; 91 — I. longicaudata adamovici; 92 — I. l. longicaudata; 93 — I. m. modesta (= intermedia); 94 — I. rhodopensis leonorae; 95 — I. rh. rhodopensis; 96 — I. rh. petkovi; 97 — I. modestior; 98 — I. dobrogensis; 99 — I. zubowskii; 100 — I. aff. camptoxypha; 101 — I. gulae; 102 — I. obtusa; 103 — I. amplipennis; 104 — I. speciosa. Scale (if present) = 2 mm." httpUri="https://zenodo.org/record/246555/files/figure.png" pageId="39" pageNumber="40">98</figureCitation>
,
<figureCitation id="E2FD9CBB392FFFB5B08E0D49FE069F17" box="[425,469,1866,1890]" captionStart="FIGURES 105 129" captionStartId="17.[151,264,1173,1196]" captionTargetBox="[151,1436,195,1152]" captionTargetId="figure@17.[151,1436,195,1152]" captionTargetPageId="17" captionText="FIGURES 105 129. Female abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55 unless specifically given); figures of the whole ovipositor are followed by enlarged figure of the ovipositor base: 105 — I. hospodar (BG: Dolni Glavanak, 25.04. 2005, CC); 106 — I. rectipennis (BG: Byala Vill., 27.06. 2002, CC); 107 — I. pavelii (BG: Chernogorovo, 29.05. 2006, CC); 108 — I. thracica; 109 — I. bureschi (BG: Bansko, 8.08. 2006, CC); 110 — I. yaraligozi; 111 — I. tosevski; 112 — I. andreevae (BG: Kresnensko Hanche, 19.04. 2006, CC); 113 — I. clara; 114 — I. miksici (BG: Gorski Dom Lodge, 24.06. 2006, CC); 115 — I. plevnensis (BG: Levishte, 14.07. 2009, CC); 116 — I. longicaudata adamovici (BG: Karandila, 24.06. 2008, CC); 117 — I. l. longicaudata (BG: Sofia University Botanical garden, 26.06. 1.07.2004, CC); 118 — I. m. modesta (= intermedia); 119 — I. rhodopensis leonorae (BG: Livade, 9.08. 2006, CC); 120 — I. rh. rhodopensis (BG: Bachkovo, 600 m, 23.05. 2004, CC); 121 — I. rh. petkovi (BG: Gluhite Kamani, 23.06. 2008, CC); 122 — I. modestior (BG: Bankya, 24.05. 2006, CC); 123 — I. dobrogensis; 124 — I. zubowskii; 125 — I. aff. camptoxypha (data as in Fig. 75); 126 — I. gulae; 127 — I. obtusa (BG: Ravnets Ridge, 14.07. 2009, CC); 128 — I. amplipennis (as in Fig. 78); 129 — I. speciosa (BG: Bankya, 24.05. 2006, CC). Indications on Fig. 127 show, respectively: G—gonangulum; L—lamella; P—lateral pit between lamella and gonangulum. Scale (if present) = 5 mm." httpUri="https://zenodo.org/record/246556/files/figure.png" pageId="39" pageNumber="40">123</figureCitation>
,
<figureCitation id="E2FD9CBB392FFFB5B0C60D49FDDE9F17" box="[481,525,1866,1890]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="39" pageNumber="40">174</figureCitation>
,
<figureCitation id="E2FD9CBB392FFFB5B33E0D49FD969F17" box="[537,581,1866,1890]" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="39" pageNumber="40">184</figureCitation>
,
<figureCitation id="E2FD9CBB392FFFB5B3760D49FDAF9F17" box="[593,636,1866,1890]" captionStart="FIGURE 194" captionStartId="49.[151,250,1945,1968]" captionTargetBox="[151,1434,1141,1924]" captionTargetId="figure@49.[151,1436,1141,1924]" captionTargetPageId="49" captionText="FIGURE 194. Distribution of I. modestior, I. dobrogensis, I. zubowskii, I. aff. camptoxypha, I. gulae, and I. obtusa on the Balkan Peninsula." httpUri="https://zenodo.org/record/246571/files/figure.png" pageId="39" pageNumber="40">194</figureCitation>
)
</paragraph>
<paragraph id="7A79803E392FFFB5B1B00D91FD9A9FDC" blockId="39.[151,585,1938,1961]" box="[151,585,1938,1961]" pageId="39" pageNumber="40">
<taxonomicName id="BDC6FBBD392FFFB5B1B00D91FE469FDD" authority="Kis" authorityName="Kis" box="[151,405,1938,1960]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="dobrogensis">
<emphasis id="48B25C2C392FFFB5B1B00D91FEBB9FDD" box="[151,360,1938,1960]" italics="true" pageId="39" pageNumber="40">Isophya dobrogensis</emphasis>
Kis
</taxonomicName>
: Kis 1994 (sp.n.).
</paragraph>
</subSubSection>
<subSubSection id="32DCD3B5392FFFBAB1B00DD6FB9F9B42" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
<paragraph id="7A79803E392FFFB5B1B00DD6FC899F98" blockId="39.[151,858,2005,2030]" box="[151,858,2005,2030]" pageId="39" pageNumber="40">
<emphasis id="48B25C2C392FFFB5B1B00DD6FE179F98" box="[151,452,2005,2029]" italics="true" pageId="39" pageNumber="40">Morphological description</emphasis>
: Kis 1994.
<emphasis id="48B25C2C392FFFB5B3610DD5FD039F98" box="[582,720,2006,2029]" italics="true" pageId="39" pageNumber="40">Biacoustics:</emphasis>
Iorgu 2012.
</paragraph>
<paragraph id="7A79803E3920FFBAB1E00A94FCE898DA" blockId="40.[151,1436,151,895]" box="[199,827,151,175]" pageId="40" pageNumber="41">The species is well characterised in the cited references.</paragraph>
<paragraph id="7A79803E3920FFBAB1E00ABFFEB69981" blockId="40.[151,1436,151,895]" pageId="40" pageNumber="41">
<emphasis id="48B25C2C3920FFBAB1E00ABFFD1998A1" box="[199,714,188,212]" italics="true" pageId="40" pageNumber="41">Supplement to the description and diagnosis</emphasis>
: Fastigium verticis is about 1/2 of the width of scapus. Male tegmina are slightly shorter than pronotum, with right anal tegminal angle. Their Costo-Medial parts are short (
<figureCitation id="E2FD9CBB3920FFBAB4490ADCFF649969" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="40" pageNumber="41">Fig. 49</figureCitation>
). Male stridulatory file (
<figureCitation id="E2FD9CBB3920FFBAB0E10B07FDF79969" box="[454,548,260,284]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="40" pageNumber="41">Fig. 174</figureCitation>
A) is
<quantity id="BD3E2DDB3920FFBAB3450B07FD009969" box="[610,723,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.6" metricValueMin="4.0" pageId="40" pageNumber="41" unit="mm" value="4.3" valueMax="4.6" valueMin="4.0">44.6 mm</quantity>
long and bears 261275 teeth. Male cercus (
<figureCitation id="E2FD9CBB3920FFBAB5E10B07FAC49969" box="[1222,1303,260,284]" captionStart="FIGURES 80 104" captionStartId="14.[151,264,1835,1858]" captionTargetBox="[226,1360,681,1815]" captionTargetId="figure@14.[226,1362,681,1816]" captionTargetPageId="14" captionText="FIGURES 80 104. Male abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55): 80 — I. hospodar; 81 — I. rectipennis; 82 — I. pavelii; 83 — I. thracica; 84 — I. bureschi; 85 — I. yaraligozi; 86 — I. tosevski; 87 — I. andreevae; 88 — I. clara; 89 — I. miksici; 90 — I. plevnensis; 91 — I. longicaudata adamovici; 92 — I. l. longicaudata; 93 — I. m. modesta (= intermedia); 94 — I. rhodopensis leonorae; 95 — I. rh. rhodopensis; 96 — I. rh. petkovi; 97 — I. modestior; 98 — I. dobrogensis; 99 — I. zubowskii; 100 — I. aff. camptoxypha; 101 — I. gulae; 102 — I. obtusa; 103 — I. amplipennis; 104 — I. speciosa. Scale (if present) = 2 mm." httpUri="https://zenodo.org/record/246555/files/figure.png" pageId="40" pageNumber="41">Fig. 98</figureCitation>
) is strongly incurved in the apical 1/3, with a distinct terminal tooth. Female tegmen is about 1/3 of the pronotum length (
<figureCitation id="E2FD9CBB3920FFBAB4490B24FF649911" captionStart="FIGURES 56 79" captionStartId="13.[151,264,1769,1792]" captionTargetBox="[178,1421,662,1756]" captionTargetId="figure@13.[178,1422,662,1756]" captionTargetPageId="13" captionText="FIGURES 56 79. Morphology of female head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55 unless specifically given): 56 — I. hospodar; 57 — I. rectipennis; 58 — I. pavelii (paratype of I. rammei Peshev); 59 — I. thracica; 60 — I. bureschi; 61 — I. yaraligozi; 62 — I. tosevski; 63 — I. andreevae (paratype); 64 — I. clara; 65 — I. miksici (paratype); 66 — I. plevnensis (paratype); 67 — I. longicaudata adamovici (paratype, BG: Sliven, 1000 m, NMNHS); 68 — I. l. longicaudata; 69 — I. rhodopensis leonorae (BG: “ Alibotush, 1500 m ”, 13.07. 1959, NMNHS); 70 — I. rh. rhodopensis (as in Fig. 46 but 17.07.1956); 71 — I. rh. petkovi; 72 — I. modestior; 73 — I. dobrogensis; 74 — I. zubowskii; 75 — I. aff. camptoxypha (MK: Bistra Mt., 2.08. 2004, CC); 76 — I. gulae (paratype, BG: Dolna Topchiya, 10.07. 1974, NMNHS); 77 — I. obtusa (BG: Vezhen Peak, 15.07. 1969, NMNHS); 78 — I. amplipennis (TR: “ Turkey | 1960 | Sureya Bey ”, NHM); 79 — I. speciosa (BG: Sakar Mt., 14.06. 1952, NMNHS). Scale (if present; black line right of the specimen) = 10 mm." httpUri="https://zenodo.org/record/246554/files/figure.png" pageId="40" pageNumber="41">Fig. 73</figureCitation>
). Female stridulatory apparatus is shown in
<figureCitation id="E2FD9CBB3920FFBAB3820B4FFCD69911" box="[677,773,332,356]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="40" pageNumber="41">Fig. 174</figureCitation>
C. Ovipositor is long—
<quantity id="BD3E2DDB3920FFBAB5280B4FFB519911" box="[1039,1154,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="40" pageNumber="41" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">1113 mm</quantity>
(
<figureCitation id="E2FD9CBB3920FFBAB5B60B4FFB359911" box="[1169,1254,332,356]" captionStart="FIGURES 80 104" captionStartId="14.[151,264,1835,1858]" captionTargetBox="[226,1360,681,1815]" captionTargetId="figure@14.[226,1362,681,1816]" captionTargetPageId="14" captionText="FIGURES 80 104. Male abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55): 80 — I. hospodar; 81 — I. rectipennis; 82 — I. pavelii; 83 — I. thracica; 84 — I. bureschi; 85 — I. yaraligozi; 86 — I. tosevski; 87 — I. andreevae; 88 — I. clara; 89 — I. miksici; 90 — I. plevnensis; 91 — I. longicaudata adamovici; 92 — I. l. longicaudata; 93 — I. m. modesta (= intermedia); 94 — I. rhodopensis leonorae; 95 — I. rh. rhodopensis; 96 — I. rh. petkovi; 97 — I. modestior; 98 — I. dobrogensis; 99 — I. zubowskii; 100 — I. aff. camptoxypha; 101 — I. gulae; 102 — I. obtusa; 103 — I. amplipennis; 104 — I. speciosa. Scale (if present) = 2 mm." httpUri="https://zenodo.org/record/246555/files/figure.png" pageId="40" pageNumber="41">Fig. 97</figureCitation>
). Colouration in both sexes is variegated—green, with yellow, orange, reddish or violet lateral body stripes, antennae and tibiae. The disc of tegmen, CuA and CuP veins are brownish, reddish or green. Lateral parts of tegmina are banded with white. The species calling song (
<figureCitation id="E2FD9CBB3920FFBAB33D0BB4FDAF99BA" box="[538,636,439,463]" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="40" pageNumber="41">Fig. 184</figureCitation>
) consists of groups of long elaborate syllables, similar in structure to these of
<taxonomicName id="BDC6FBBD3920FFBAB1D20BDEFE8E9981" box="[245,349,477,500]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="48B25C2C3920FFBAB1D20BDEFE8E9981" box="[245,349,477,500]" italics="true" pageId="40" pageNumber="41">I. costata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="7A79803E3920FFBAB1E00802FB9F9B42" blockId="40.[151,1436,151,895]" pageId="40" pageNumber="41">
<emphasis id="48B25C2C3920FFBAB1E00802FE859A6D" box="[199,342,513,536]" italics="true" pageId="40" pageNumber="41">Bioacoustics</emphasis>
: The song was recorded at 26°С. The grouped syllables were separated by intervals of 25 s. First (main) part of the syllable consists of a compact impulse series of 300420 ms with 60100 impulses and more or less constant impulse interval of average 46 ms. The second part of the syllable follows at an interval of 90230 ms and has quite a peculiar structure, consisting of 36 small groups of impulses, separated by intervals of 5090 ms; each group includes 26 impulses with impulse period of 79 ms. The total duration of the syllables was 730 1100 ms. Very interesting in this species is the male-female duet before mating, with one of the longest and more elaborate known female answers to male song in this genus. Female song occurs after the second part of male syllable and consists of variable series of 927 impulses, recorded for a period of 48 s. It begins with a group of 8 13 impulses, lasting 301596 ms and is followed by several widely spaced impulses.
</paragraph>
</subSubSection>
<subSubSection id="32DCD3B53920FFBAB1E00947FED19F52" pageId="40" pageNumber="41" type="distribution">
<paragraph id="7A79803E3920FFBAB1E00947FB849B0A" blockId="40.[151,1436,151,895]" pageId="40" pageNumber="41">
<emphasis id="48B25C2C3920FFBAB1E00947FD569B29" box="[199,645,836,860]" italics="true" pageId="40" pageNumber="41">
Distribution (
<figureCitation id="E2FD9CBB3920FFBAB0430947FE1B9B29" box="[356,456,836,860]" captionStart="FIGURE 194" captionStartId="49.[151,250,1945,1968]" captionTargetBox="[151,1434,1141,1924]" captionTargetId="figure@49.[151,1436,1141,1924]" captionTargetPageId="49" captionText="FIGURE 194. Distribution of I. modestior, I. dobrogensis, I. zubowskii, I. aff. camptoxypha, I. gulae, and I. obtusa on the Balkan Peninsula." httpUri="https://zenodo.org/record/246571/files/figure.png" pageId="40" pageNumber="41">Fig. 194</figureCitation>
) and phenology
</emphasis>
:
<taxonomicName id="BDC6FBBD3920FFBAB3BF0947FC579B29" box="[664,900,836,860]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="dobrogensis">
<emphasis id="48B25C2C3920FFBAB3BF0947FC579B29" box="[664,900,836,860]" italics="true" pageId="40" pageNumber="41">Isophya dobrogensis</emphasis>
</taxonomicName>
is a local endemite known only from Popina Island in Razelm Lake (SE
<collectingCountry id="02D1C0AE3920FFBAB0EB0964FDE59B0A" box="[460,566,871,895]" name="Romania" pageId="40" pageNumber="41">Romania</collectingCountry>
), at
<quantity id="BD3E2DDB3920FFBAB3440964FD6B9B0A" box="[611,696,871,895]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="3.0" metricValueMin="0.1" pageId="40" pageNumber="41" unit="m" value="15.5" valueMax="30.0" valueMin="1.0">130 m</quantity>
alt. Nymphs—IVV, imago—VVI.
</paragraph>
<paragraph id="7A79803E3920FFBAB1B009B3FDCA9BBD" blockId="40.[151,1437,943,1831]" box="[151,537,943,968]" pageId="40" pageNumber="41">
<emphasis id="48B25C2C3920FFBAB1B009B3FDCA9BBD" bold="true" box="[151,537,943,968]" pageId="40" pageNumber="41">
5.
<taxonomicName id="BDC6FBBD3920FFBAB19409ACFEA29BBD" box="[179,369,943,968]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="kraussii">
<emphasis id="48B25C2C3920FFBAB19409ACFEA29BBD" bold="true" box="[179,369,943,968]" italics="true" pageId="40" pageNumber="41">Isophya kraussii</emphasis>
</taxonomicName>
species group
</emphasis>
</paragraph>
<paragraph id="7A79803E3920FFBAB1B009D7FB849C2D" blockId="40.[151,1437,943,1831]" pageId="40" pageNumber="41">
This group shows similar characters to the
<taxonomicName id="BDC6FBBD3920FFBAB3B909D6FCDF9B99" box="[670,780,981,1004]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="48B25C2C3920FFBAB3B909D6FCDF9B99" box="[670,780,981,1004]" italics="true" pageId="40" pageNumber="41">I. costata</emphasis>
</taxonomicName>
group but we regard it separately on account of some differences in the song structure, smaller body size, short ovipositor, and different morphology of the sex chromosome. However, it is possible in future, when new acoustic, molecular, and karyological data is accumulated, to reconsider this group as well as the
<taxonomicName id="BDC6FBBD3920FFBAB3F80E42FC9B9C2D" box="[735,840,1089,1112]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="48B25C2C3920FFBAB3F80E42FC9B9C2D" box="[735,840,1089,1112]" italics="true" pageId="40" pageNumber="41">I. costata</emphasis>
</taxonomicName>
and
<taxonomicName id="BDC6FBBD3920FFBAB2590E42FC2F9C2D" box="[894,1020,1089,1112]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pyrenaea">
<emphasis id="48B25C2C3920FFBAB2590E42FC2F9C2D" box="[894,1020,1089,1112]" italics="true" pageId="40" pageNumber="41">I. pyrenaea</emphasis>
</taxonomicName>
groups.
</paragraph>
<paragraph id="7A79803E3920FFBAB1E00E67FED19F52" blockId="40.[151,1437,943,1831]" pageId="40" pageNumber="41">
Species of moderate size.
<collectingCountry id="02D1C0AE3920FFBAB0C80E67FDF49C09" box="[495,551,1124,1148]" name="India" pageId="40" pageNumber="41">Hind</collectingCountry>
femur is shorter than
<quantity id="BD3E2DDB3920FFBAB2030E67FCA49C09" box="[804,887,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="40" pageNumber="41" unit="mm" value="18.0">18 mm</quantity>
. Fastigium verticis is narrower than scapus (less than 1/2 of its width). The pronotal disc is long; the side keels of metazone are sinuately diverging. Male tegmina are distinctly shortened, usually shorter than pronotum. CuP is very long (about 4/5 times of the hind margin of metazone or longer), moderately widened, slightly bulged over the pronotal surface; CuP and CuA veins are very closely approximated as a result of bending the area between them and uplifting of CuA over the tegminal surface. In comparison to
<taxonomicName id="BDC6FBBD3920FFBAB0450F1AFE1F9D45" box="[354,460,1305,1328]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="48B25C2C3920FFBAB0450F1AFE1F9D45" box="[354,460,1305,1328]" italics="true" pageId="40" pageNumber="41">I. costata</emphasis>
</taxonomicName>
group CuA is situated higher than CuP. The stridulatory file has a very high number (180305) of fine dense teeth. Female tegmina have truncate or concave apical margins and reticulate venation. The lower keels of the hind femur lack spines. Male cerci are moderately stout to moderately slender; their apical part is gradually incurved, rounded or more or less pointed. The cercal tooth is apically positioned, small, slender, pointed. The female ovipositor is short (usually
<quantity id="BD3E2DDB3920FFBAB38A0FA4FCCB9DB5" box="[685,792,1447,1472]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="40" pageNumber="41" unit="mm" value="9.5" valueMax="12.0" valueMin="7.0">712 mm</quantity>
to
<quantity id="BD3E2DDB3920FFBAB21D0FA4FC5B9DB5" box="[826,904,1447,1472]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="40" pageNumber="41" unit="mm" value="14.0">14 mm</quantity>
in
<taxonomicName id="BDC6FBBD3920FFBAB28D0FAAFBFC9DB5" box="[938,1071,1448,1472]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="zubowskii">
<emphasis id="48B25C2C3920FFBAB28D0FAAFBFC9DB5" box="[938,1071,1448,1472]" italics="true" pageId="40" pageNumber="41">I. zubowskii</emphasis>
</taxonomicName>
). The body colouration is green; melanism is always absent. The disc of tegmina is green or brownish with dark stridulatory area. The lateral margins of pronotum in metazone have reddish stripe situated above the light band. The song consists of groups or long sequences of syllables lasting 120400 ms. The X-chromosome is submetacentric (
<typeStatus id="A57D3E9C3920FFBAB5950C17FB379E59" box="[1202,1252,1556,1580]" pageId="40" pageNumber="41">type</typeStatus>
3 according to Warchałowska-Śliwa
<emphasis id="48B25C2C3920FFBAB0AB0C3AFE169E25" box="[396,453,1592,1616]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
2008), similar to that observed within the
<taxonomicName id="BDC6FBBD3920FFBAB2820C3AFBF79E25" box="[933,1060,1593,1616]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pyrenaea">
<emphasis id="48B25C2C3920FFBAB2820C3AFBF79E25" box="[933,1060,1593,1616]" italics="true" pageId="40" pageNumber="41">I. pyrenaea</emphasis>
</taxonomicName>
group. We regard the group with four species (seven taxa)—
<emphasis id="48B25C2C3920FFBAB0EB0C5EFC919E01" box="[460,834,1628,1652]" italics="true" pageId="40" pageNumber="41">
<taxonomicName id="BDC6FBBD3920FFBAB0EB0C5EFDB79E01" box="[460,612,1628,1652]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="brevicauda">I. brevicauda</taxonomicName>
,
<taxonomicName id="BDC6FBBD3920FFBAB3570C5EFC919E01" authority="Brunner" authorityName="Brunner" box="[624,834,1628,1652]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="kraussii" subSpecies="kraussii">I. kraussii kraussii</taxonomicName>
</emphasis>
Brunner von Wattenwyl, 1878,
<taxonomicName id="BDC6FBBD3920FFBAB5890C5EFE579EE2" authority="Iorgu &amp; Heller, 2013" authorityName="Iorgu &amp; Heller" authorityYear="2013" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="kraussii" subSpecies="moldavica">
<emphasis id="48B25C2C3920FFBAB5890C5EFA489E01" box="[1198,1435,1628,1652]" italics="true" pageId="40" pageNumber="41">I. kraussii moldavica</emphasis>
Iorgu &amp; Heller, 2013
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3920FFBAB0B50C82FCCE9EE2" authority="Maran, 1954" authorityName="Maran" authorityYear="1954" box="[402,797,1663,1688]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="pienensis" subSpecies="pienensis">
<emphasis id="48B25C2C3920FFBAB0B50C82FD519EED" box="[402,642,1665,1688]" italics="true" pageId="40" pageNumber="41">I. pienensis pienensis</emphasis>
Mařan, 1954
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3920FFBAB20E0C82FADE9EE2" authority="Chladek, 2010" authorityName="Chladek" authorityYear="2010" box="[809,1293,1663,1688]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="pienensis" subSpecies="austromoravica">
<emphasis id="48B25C2C3920FFBAB20E0C82FBB29EED" box="[809,1121,1665,1688]" italics="true" pageId="40" pageNumber="41">I. pienensis austromoravica</emphasis>
Chládek, 2010
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3920FFBAB43B0C82FE709EC9" authority="Chladek, 2011" authorityName="Chladek" authorityYear="2011" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="pienensis" subSpecies="sudetica">
<emphasis id="48B25C2C3920FFBAB43B0C82FF279EC9" italics="true" pageId="40" pageNumber="41">I. pienensis sudetica</emphasis>
Chládek, 2011
</taxonomicName>
, and
<taxonomicName id="BDC6FBBD3920FFBAB0C10CA6FDBD9EC9" box="[486,622,1700,1724]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="zubowskii">
<emphasis id="48B25C2C3920FFBAB0C10CA6FDBD9EC9" box="[486,622,1700,1724]" italics="true" pageId="40" pageNumber="41">I. zubowskii</emphasis>
</taxonomicName>
, though further reconsideration of the groups constitution is possible. The range of the group covers the northeasternmost part of the Balkan Peninsula, Central Europe and the Carpathian basin (incl.
<collectingCountry id="02D1C0AE3920FFBAB0930CEFFDDA9F71" box="[436,521,1772,1796]" name="Croatia" pageId="40" pageNumber="41">Croatia</collectingCountry>
,
<collectingCountry id="02D1C0AE3920FFBAB33B0CEFFD529F71" box="[540,641,1772,1796]" name="Slovenia" pageId="40" pageNumber="41">Slovenia</collectingCountry>
,
<collectingCountry id="02D1C0AE3920FFBAB3B30CEFFD3A9F71" box="[660,745,1772,1796]" name="Austria" pageId="40" pageNumber="41">Austria</collectingCountry>
,
<collectingCountry id="02D1C0AE3920FFBAB3DB0CEEFC8C9F70" box="[764,863,1773,1797]" name="Hungary" pageId="40" pageNumber="41">Hungary</collectingCountry>
,
<collectingCountry id="02D1C0AE3920FFBAB2550CEFFC049F71" box="[882,983,1772,1796]" name="Slovakia" pageId="40" pageNumber="41">Slovakia</collectingCountry>
, S
<collectingCountry id="02D1C0AE3920FFBAB52F0CEFFBA19F71" box="[1032,1138,1772,1796]" name="Germany" pageId="40" pageNumber="41">Germany</collectingCountry>
, S
<collectingCountry id="02D1C0AE3920FFBAB5840CEFFB279F71" box="[1187,1268,1772,1796]" name="Poland" pageId="40" pageNumber="41">Poland</collectingCountry>
,
<collectingCountry id="02D1C0AE3920FFBAB4210CEFFABD9F71" box="[1286,1390,1772,1796]" name="Romania" pageId="40" pageNumber="41">Romania</collectingCountry>
, W
<collectingCountry id="02D1C0AE3920FFBAB1B00D0CFF249F52" box="[151,247,1807,1831]" name="Ukraine" pageId="40" pageNumber="41">Ukraine</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>