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<document id="CA7D06F657247F3CDF2B71C315AC3514" ID-DOI="10.1080/00222933.2019.1596328" ID-GBIF-Dataset="31e9dec8-ac91-43e6-aef3-c43fb0ed82ad" ID-ISSN="1464-5262" ID-Zenodo-Dep="3675703" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1582134628034" checkinUser="valdenar" docAuthor="Molina-Acevedo, Isabel C." docDate="2019" docId="EB1A9B52FFB3FFB371FB54CAFD4BFDA5" docLanguage="en" docName="JNATHIST.53.910.517540.pdf.imf" docOrigin="Journal of Natural History 53 (9)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Treadwellphysa villalobosi Molina-Acevedo 2019, sp. nov." docType="treatment" docVersion="10" lastPageNumber="538" masterDocId="1723E32AFFA1FFA470405151FF2DFFA4" masterDocTitle="Redescription of two species previously regarded as Marphysa de Quatrefagesı 1865 ı with the description of new species of Treadwellphysa Molina-Acevedo and Carrera-Parraı 2017" masterLastPageNumber="540" masterPageNumber="517" pageNumber="534" updateTime="1698720035678" updateUser="plazi">
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<mods:title id="F43E4A7355A09E39AA0AF3DD906A6E7D">Redescription of two species previously regarded as Marphysa de Quatrefagesı 1865 ı with the description of new species of Treadwellphysa Molina-Acevedo and Carrera-Parraı 2017</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="72D850D41AD8A6B0F71677295A8E1A68">Author</mods:roleTerm>
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<mods:namePart id="7286AD9C3D44105E6A55125AAABFB555">Molina-Acevedo, Isabel C.</mods:namePart>
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<mods:typeOfResource id="055309CB690E816D35E60096BE1D5636">text</mods:typeOfResource>
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<mods:title id="0CC38EB2F03E80D66C81DFCC16A8ED80">Journal of Natural History</mods:title>
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<mods:part id="E162586F10E746915B5D597B16E44A46">
<mods:date id="C1C126F9A2591DCF9F734CF020A5FF69">2019</mods:date>
<mods:detail id="DD0B6479EAC733EC91FF00844AE02FC3" type="pubDate">
<mods:number id="D3C295B2E01CA5676BCDC3BB81F508CC">2019-04-30</mods:number>
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<mods:detail id="4E9C8AD1B8A4EC98CD211B086ABA1B31" type="volume">
<mods:number id="BB9320EFC4FF6F38129FAF72FAD4B529">53</mods:number>
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<mods:number id="65E25636D5076B144A10BCE5FD906F60">9</mods:number>
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<mods:classification id="E7011A16C0C066935C47F3F9C60FC559">journal article</mods:classification>
<mods:identifier id="74D2B81A31FD7277986EAC5BA23E3132" type="DOI">10.1080/00222933.2019.1596328</mods:identifier>
<mods:identifier id="EADC5CEAFB113B21D51ED0E008C10E63" type="GBIF-Dataset">31e9dec8-ac91-43e6-aef3-c43fb0ed82ad</mods:identifier>
<mods:identifier id="17F5E3014991300FDC74E75291AA1EEC" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="3566A508969865BAC41ED75E6155E152" type="Zenodo-Dep">3675703</mods:identifier>
<mods:identifier id="60174A8DE937F976DA7311E30168D3DD" type="ZooBank">693D9279-682F-44E1-B1D5-15A9A5029FB6</mods:identifier>
</mods:mods>
<treatment id="EB1A9B52FFB3FFB371FB54CAFD4BFDA5" ID-DOI="http://doi.org/10.5281/zenodo.3680430" ID-GBIF-Taxon="162173994" ID-Zenodo-Dep="3680430" LSID="urn:lsid:plazi:treatment:EB1A9B52FFB3FFB371FB54CAFD4BFDA5" httpUri="http://treatment.plazi.org/id/EB1A9B52FFB3FFB371FB54CAFD4BFDA5" lastPageId="23" lastPageNumber="538" pageId="18" pageNumber="534">
<subSubSection id="2BA979CFFFB3FFB671FB54CAFC44FA10" box="[443,873,1435,1460]" pageId="18" pageNumber="534" type="nomenclature">
<paragraph id="630C2A44FFB3FFB671FB54CAFC44FA10" blockId="18.[443,873,1435,1495]" box="[443,873,1435,1460]" pageId="18" pageNumber="534">
<heading id="38449D28FFB3FFB671FB54CAFC44FA10" box="[443,873,1435,1460]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="534" reason="2">
<taxonomicName id="A4B351C7FFB3FFB671FB54CAFC2EFA10" ID-CoL="85S5T" authority="Molina-Acevedo, 2019" authorityName="Molina-Acevedo" authorityYear="2019" box="[443,771,1435,1460]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="534" phylum="Annelida" rank="species" species="villalobosi" status="sp. nov.">
<emphasis id="51C7F656FFB3FFB671FB54CAFC2EFA10" bold="true" box="[443,771,1435,1460]" italics="true" pageId="18" pageNumber="534">Treadwellphysa villalobosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFB3FFB6734C54CAFC44FA10" box="[780,873,1435,1460]" pageId="18" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2BA979CFFFB3FFB6720554EFFDF3FA73" box="[581,734,1470,1495]" pageId="18" pageNumber="534" type="description">
<paragraph id="630C2A44FFB3FFB6720554EFFDF3FA73" blockId="18.[443,873,1435,1495]" box="[581,734,1470,1495]" pageId="18" pageNumber="534">
<heading id="38449D28FFB3FFB6720554EFFDF3FA73" box="[581,734,1470,1495]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="534" reason="5">
(
<figureCitation id="FB8836C1FFB3FFB6720D54EFFDFBFA73" box="[589,726,1470,1495]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStartId-0="19.[160,225,1408,1431]" captionStartId-1="21.[160,225,1512,1535]" captionTargetBox-0="[164,1152,143,1357]" captionTargetBox-1="[164,1152,143,1480]" captionTargetId-0="figure@19.[163,1153,142,1376]" captionTargetId-1="figure@21.[163,1153,142,1481]" captionTargetPageId-0="19" captionTargetPageId-1="21" captionText-0="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." captionText-1="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi-0="http://doi.org/10.5281/zenodo.3675720" figureDoi-1="http://doi.org/10.5281/zenodo.3675722" httpUri-0="https://zenodo.org/record/3675720/files/figure.png" httpUri-1="https://zenodo.org/record/3675722/files/figure.png" pageId="18" pageNumber="534">
Figures 8
<emphasis id="51C7F656FFB3FFB672FA54EFFDE5FA73" box="[698,712,1470,1495]" italics="true" pageId="18" pageNumber="534"></emphasis>
9
</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2BA979CFFFB3FFB770E0574DFCA8F9E6" lastPageId="19" lastPageNumber="535" pageId="18" pageNumber="534" type="materials_examined">
<paragraph id="630C2A44FFB3FFB670E0574DFEADF991" blockId="18.[160,1156,1564,1727]" box="[160,384,1564,1589]" pageId="18" pageNumber="534">
<emphasis id="51C7F656FFB3FFB670E0574DFEADF991" bold="true" box="[160,384,1564,1589]" italics="true" pageId="18" pageNumber="534">Material examined</emphasis>
</paragraph>
<paragraph id="630C2A44FFB3FFB670E0576FFE26F91B" blockId="18.[160,1156,1564,1727]" pageId="18" pageNumber="534">
<emphasis id="51C7F656FFB3FFB670E0576FFE60F9F3" bold="true" box="[160,333,1598,1623]" italics="true" pageId="18" pageNumber="534">Type material.</emphasis>
<materialsCitation id="D3DB2019FFB3FFB6711C576EFE57F939" ID-GBIF-Occurrence="2569345062" collectingDate="2013-11-10" collectionCode="T" collectorName="F. Villalobos-Guerrero" county="Blancoi Cabuyai Punta Arenasi Costa" elevation="1" location="Punta Arenasi Costa Ricai Pacific" municipality="Cabuyai Punta Arenasi Costa Ricai" pageId="18" pageNumber="534" specimenCode="ECOSUR206" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<typeStatus id="BC0894E6FFB3FFB6711C576EFEEEF9F3" box="[348,451,1599,1623]" pageId="18" pageNumber="534" type="holotype">Holotype</typeStatus>
<specimenCode id="3315823FFFB3FFB6718D576EFD79F9F3" box="[461,596,1599,1623]" collectionCode="ECOSUR" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="18" pageNumber="534" type="Museum">ECOSUR206</specimenCode>
;
<collectingRegion id="A177E4A6FFB3FFB67221576EFDB6F9F3" box="[609,667,1599,1623]" country="South Africa" name="Western Cape" pageId="18" pageNumber="534">Cabo</collectingRegion>
<collectingCounty id="8A6D52C8FFB3FFB672E5576EFB6BF9F3" box="[677,1094,1599,1623]" pageId="18" pageNumber="534">
Blanco,
<collectingMunicipality id="8368B03EFFB3FFB672BF576EFB6BF9F3" box="[767,1094,1599,1623]" pageId="18" pageNumber="534">
Cabuya,
<location id="666C7C9FFFB3FFB67322576EFB6BF9F3" LSID="urn:lsid:plazi:treatment:EB1A9B52FFB3FFB371FB54CAFD4BFDA5:666C7C9FFFB3FFB67322576EFB6BF9F3" box="[866,1094,1599,1623]" county="Blancoi Cabuyai Punta Arenasi Costa" municipality="Cabuyai Punta Arenasi Costa Ricai" name="Punta Arenasi Costa Ricai Pacific" pageId="18" pageNumber="534" stateProvince="Western Cape">
Punta
<location id="666C7C9FFFB3FFB673ED576EFB6BF9F3" LSID="urn:lsid:plazi:treatment:EB1A9B52FFB3FFB371FB54CAFD4BFDA5:666C7C9FFFB3FFB673ED576EFB6BF9F3" box="[941,1094,1599,1623]" county="Blancoi Cabuyai Punta Arenasi Costa" municipality="Cabuyai Punta Arenasi Costa Ricai" name="Arenasi Costa Ricai Pacific Ocean" pageId="18" pageNumber="534" stateProvince="Western Cape">
Arenas,
<collectingCountry id="1BA46AD4FFB3FFB67448576EFB6BF9F3" box="[1032,1094,1599,1623]" name="Costa Rica" pageId="18" pageNumber="534">Costa</collectingCountry>
</location>
</location>
</collectingMunicipality>
</collectingCounty>
Rica, Pacific Ocean;
<geoCoordinate id="06874C83FFB3FFB6710A5733FE8CF9DE" box="[330,417,1634,1658]" degrees="9.586" direction="north" orientation="latitude" pageId="18" pageNumber="534" precision="55" value="9.586">9.586°N</geoCoordinate>
-
<geoCoordinate id="06874C83FFB3FFB671F35733FD0CF9DE" box="[435,545,1634,1658]" degrees="85.089" direction="west" orientation="longitude" pageId="18" pageNumber="534" precision="55" value="-85.089">85.089°W</geoCoordinate>
;
<date id="170D0C84FFB3FFB672705733FC3DF9DE" box="[560,784,1634,1658]" pageId="18" pageNumber="534" value="2013-11-10">
<collectingDate id="0749F56CFFB3FFB672705733FC3DF9DE" box="[560,784,1634,1658]" pageId="18" pageNumber="534" value="2013-11-10">10 November 2013</collectingDate>
</date>
; in volcanic rock;
<quantity id="A44B87A1FFB3FFB673A65733FB3AF9DE" box="[998,1047,1634,1658]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="534" unit="m" value="1.0">
<elevation id="E89ECD77FFB3FFB673A65733FB3AF9DE" box="[998,1047,1634,1658]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="534" unit="m" value="1.0">1 m</elevation>
</quantity>
; coll.
<collectionCode id="05A2B281FFB3FFB6741E5733FB42F9DE" box="[1118,1135,1634,1658]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="534">T</collectionCode>
.
<collectorName id="CE464F92FFB3FFB674325733FE57F939" pageId="18" pageNumber="534">F. Villalobos-Guerrero</collectorName>
</materialsCitation>
.
<materialsCitation id="D3DB2019FFB3FFB671C557D4FE26F91B" ID-GBIF-Occurrence="2570076583" pageId="18" pageNumber="534" typeStatus="paratype">
<typeStatus id="BC0894E6FFB3FFB671C557D4FEDFF939" box="[389,498,1669,1693]" pageId="18" pageNumber="534" type="paratype">Paratypes</typeStatus>
<specimenCode id="3315823FFFB3FFB671BA57D4FD53F939" box="[506,638,1669,1693]" pageId="18" pageNumber="534">
<collectionCode id="05A2B281FFB3FFB671BA57D4FD77F939" box="[506,602,1669,1693]" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="18" pageNumber="534" type="Museum">ECOSUR</collectionCode>
207
</specimenCode>
(1),
<collectionCode id="05A2B281FFB3FFB672F057D4FC04F939" box="[688,809,1669,1693]" pageId="18" pageNumber="534">LACM-AHF</collectionCode>
<specimenCode id="3315823FFFB3FFB6737157D4FCA0F939" box="[817,909,1669,1693]" pageId="18" pageNumber="534">
<collectionCode id="05A2B281FFB3FFB6737157D4FC40F939" box="[817,877,1669,1693]" pageId="18" pageNumber="534">POLY</collectionCode>
11
</specimenCode>
,124 (1); same data as
<typeStatus id="BC0894E6FFB3FFB670E057F6FE2AF91B" box="[160,263,1703,1727]" pageId="18" pageNumber="534" type="holotype">holotype</typeStatus>
.
</materialsCitation>
</paragraph>
<caption id="37CC7ACCFFB2FFB770E054D1FB0CFA4B" ID-DOI="http://doi.org/10.5281/zenodo.3675720" ID-Zenodo-Dep="3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="19" pageNumber="535" startId="19.[160,225,1408,1431]" targetBox="[164,1152,143,1357]" targetPageId="19">
<paragraph id="630C2A44FFB2FFB770E054D1FB0CFA4B" blockId="19.[160,1156,1408,1519]" pageId="19" pageNumber="535">
<emphasis id="51C7F656FFB2FFB770E054D1FFD6FA33" bold="true" box="[160,251,1408,1431]" pageId="19" pageNumber="535">Figure 8.</emphasis>
<taxonomicName id="A4B351C7FFB2FFB7714354D1FED3FA33" authority="Molina-Acevedo, 2019" authorityName="Molina-Acevedo" authorityYear="2019" box="[259,510,1408,1431]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="19" pageNumber="535" phylum="Annelida" rank="species" species="villalobosi" status="sp. nov.">
<emphasis id="51C7F656FFB2FFB7714354D1FED3FA33" box="[259,510,1408,1431]" italics="true" pageId="19" pageNumber="535">Treadwellphysa villalobosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFB2FFB7724554D1FD7BFA33" box="[517,598,1408,1431]" pageId="19" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm.
</paragraph>
</caption>
<paragraph id="630C2A44FFB2FFB770E05778FCA8F9E6" blockId="19.[160,901,1577,1602]" box="[160,901,1577,1602]" pageId="19" pageNumber="535">
<emphasis id="51C7F656FFB2FFB770E05778FEA2F9E6" bold="true" box="[160,399,1577,1602]" italics="true" pageId="19" pageNumber="535">Additional material.</emphasis>
ECOSUR-P2944 (2); same data as holotype.
</paragraph>
</subSubSection>
<subSubSection id="2BA979CFFFB5FFB270E051DFFC29FB58" lastPageId="22" lastPageNumber="538" pageId="20" pageNumber="536" type="description">
<paragraph id="630C2A44FFB5FFB070E051DFFE0BFF03" blockId="20.[160,1156,142,1727]" box="[160,294,142,167]" pageId="20" pageNumber="536">
<emphasis id="51C7F656FFB5FFB070E051DFFE0BFF03" bold="true" box="[160,294,142,167]" italics="true" pageId="20" pageNumber="536">Description</emphasis>
</paragraph>
<paragraph id="630C2A44FFB5FFB070E051E3FE6CFE96" blockId="20.[160,1156,142,1727]" pageId="20" pageNumber="536">
<typeStatus id="BC0894E6FFB5FFB070E051E3FE2AFF6E" box="[160,263,178,202]" pageId="20" pageNumber="536" type="holotype">Holotype</typeStatus>
complete, laterally dissected, with 417 chaetigers, L10 = 6 mm, W10 = 2.1 mm, TL =
<quantity id="A44B87A1FFB5FFB0709D5185FE12FF48" box="[221,319,212,236]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" pageId="20" pageNumber="536" unit="mm" value="135.0">135 mm</quantity>
. Anterior region of body with convex dorsum and flat ventrum, without groove; body depressed from chaetiger 12, widest at chaetiger 62, tapering after chaetiger 182.
</paragraph>
<paragraph id="630C2A44FFB5FFB07080506DFEFAFD80" blockId="20.[160,1156,142,1727]" pageId="20" pageNumber="536">
Prostomium bilobed,
<quantity id="A44B87A1FFB5FFB071FF506DFD34FEF0" box="[447,537,316,340]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="20" pageNumber="536" unit="mm" value="1.3">1.3 mm</quantity>
long,
<quantity id="A44B87A1FFB5FFB0722B506DFD9FFEF0" box="[619,690,316,340]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="536" unit="mm" value="2.0">2 mm</quantity>
wide; lobes frontally rounded; median sulcus shallow (
<figureCitation id="FB8836C1FFB5FFB07113500EFE9DFED3" box="[339,432,351,375]" captionStart="Figure 8" captionStartId="19.[160,225,1408,1431]" captionTargetBox="[164,1152,143,1357]" captionTargetId="figure@19.[163,1153,142,1376]" captionTargetPageId="19" captionText="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="20" pageNumber="536">Figure 8</figureCitation>
(a
<emphasis id="51C7F656FFB5FFB07184500EFEFCFED3" box="[452,465,351,375]" italics="true" pageId="20" pageNumber="536"></emphasis>
c)), ventral sulcus deep (
<figureCitation id="FB8836C1FFB5FFB072A8500EFC6AFED3" box="[744,839,351,375]" captionStart="Figure 8" captionStartId="19.[160,225,1408,1431]" captionTargetBox="[164,1152,143,1357]" captionTargetId="figure@19.[163,1153,142,1376]" captionTargetPageId="19" captionText="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="20" pageNumber="536">Figure 8</figureCitation>
(b)). Prostomial appendages in a semicircle, median antenna isolated by a gap. Palps reaching second peristomial ring; lateral antennae reaching middle of second chaetiger; median antenna reaching third chaetiger. Palpophores and ceratophores ring-shaped, short, slender; palpostyles and ceratostyles tapering, slender, without articulation. Eyes reniform, brown, between palps and lateral antennae.
</paragraph>
<paragraph id="630C2A44FFB5FFB07080537EFCB5FD28" blockId="20.[160,1156,142,1727]" pageId="20" pageNumber="536">
Peristomium of similar length than prostomium (
<quantity id="A44B87A1FFB5FFB072A4537EFC14FDE3" box="[740,825,559,583]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="20" pageNumber="536" unit="mm" value="1.3">1.3 mm</quantity>
long,
<quantity id="A44B87A1FFB5FFB073C0537EFCF9FDE3" box="[896,980,559,583]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="20" pageNumber="536" unit="mm" value="2.8">2.8 mm</quantity>
wide); first ring 2 times longer than second ring, separation between rings distinct dorsal and ventrally (
<figureCitation id="FB8836C1FFB5FFB070E85325FE28FD28" box="[168,261,628,652]" captionStart="Figure 8" captionStartId="19.[160,225,1408,1431]" captionTargetBox="[164,1152,143,1357]" captionTargetId="figure@19.[163,1153,142,1376]" captionTargetPageId="19" captionText="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="20" pageNumber="536">Figure 8</figureCitation>
(a
<emphasis id="51C7F656FFB5FFB071595325FE0BFD28" box="[281,294,628,652]" italics="true" pageId="20" pageNumber="536"></emphasis>
c)). Ventral lip without central depression (
<figureCitation id="FB8836C1FFB5FFB0734E5325FC40FD28" box="[782,877,628,652]" captionStart="Figure 8" captionStartId="19.[160,225,1408,1431]" captionTargetBox="[164,1152,143,1357]" captionTargetId="figure@19.[163,1153,142,1376]" captionTargetPageId="19" captionText="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="20" pageNumber="536">Figure 8</figureCitation>
(b)).
</paragraph>
<paragraph id="630C2A44FFB5FFB0708053C6FC39FBEB" blockId="20.[160,1156,142,1727]" pageId="20" pageNumber="536">
Maxillary apparatus with MF = 1+1, 3+3, 5+0, 4+5, 1+1 (
<figureCitation id="FB8836C1FFB5FFB073C053C6FCC8FD0B" box="[896,997,663,687]" captionStart="Figure 8" captionStartId="19.[160,225,1408,1431]" captionTargetBox="[164,1152,143,1357]" captionTargetId="figure@19.[163,1153,142,1376]" captionTargetPageId="19" captionText="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="20" pageNumber="536">Figure 8</figureCitation>
(d)). Maxillary carriers 2.8 times shorter than length of MI. MI forceps-like; closing system 3.4 times shorter than length of MI; ligament between MI and MII sclerotised (
<figureCitation id="FB8836C1FFB5FFB0738B538DFB06FD50" box="[971,1067,732,756]" captionStart="Figure 8" captionStartId="19.[160,225,1408,1431]" captionTargetBox="[164,1152,143,1357]" captionTargetId="figure@19.[163,1153,142,1376]" captionTargetPageId="19" captionText="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="20" pageNumber="536">Figure 8</figureCitation>
(d)). MII with recurved teeth; cavity opening oval, 2.9 times shorter than length of MII; ligament between MII and MIII and right MIV, sclerotised (
<figureCitation id="FB8836C1FFB5FFB073205273FCEFFC9E" box="[864,962,802,826]" captionStart="Figure 8" captionStartId="19.[160,225,1408,1431]" captionTargetBox="[164,1152,143,1357]" captionTargetId="figure@19.[163,1153,142,1376]" captionTargetPageId="19" captionText="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="20" pageNumber="536">Figure 8</figureCitation>
(d,e)). MIII short; with blunt teeth; with irregular attachment lamella, situated in centre of the right edge of maxilla, sclerotised (
<figureCitation id="FB8836C1FFB5FFB071BA5236FD70FCDB" box="[506,605,871,895]" captionStart="Figure 8" captionStartId="19.[160,225,1408,1431]" captionTargetBox="[164,1152,143,1357]" captionTargetId="figure@19.[163,1153,142,1376]" captionTargetPageId="19" captionText="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="20" pageNumber="536">Figure 8</figureCitation>
(d,e)). Left MIV with three larger lateral teeth; attachment lamella semicircular, wide, better developed on left side, situated along anterior edge of maxilla (
<figureCitation id="FB8836C1FFB5FFB071A152FDFD6AFC60" box="[481,583,940,964]" captionStart="Figure 8" captionStartId="19.[160,225,1408,1431]" captionTargetBox="[164,1152,143,1357]" captionTargetId="figure@19.[163,1153,142,1376]" captionTargetPageId="19" captionText="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="20" pageNumber="536">Figure 8</figureCitation>
(d,e)). Right MIV with two larger lateral teeth; attachment lamella semicircular, wide, better developed in central portion, situated along of anterior edge of maxilla (
<figureCitation id="FB8836C1FFB5FFB0720B52A3FD82FBAE" box="[587,687,1010,1034]" captionStart="Figure 8" captionStartId="19.[160,225,1408,1431]" captionTargetBox="[164,1152,143,1357]" captionTargetId="figure@19.[163,1153,142,1376]" captionTargetPageId="19" captionText="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="20" pageNumber="536">Figure 8</figureCitation>
(d)). MV square, with a short, rounded tooth (
<figureCitation id="FB8836C1FFB5FFB070B05545FE7DFB88" box="[240,336,1044,1068]" captionStart="Figure 8" captionStartId="19.[160,225,1408,1431]" captionTargetBox="[164,1152,143,1357]" captionTargetId="figure@19.[163,1153,142,1376]" captionTargetPageId="19" captionText="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="20" pageNumber="536">Figure 8</figureCitation>
(d,e)). Mandibles amber (
<figureCitation id="FB8836C1FFB5FFB072345545FDF9FB88" box="[628,724,1044,1068]" captionStart="Figure 8" captionStartId="19.[160,225,1408,1431]" captionTargetBox="[164,1152,143,1357]" captionTargetId="figure@19.[163,1153,142,1376]" captionTargetPageId="19" captionText="Figure 8. Treadwellphysa villalobosi sp. nov., holotype ECOSUR206. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation in the base of MII; (e) left MIV, lateral view, arrow indicates ligament between maxillae; (f) mandibles. Scale bars: ac = 1.5 mm; de = 0.6 mm; f = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.3675720" httpUri="https://zenodo.org/record/3675720/files/figure.png" pageId="20" pageNumber="536">Figure 8</figureCitation>
(f)); calcareous cutting plates broken, cutting plates dark, oval, with up to 11 growth rings.
</paragraph>
<paragraph id="630C2A44FFB5FFB07080550BFEEEFB58" blockId="20.[160,1156,142,1727]" pageId="20" pageNumber="536">
Branchiae palmate with up to three filaments, in chaetigers 52 to 365R
<emphasis id="51C7F656FFB5FFB073965508FCCEFBD5" box="[982,995,1113,1137]" italics="true" pageId="20" pageNumber="536"></emphasis>
376L (
<figureCitation id="FB8836C1FFB5FFB07468550BFBAEFBD6" box="[1064,1155,1114,1138]" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="20" pageNumber="536">Figure 9</figureCitation>
(d,e)). One filament along chaetigers 52L to 54L; two filaments along chaetigers 55L to 113L; three filaments along chaetigers 114L to 324L; two filaments along chaetigers 325L to 365L; one filament along chaetigers 366L to
<specimenCount id="75B5E1CDFFB5FFB072925593FC23FB7E" box="[722,782,1218,1242]" pageId="20" pageNumber="536" type="generic">376L.</specimenCount>
Branchial filaments longer than dorsal cirri (
<figureCitation id="FB8836C1FFB5FFB0716755B5FEA9FB58" box="[295,388,1252,1276]" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="20" pageNumber="536">Figure 9</figureCitation>
(d,e)).
</paragraph>
<paragraph id="630C2A44FFB5FFB070805456FFC0F91B" blockId="20.[160,1156,142,1727]" pageId="20" pageNumber="536">
First four parapodia smaller; best developed in chaetigers 15
<emphasis id="51C7F656FFB5FFB073DC5456FC84FABB" box="[924,937,1287,1311]" italics="true" pageId="20" pageNumber="536"></emphasis>
61, following ones becoming gradually smaller. Notopodial cirri conical, longer than ventral cirri in first 10 chaetigers, in chaetigers 11 to 38 of similar sizes, following ones gradually decreasing in size, shorter than ventral cirri (
<figureCitation id="FB8836C1FFB5FFB07251543EFD5FFA23" box="[529,626,1391,1415]" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="20" pageNumber="536">Figure 9</figureCitation>
(a
<emphasis id="51C7F656FFB5FFB072C6543EFDBEFA23" box="[646,659,1391,1415]" italics="true" pageId="20" pageNumber="536"></emphasis>
f)). Prechaetal lobes short, appearing as a transverse fold in all chaetigers (
<figureCitation id="FB8836C1FFB5FFB0726B54C3FDA0FA0E" box="[555,653,1426,1450]" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="20" pageNumber="536">Figure 9</figureCitation>
(a
<emphasis id="51C7F656FFB5FFB072E254C3FD82FA0E" box="[674,687,1426,1450]" italics="true" pageId="20" pageNumber="536"></emphasis>
f)). Chaetal lobes in chaetigers 1 to 25 rounded, slightly shorter than postchaetal lobes, with aciculae emerging dorsal to midline; from chaetiger 26 triangular, longer than other lobes, with aciculae emerging in midline (
<figureCitation id="FB8836C1FFB5FFB0714454ABFE4CF9B6" box="[260,353,1530,1554]" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="20" pageNumber="536">Figure 9</figureCitation>
(a
<emphasis id="51C7F656FFB5FFB0713654A8FEAEF9B5" box="[374,387,1529,1553]" italics="true" pageId="20" pageNumber="536"></emphasis>
f)). Postchaetal lobes poorly developed in chaetigers 1
<emphasis id="51C7F656FFB5FFB073AC54A8FCD4F9B5" box="[1004,1017,1529,1553]" italics="true" pageId="20" pageNumber="536"></emphasis>
37, rounded in first 10 chaetigers; tapered with blunt tip, progressively decreasing in size from chaetigers 11 to 37; following ones inconspicuous (
<figureCitation id="FB8836C1FFB5FFB072B3576EFC7FF9F3" box="[755,850,1599,1623]" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="20" pageNumber="536">Figure 9</figureCitation>
(a
<emphasis id="51C7F656FFB5FFB07326576EFC5EF9F3" box="[870,883,1599,1623]" italics="true" pageId="20" pageNumber="536"></emphasis>
f)). Ventral cirri in chaetigers 1
<emphasis id="51C7F656FFB5FFB070B35730FE2DF9DD" box="[243,256,1633,1657]" italics="true" pageId="20" pageNumber="536"></emphasis>
4 digitiform; in chaetigers 5 to 284 with a swollen base as a transverse welt with a short digitiform tip; from chaetiger 285 digitiform, gradually decreasing in size (
<figureCitation id="FB8836C1FFB5FFB0747C57D5FF83F91B" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="20" pageNumber="536">Figure 9</figureCitation>
(a
<emphasis id="51C7F656FFB5FFB0708257F6FFE2F91B" box="[194,207,1703,1727]" italics="true" pageId="20" pageNumber="536"></emphasis>
f)).
</paragraph>
<caption id="37CC7ACCFFB4FFB170E054B9FCE0F8AC" ID-DOI="http://doi.org/10.5281/zenodo.3675722" ID-Zenodo-Dep="3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="21" pageNumber="537" startId="21.[160,225,1512,1535]" targetBox="[164,1152,143,1480]" targetPageId="21">
<paragraph id="630C2A44FFB4FFB170E054B9FCE0F8AC" blockId="21.[160,1156,1512,1800]" pageId="21" pageNumber="537">
<emphasis id="51C7F656FFB4FFB170E054B9FFD1FA5B" bold="true" box="[160,252,1512,1535]" pageId="21" pageNumber="537">Figure 9.</emphasis>
<taxonomicName id="A4B351C7FFB4FFB1714654B8FD2CF9A4" authority="Molina-Acevedo, 2019" authorityName="Molina-Acevedo" authorityYear="2019" box="[262,513,1513,1536]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="537" phylum="Annelida" rank="species" species="villalobosi" status="sp. nov.">
<emphasis id="51C7F656FFB4FFB1714654B8FD2CF9A4" box="[262,513,1513,1536]" italics="true" pageId="21" pageNumber="537">Treadwellphysa villalobosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFB4FFB1724A54B8FD77F9A4" box="[522,602,1513,1536]" pageId="21" pageNumber="537" rank="species">sp. nov.</taxonomicNameLabel>
(a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view.
</paragraph>
</caption>
<paragraph id="630C2A44FFB7FFB2708051DFFC35FF48" blockId="22.[160,1156,142,825]" pageId="22" pageNumber="538">
Aciculae blunt, reddish along most of the length, translucent on the distal tip (
<figureCitation id="FB8836C1FFB7FFB273B251DFFB65FF02" box="[1010,1096,142,166]" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="22" pageNumber="538">Figure 9</figureCitation>
(a
<emphasis id="51C7F656FFB7FFB2741C51DFFB44FF02" box="[1116,1129,142,166]" italics="true" pageId="22" pageNumber="538"></emphasis>
f)). First two chaetigers with two aciculae, chaetigers 3
<emphasis id="51C7F656FFB7FFB2728551E0FDFFFF6D" box="[709,722,177,201]" italics="true" pageId="22" pageNumber="538"></emphasis>
49 with three aciculae; chaetigers 50
<emphasis id="51C7F656FFB7FFB2741C51E0FB44FF6D" box="[1116,1129,177,201]" italics="true" pageId="22" pageNumber="538"></emphasis>
62 with two aciculae; from chaetiger 63 with only one acicula.
</paragraph>
<paragraph id="630C2A44FFB7FFB2708051A7FFFBFD50" blockId="22.[160,1156,142,825]" pageId="22" pageNumber="538">
Limbate chaetae of two lengths in the same chaetiger, longer in the dorsal region of the parapodia, reduced in number in anterior chaetiger. Three
<typeStatus id="BC0894E6FFB7FFB273775048FC5CFE95" box="[823,881,281,305]" pageId="22" pageNumber="538">types</typeStatus>
of pectinate chaetae; in anterior chaetigers, isodont narrow, with short and slender teeth, with 1
<emphasis id="51C7F656FFB7FFB273E0506AFC80FEF7" box="[928,941,315,339]" italics="true" pageId="22" pageNumber="538"></emphasis>
2 pectinate, with up to 16 teeth (
<figureCitation id="FB8836C1FFB7FFB27163500FFE56FED2" box="[291,379,350,374]" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="22" pageNumber="538">Figure 9</figureCitation>
(g)); in median-posterior chaetigers, isodont wide, with short and slender teeth, with 4
<emphasis id="51C7F656FFB7FFB2717150D0FE13FE3D" box="[305,318,385,409]" italics="true" pageId="22" pageNumber="538"></emphasis>
6 pectinate, with up to 22
<emphasis id="51C7F656FFB7FFB2722B50D0FD55FE3D" box="[619,632,385,409]" italics="true" pageId="22" pageNumber="538"></emphasis>
24 teeth (
<figureCitation id="FB8836C1FFB7FFB272A950D0FC6BFE3D" box="[745,838,385,409]" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="22" pageNumber="538">Figure 9</figureCitation>
(h,i)); in posterior chaetigers, anodont wide, with long and thick teeth, with 2
<emphasis id="51C7F656FFB7FFB272E550F2FD9FFE1F" box="[677,690,419,443]" italics="true" pageId="22" pageNumber="538"></emphasis>
3 pectinate, with up to 10
<emphasis id="51C7F656FFB7FFB2738A50F2FCFAFE1F" box="[970,983,419,443]" italics="true" pageId="22" pageNumber="538"></emphasis>
14 teeth (
<figureCitation id="FB8836C1FFB7FFB2747F50F5FF83FE7A" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="22" pageNumber="538">Figure 9</figureCitation>
(h,i)). Compound spinifalciger and spiniger absent. Compound falcigers present in all chaetigers; in anterior region with blades of two lengths (longer 70 µm,
<figureCitation id="FB8836C1FFB7FFB2733650B8FCE3FDA5" box="[886,974,489,513]" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="22" pageNumber="538">Figure 9</figureCitation>
(l,m), smaller 46
<emphasis id="51C7F656FFB7FFB2743650B8FBAEFDA5" box="[1142,1155,489,513]" italics="true" pageId="22" pageNumber="538"></emphasis>
49 µm,
<figureCitation id="FB8836C1FFB7FFB270AC535DFE6CFD80" box="[236,321,524,548]" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="22" pageNumber="538">Figure 9</figureCitation>
(j,k)); with triangular teeth, of similar size, distal tooth directed upward, proximal tooth directed laterally. In median-posterior chaetigers all blades of similar length, shorter than blades of anterior chaetigers (32 µm,
<figureCitation id="FB8836C1FFB7FFB2727A5300FDB8FDCD" box="[570,661,593,617]" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="22" pageNumber="538">Figure 9</figureCitation>
(n)); with triangular teeth, distal tooth shorter than proximal, directed upward, proximal tooth directed laterally. Subacicular hooks bidentate, translucent, starting from chaetigers 50R
<emphasis id="51C7F656FFB7FFB272CF53C7FDB1FD0A" box="[655,668,662,686]" italics="true" pageId="22" pageNumber="538"></emphasis>
51L, one per chaetiger; with triangular teeth, distal tooth smaller than proximal, directed upward; proximal tooth directed laterally; (
<figureCitation id="FB8836C1FFB7FFB2747F53E8FF83FD50" captionStart="Figure 9" captionStartId="21.[160,225,1512,1535]" captionTargetBox="[164,1152,143,1480]" captionTargetId="figure@21.[163,1153,142,1481]" captionTargetPageId="21" captionText="Figure 9. Treadwellphysa villalobosi sp. nov. (a) Parapodium 3; (b) parapodium 14; (c) parapodium 42; (d) parapodium 104; (e) parapodium 209; (f) parapodium 394; (g) chaetiger 20, isodont pectinate narrow, with short and slender teeth; (h) chaetiger 249, isodont pectinate wide, with short and slender teeth; (i) chaetiger 249, anodont pectinate wide, with long and thick teeth; (j) chaetiger 17, compound falciger with short blade; (k) chaetiger 8, compound falciger with short blade; (l) chaetiger 10, compound falciger with long blade; (m) chaetiger 8, compound falciger with long blade; (n) chaetiger 249, compound falciger with short blade; (o) chaetiger 40, subacicular hook. af, k, m from holotype ECOSUR206; g, j, l from paratype ECOSUR207; h, i, n, o from paratype ECOSUR00000. Scale bars: a, f = 80 µm; be = 0.16 mm; g = 3.5 µm; h = 8.4 µm; i = 7.2 µm; j = 9.7 µm; k, m = 14 µm; l, n = 5.7 µm; o = 16 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675722" httpUri="https://zenodo.org/record/3675722/files/figure.png" pageId="22" pageNumber="538">Figure 9</figureCitation>
(o)).
</paragraph>
<paragraph id="630C2A44FFB7FFB2708053AFFC7EFC9D" blockId="22.[160,1156,142,825]" pageId="22" pageNumber="538">Pygidium with two pairs of anal cirri, without articulation; dorsal pair as long as last three chaetigers; ventral pair short, as long as last chaetiger.</paragraph>
<paragraph id="630C2A44FFB7FFB270E05237FE34FCDB" blockId="22.[160,1155,870,1276]" box="[160,281,870,895]" pageId="22" pageNumber="538">
<emphasis id="51C7F656FFB7FFB270E05237FE34FCDB" bold="true" box="[160,281,870,895]" italics="true" pageId="22" pageNumber="538">Variations</emphasis>
</paragraph>
<paragraph id="630C2A44FFB7FFB270E052D8FC29FB58" blockId="22.[160,1155,870,1276]" pageId="22" pageNumber="538">
Material examined with L10 =
<quantity id="A44B87A1FFB7FFB271A552D8FD4FFC05" box="[485,610,905,929]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" metricValueMax="5.7" metricValueMin="3.7" pageId="22" pageNumber="538" unit="mm" value="4.7" valueMax="5.7" valueMin="3.7">
3.7
<emphasis id="51C7F656FFB7FFB2724452D8FD3CFC05" box="[516,529,905,929]" italics="true" pageId="22" pageNumber="538"></emphasis>
5.7 mm
</quantity>
and W10 =
<quantity id="A44B87A1FFB7FFB272A452D8FC4EFC05" box="[740,867,905,929]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="22" pageNumber="538" unit="mm" value="1.9" valueMax="2.2" valueMin="1.6">
1.6
<emphasis id="51C7F656FFB7FFB2734352D8FC3DFC05" box="[771,784,905,929]" italics="true" pageId="22" pageNumber="538"></emphasis>
2.2 mm
</quantity>
.
<location id="666C7C9FFFB7FFB2732C52D8FCB7FC05" LSID="urn:lsid:plazi:treatment:EB1A9B52FFB3FFB371FB54CAFD4BFDA5:666C7C9FFFB7FFB2732C52D8FCB7FC05" box="[876,922,905,929]" name="Two" pageId="22" pageNumber="538">Two</location>
<specimenCount id="75B5E1CDFFB7FFB273E052D8FB3FFC05" box="[928,1042,905,929]" pageId="22" pageNumber="538" type="generic">specimens</specimenCount>
complete,
<typeStatus id="BC0894E6FFB7FFB270E052FDFE2CFC60" box="[160,257,940,964]" pageId="22" pageNumber="538" type="holotype">holotype</typeStatus>
with 417 and
<typeStatus id="BC0894E6FFB7FFB271E652FDFD2BFC60" box="[422,518,940,964]" pageId="22" pageNumber="538" type="paratype">paratype</typeStatus>
<specimenCode id="3315823FFFB7FFB2724F52FDFDB1FC60" box="[527,668,940,964]" pageId="22" pageNumber="538">ECOSUR0000</specimenCode>
with 271 chaetigers. Palps reaching second peristomial ring or first chaetiger. Lateral antennae reaching second or third chaetiger. Median antenna reaching third or fourth chaetiger.
<collectingMunicipality id="8368B03EFFB7FFB2722652A0FDA3FBAD" box="[614,654,1009,1033]" pageId="22" pageNumber="538">The</collectingMunicipality>
maxillary formula variations are
<specimenCode id="3315823FFFB7FFB273A152A0FB03FBAD" box="[993,1070,1009,1033]" collectionCode="MII" country="Ireland" lsid="urn:lsid:biocol.org:col:13666" name="Museum of Irish Industry" pageId="22" pageNumber="538" type="Herbarium">
MII 2
<emphasis id="51C7F656FFB7FFB2745352A0FB0DFBAD" box="[1043,1056,1009,1033]" italics="true" pageId="22" pageNumber="538"></emphasis>
3
</specimenCode>
+ 3,
<specimenCode id="3315823FFFB7FFB2741C52A0FFE7FB88" collectionCode="MIII" pageId="22" pageNumber="538">
MIII 4
<emphasis id="51C7F656FFB7FFB270ED5545FF97FB88" box="[173,186,1044,1068]" italics="true" pageId="22" pageNumber="538"></emphasis>
5
</specimenCode>
,
<specimenCode id="3315823FFFB7FFB270925545FE22FB88" box="[210,271,1044,1068]" collectionCode="MIV" country="Italy" httpUri="http://grbio.org/cool/mev6-t33n" name="Universita' degli Studi di Milano, Medicina Veterinaria, Sez. Parassitologia" pageId="22" pageNumber="538" type="University or college">MIV 4</specimenCode>
+ 5
<emphasis id="51C7F656FFB7FFB271775545FE69FB88" box="[311,324,1044,1068]" italics="true" pageId="22" pageNumber="538"></emphasis>
6. The proportions of the maxillary apparatus vary as follows: maxillary carriers 2.8
<emphasis id="51C7F656FFB7FFB270FF5567FFE1FBEA" box="[191,204,1078,1102]" italics="true" pageId="22" pageNumber="538"></emphasis>
3.6 times shorter than length of MI; closing system 3.4
<emphasis id="51C7F656FFB7FFB273425567FC22FBEA" box="[770,783,1078,1102]" italics="true" pageId="22" pageNumber="538"></emphasis>
4.4 times shorter than the length of the MI; cavity opening 2.9
<emphasis id="51C7F656FFB7FFB271885508FEF8FBD5" box="[456,469,1113,1137]" italics="true" pageId="22" pageNumber="538"></emphasis>
3.5 times shorter than the length of the MII. Branchiae from chaetigers 39
<emphasis id="51C7F656FFB7FFB27171552DFE13FB30" box="[305,318,1148,1172]" italics="true" pageId="22" pageNumber="538"></emphasis>
53 to 37
<emphasis id="51C7F656FFB7FFB271D9552DFE8BFB30" box="[409,422,1148,1172]" italics="true" pageId="22" pageNumber="538"></emphasis>
40 chaetigers before pygidium. Maximum number of branchial filaments varied from two to four. Poorly developed postchaetal lobe present in first 27
<emphasis id="51C7F656FFB7FFB2741C55CFFB44FB12" box="[1116,1129,1182,1206]" italics="true" pageId="22" pageNumber="538"></emphasis>
56 chaetigers. Ventral cirri with a swollen base from chaetigers 5 to 132
<emphasis id="51C7F656FFB7FFB273CA5590FCBAFB7D" box="[906,919,1217,1241]" italics="true" pageId="22" pageNumber="538"></emphasis>
136 chaetigers before pygidium. Start of subacicular hooks in chaetigers 31
<emphasis id="51C7F656FFB7FFB2729755B5FDC9FB58" box="[727,740,1252,1276]" italics="true" pageId="22" pageNumber="538"></emphasis>
46.
</paragraph>
</subSubSection>
<subSubSection id="2BA979CFFFB7FFB270E05478FD61FAC0" box="[160,588,1321,1380]" pageId="22" pageNumber="538" type="distribution">
<paragraph id="630C2A44FFB7FFB270E05478FE01FAE6" blockId="22.[160,588,1321,1380]" box="[160,300,1321,1346]" pageId="22" pageNumber="538">
<emphasis id="51C7F656FFB7FFB270E05478FE01FAE6" bold="true" box="[160,300,1321,1346]" italics="true" pageId="22" pageNumber="538">Distribution</emphasis>
</paragraph>
<paragraph id="630C2A44FFB7FFB270E0541DFD61FAC0" blockId="22.[160,588,1321,1380]" box="[160,588,1356,1380]" pageId="22" pageNumber="538">
Cabo Blanco,
<collectingCountry id="1BA46AD4FFB7FFB2717C541DFE9DFAC0" box="[316,432,1356,1380]" name="Costa Rica" pageId="22" pageNumber="538">Costa Rica</collectingCountry>
(Pacific side).
</paragraph>
</subSubSection>
<subSubSection id="2BA979CFFFB7FFB270E054C0FC30F9B6" pageId="22" pageNumber="538" type="etymology">
<paragraph id="630C2A44FFB7FFB270E054C0FE30FA0E" blockId="22.[160,1155,1425,1554]" box="[160,285,1425,1450]" pageId="22" pageNumber="538">
<emphasis id="51C7F656FFB7FFB270E054C0FE30FA0E" bold="true" box="[160,285,1425,1450]" italics="true" pageId="22" pageNumber="538">Etymology</emphasis>
</paragraph>
<paragraph id="630C2A44FFB7FFB270E054E5FC30F9B6" blockId="22.[160,1155,1425,1554]" pageId="22" pageNumber="538">This species is named in honour of my colleague Tulio F. Villalobos-Guerrero, in recognition of his contributions to the study of errant polychaetes, especially on nereidids from the Mexican Pacific, and for collecting these specimens.</paragraph>
</subSubSection>
<subSubSection id="2BA979CFFFB7FFB370E0576FFD4BFDA5" lastPageId="23" lastPageNumber="539" pageId="22" pageNumber="538" type="discussion">
<paragraph id="630C2A44FFB7FFB270E0576FFE2BF9F3" blockId="22.[160,1155,1598,1727]" box="[160,262,1598,1623]" pageId="22" pageNumber="538">
<emphasis id="51C7F656FFB7FFB270E0576FFE2BF9F3" bold="true" box="[160,262,1598,1623]" italics="true" pageId="22" pageNumber="538">Remarks</emphasis>
</paragraph>
<paragraph id="630C2A44FFB7FFB370E05733FD4BFDA5" blockId="22.[160,1155,1598,1727]" lastBlockId="23.[160,1156,142,513]" lastPageId="23" lastPageNumber="539" pageId="22" pageNumber="538">
<taxonomicName id="A4B351C7FFB7FFB270E05733FE80F9DE" authority="Molina-Acevedo, 2019" authorityName="Molina-Acevedo" authorityYear="2019" box="[160,429,1634,1658]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="22" pageNumber="538" phylum="Annelida" rank="species" species="villalobosi" status="sp. nov.">
<emphasis id="51C7F656FFB7FFB270E05733FE80F9DE" box="[160,429,1634,1658]" italics="true" pageId="22" pageNumber="538">Treadwellphysa villalobosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFB7FFB271F25733FD28F9DE" box="[434,517,1634,1658]" pageId="22" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="A4B351C7FFB7FFB2723B5733FDC3F9DE" baseAuthorityName="Treadwelli" baseAuthorityYear="1921" box="[635,750,1634,1658]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="22" pageNumber="538" phylum="Annelida" rank="species" species="languida">
<emphasis id="51C7F656FFB7FFB2723B5733FDC3F9DE" box="[635,750,1634,1658]" italics="true" pageId="22" pageNumber="538">T. languida</emphasis>
</taxonomicName>
,
<taxonomicName id="A4B351C7FFB7FFB272B95733FC75F9DE" authority="Molina-Acevedo, 2019" authorityName="Molina-Acevedo" authorityYear="2019" box="[761,856,1634,1658]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="22" pageNumber="538" phylum="Annelida" rank="species" species="rizzoae" status="sp. nov.">
<emphasis id="51C7F656FFB7FFB272B95733FC75F9DE" box="[761,856,1634,1658]" italics="true" pageId="22" pageNumber="538">T. rizzoae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFB7FFB2731E5733FC9DF9DE" box="[862,944,1634,1658]" pageId="22" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="A4B351C7FFB7FFB273A35733FBAEF9DE" baseAuthorityName="de Leon-Gonzalez and Diaz-Castanedai" baseAuthorityYear="2006" box="[995,1155,1634,1658]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="22" pageNumber="538" phylum="Annelida" rank="species" species="veracruzensis">
<emphasis id="51C7F656FFB7FFB273A35733FBAEF9DE" box="[995,1155,1634,1658]" italics="true" pageId="22" pageNumber="538">T. veracruzensis</emphasis>
</taxonomicName>
in having compound falcigers, whereas compound spinigers are absent. However,
<taxonomicName id="A4B351C7FFB7FFB2744657D4FBAEF939" authority="Molina-Acevedo, 2019" authorityName="Molina-Acevedo" authorityYear="2019" box="[1030,1155,1669,1693]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="22" pageNumber="538" phylum="Annelida" rank="species" species="villalobosi" status="sp. nov.">
<emphasis id="51C7F656FFB7FFB2744657D4FBAEF939" box="[1030,1155,1669,1693]" italics="true" pageId="22" pageNumber="538">T. villalobosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFB7FFB270E057F6FFD5F91B" box="[160,248,1703,1727]" pageId="22" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
lacks compound spinifalcigers, whereas in
<taxonomicName id="A4B351C7FFB7FFB272A057F6FC74F91B" baseAuthorityName="Treadwelli" baseAuthorityYear="1921" box="[736,857,1703,1727]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="22" pageNumber="538" phylum="Annelida" rank="species" species="languida">
<emphasis id="51C7F656FFB7FFB272A057F6FC74F91B" box="[736,857,1703,1727]" italics="true" pageId="22" pageNumber="538">T. languida</emphasis>
</taxonomicName>
and
<taxonomicName id="A4B351C7FFB7FFB273D857F6FB10F91B" baseAuthorityName="de Leon-Gonzalez and Diaz-Castanedai" baseAuthorityYear="2006" box="[920,1085,1703,1727]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="22" pageNumber="538" phylum="Annelida" rank="species" species="veracruzensis">
<emphasis id="51C7F656FFB7FFB273D857F6FB10F91B" box="[920,1085,1703,1727]" italics="true" pageId="22" pageNumber="538">T. veracruzensis</emphasis>
</taxonomicName>
these compound chaetae are present. Furthermore, in
<taxonomicName id="A4B351C7FFB6FFB372F651DFFC01FF02" baseAuthorityName="Treadwelli" baseAuthorityYear="1921" box="[694,812,142,166]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="23" pageNumber="539" phylum="Annelida" rank="species" species="languida">
<emphasis id="51C7F656FFB6FFB372F651DFFC01FF02" box="[694,812,142,166]" italics="true" pageId="23" pageNumber="539">T. languida</emphasis>
</taxonomicName>
(holotype, L10:
<quantity id="A44B87A1FFB6FFB3739D51DFFB1CFF02" box="[989,1073,142,166]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="23" pageNumber="539" unit="mm" value="4.5">4.5 mm</quantity>
) and
<taxonomicName id="A4B351C7FFB6FFB3743151DFFE04FF6D" baseAuthorityName="de Leon-Gonzalez and Diaz-Castanedai" baseAuthorityYear="2006" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="23" pageNumber="539" phylum="Annelida" rank="species" species="veracruzensis">
<emphasis id="51C7F656FFB6FFB3743151DFFE04FF6D" italics="true" pageId="23" pageNumber="539">T. veracruzensis</emphasis>
</taxonomicName>
(ECOSUR-OH-P675, L10:
<quantity id="A44B87A1FFB6FFB3720351E0FDA5FF6D" box="[579,648,177,201]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="539" unit="mm" value="3.0">3 mm</quantity>
) the branchiae start in chaetiger 60 and 35, respectively, while in
<taxonomicName id="A4B351C7FFB6FFB371D55185FD34FF48" authority="Molina-Acevedo, 2019" authorityName="Molina-Acevedo" authorityYear="2019" box="[405,537,212,236]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="23" pageNumber="539" phylum="Annelida" rank="species" species="villalobosi" status="sp. nov.">
<emphasis id="51C7F656FFB6FFB371D55185FD34FF48" box="[405,537,212,236]" italics="true" pageId="23" pageNumber="539">T. villalobosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFB6FFB372655185FD50FF48" box="[549,637,212,236]" pageId="23" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
(holotype, L10:
<quantity id="A44B87A1FFB6FFB373775185FC51FF48" box="[823,892,212,236]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="539" unit="mm" value="6.0">6 mm</quantity>
) the branchiae start in chaetiger 52. Moreover,
<taxonomicName id="A4B351C7FFB6FFB371E551A7FD09FEAA" authority="Molina-Acevedo, 2019" authorityName="Molina-Acevedo" authorityYear="2019" box="[421,548,246,270]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="23" pageNumber="539" phylum="Annelida" rank="species" species="villalobosi" status="sp. nov.">
<emphasis id="51C7F656FFB6FFB371E551A7FD09FEAA" box="[421,548,246,270]" italics="true" pageId="23" pageNumber="539">T. villalobosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFB6FFB3726B51A7FD52FEAA" box="[555,639,246,270]" pageId="23" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="A4B351C7FFB6FFB3734851A7FC47FEAA" authority="Molina-Acevedo, 2019" authorityName="Molina-Acevedo" authorityYear="2019" box="[776,874,246,270]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="23" pageNumber="539" phylum="Annelida" rank="species" species="rizzoae" status="sp. nov.">
<emphasis id="51C7F656FFB6FFB3734851A7FC47FEAA" box="[776,874,246,270]" italics="true" pageId="23" pageNumber="539">T. rizzoae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFB6FFB3733051A7FCE9FEAA" box="[880,964,246,270]" pageId="23" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
in the colour, the starting chaetiger of the subacicular hooks, and the starting and ending of the branchiae (see remarks section of
<taxonomicName id="A4B351C7FFB6FFB37138506DFEF0FEF0" authority="Molina-Acevedo, 2019" authorityName="Molina-Acevedo" authorityYear="2019" box="[376,477,316,340]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="23" pageNumber="539" phylum="Annelida" rank="species" species="rizzoae" status="sp. nov.">
<emphasis id="51C7F656FFB6FFB37138506DFEF0FEF0" box="[376,477,316,340]" italics="true" pageId="23" pageNumber="539">T. rizzoae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFB6FFB371A8506DFD6FFEF0" box="[488,578,316,340]" pageId="23" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
). Likewise, the two species are different because
<taxonomicName id="A4B351C7FFB6FFB37431506DFE2AFED2" authority="Molina-Acevedo, 2019" authorityName="Molina-Acevedo" authorityYear="2019" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="23" pageNumber="539" phylum="Annelida" rank="species" species="villalobosi" status="sp. nov.">
<emphasis id="51C7F656FFB6FFB37431506DFE2AFED2" italics="true" pageId="23" pageNumber="539">T. villalobosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFB6FFB3714F500FFE49FED2" box="[271,356,350,374]" pageId="23" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
has tapered postchaetal lobes with blunt tip and the ventral cirri have a higher number of chaetigers with a swollen base, from chaetiger 5 to 284 (
<typeStatus id="BC0894E6FFB6FFB373A850D0FB67FE3D" box="[1000,1098,385,409]" pageId="23" pageNumber="539" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A4B351C7FFB6FFB3743150D0FE2AFE18" authorityName="Molina-Acevedo" authorityYear="2019" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="23" pageNumber="539" phylum="Annelida" rank="species" species="villalobosi">
<emphasis id="51C7F656FFB6FFB3743150D0FE2AFE18" italics="true" pageId="23" pageNumber="539">T. villalobosi</emphasis>
</taxonomicName>
has 417 chaetigers), whereas in
<taxonomicName id="A4B351C7FFB6FFB3721B50F5FD97FE18" authority="Molina-Acevedo, 2019" authorityName="Molina-Acevedo" authorityYear="2019" box="[603,698,420,444]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="23" pageNumber="539" phylum="Annelida" rank="species" species="rizzoae" status="sp. nov.">
<emphasis id="51C7F656FFB6FFB3721B50F5FD97FE18" box="[603,698,420,444]" italics="true" pageId="23" pageNumber="539">T. rizzoae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFB6FFB3728050F5FC3FFE18" box="[704,786,420,444]" pageId="23" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
the postchaetal lobes are rounded and the ventral cirri have fewer chaetigers with a swollen base, from chaetiger 6 to 85 (
<typeStatus id="BC0894E6FFB6FFB370E650B8FE25FDA5" box="[166,264,489,513]" pageId="23" pageNumber="539" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A4B351C7FFB6FFB3716A50B8FEA7FDA5" authorityName="Molina-Acevedo" authorityYear="2019" box="[298,394,489,513]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="23" pageNumber="539" phylum="Annelida" rank="species" species="rizzoae">
<emphasis id="51C7F656FFB6FFB3716A50B8FEA7FDA5" box="[298,394,489,513]" italics="true" pageId="23" pageNumber="539">T. rizzoae</emphasis>
</taxonomicName>
has 210 chaetigers).
</paragraph>
</subSubSection>
</treatment>
</document>