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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.724.21415" ID-GBIF-Dataset="99799f60-7061-439a-855d-53f781229ca9" ID-PMC="PMC5769737" ID-Pensoft-Pub="1313-2970-724-53" ID-PubMed="29362533" ID-ZBK="BCDF3F207E274CCFA47467DA61308A78" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-724-53" ModsDocOrigin="ZooKeys 724" ModsDocTitle="Aquatic dance flies (Diptera, Empididae, Clinocerinae and Hemerodromiinae) of Greece: species richness, distribution and description of five new species" checkinTime="1513881635527" checkinUser="pensoft" docAuthor="Ivkovic, Marija, Cevid, Josipa, Horvat, Bogdan &amp; Sinclair, Bradley J." docDate="2017" docId="E8E3DE57533B8CCD0D1D86318F6F7AC5" docLanguage="en" docName="ZooKeys 724: 53-100" docOrigin="ZooKeys 724" docSource="http://dx.doi.org/10.3897/zookeys.724.21415" docTitle="Chelifera horvati Ivkovic &amp; Sinclair, sp. n." docType="treatment" docUuid="9DE403F2-5A28-4E6F-A485-A4D42308165D" docUuidSource="ZooBank" docVersion="5" lastPageNumber="62" masterDocId="FFA4FFD8AC29FFE5CB1EFF8C2D70FFAC" masterDocTitle="Aquatic dance flies (Diptera, Empididae, Clinocerinae and Hemerodromiinae) of Greece: species richness, distribution and description of five new species" masterLastPageNumber="100" masterPageNumber="53" pageNumber="61" updateTime="1668165242534" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Aquatic dance flies (Diptera, Empididae, Clinocerinae and Hemerodromiinae) of Greece: species richness, distribution and description of five new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ivkovic, Marija</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cevid, Josipa</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Horvat, Bogdan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sinclair, Bradley J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>724</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>53</mods:start>
<mods:end>100</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.724.21415</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.724.21415</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-724-53</mods:identifier>
<mods:identifier type="ZBK">BCDF3F207E274CCFA47467DA61308A78</mods:identifier>
<mods:identifier type="ZooBank">BCDF3F207E274CCFA47467DA61308A78</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="139438677" LSID="urn:lsid:zoobank.org:act:9DE403F2-5A28-4E6F-A485-A4D42308165D" httpUri="http://treatment.plazi.org/id/E8E3DE57533B8CCD0D1D86318F6F7AC5" lastPageId="9" lastPageNumber="62" pageId="8" pageNumber="61">
<subSubSection pageId="8" pageNumber="61" type="nomenclature">
<paragraph pageId="8" pageNumber="61">
<taxonomicName LSID="http://zoobank.org/9DE403F2-5A28-4E6F-A485-A4D42308165D" authority="Ivkovic &amp; Sinclair" class="Insecta" family="Empididae" genus="Chelifera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chelifera horvati" order="Diptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="horvati">
Chelifera horvati
<normalizedToken originalValue="Ivković">Ivkovic</normalizedToken>
&amp; Sinclair
</taxonomicName>
<taxonomicNameLabel pageId="8" pageNumber="61">sp. n.</taxonomicNameLabel>
Figs 1, 11
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="61" type="type locality">
<paragraph pageId="8" pageNumber="61">Type locality.</paragraph>
<paragraph pageId="8" pageNumber="61">Greece: Central Greece, Etolia, Arta, Loutraki.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="61" type="type material">
<paragraph pageId="8" pageNumber="61">Type material.</paragraph>
<paragraph pageId="8" pageNumber="61">
HOLOTYPE ♂, labelled: &quot;GREECE: Central Greece/ Etolia, Arta, Loutraki/ 16.iv.1990/ leg. B. Horvat, I. Sivec&quot;; &quot;HOLOTYPE/
<taxonomicName class="Insecta" family="Empididae" genus="Chelifera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chelifera" order="Diptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="genus">Chelifera</taxonomicName>
/
<taxonomicName lsidName="horvati" pageId="8" pageNumber="61" rank="species" species="horvati">horvati</taxonomicName>
/
<normalizedToken originalValue="Ivković">Ivkovic</normalizedToken>
&amp; Sinclair&quot; (CNC, dried from alcohol).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="61" type="diagnosis">
<paragraph pageId="8" pageNumber="61">Diagnosis.</paragraph>
<paragraph pageId="8" pageNumber="61">A yellow-brown species with distinct, brown and rounded pterostigma, characterized in the male by dark brown cercus with elongate, slender forked process at mid-length, posteriorly tapered epandrium with stout inner setae and membranous distiphallus with two elongate lobes.</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="62" pageId="8" pageNumber="61" type="description">
<paragraph pageId="8" pageNumber="61">Description.</paragraph>
<paragraph pageId="8" pageNumber="61">
Male. Body length 4 mm, wing length 3.6 mm. Head dorsoventrally flattened, dark brown; ocellar triangle dark brown; all setae whitish. Eyes iridescent black; narrowly separated on face. Face with thick, whitish pubescence. One pair of postocular setae and scattered fine setae on vertex. Occiput bearing scattered fine setulae; gena with rather dense short, downwardly directed whitish pile. Antenna whitish, with scape and pedicel bearing distinct short dorsal setulae; postpedicel about 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, stylus much shorter than postpedicel.
</paragraph>
<paragraph pageId="8" pageNumber="61">Thorax elongate; yellow, all setae yellowish. Mesonotum with pair of brown vittae, extending around prescutellar depression; small dark spot posterior to postpronotal lobe and larger dark spot near wing base. Holotype missing most thoracic setae.</paragraph>
<paragraph pageId="8" pageNumber="61">
Wing (slightly damaged) membrane transparent, veins yellow; pterostigma dark, rounded, with R2+3 arched around it; fork of R4+5 less than 90°; cell r4 rather long, R5 nearly 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as R4. Halter pale.
</paragraph>
<paragraph lastPageId="9" lastPageNumber="62" pageId="8" pageNumber="61">
Legs whitish yellow, apical two tarsal segments on all legs brown. Fore coxa about 8
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide with several pale dorsoapical setae. Fore femur slightly longer than fore coxa, more than 4
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, evenly inflated, widest at middle.
<pageBreakToken pageId="9" pageNumber="62" start="start">Fore</pageBreakToken>
femur with two rows of black ventral denticles and two rows of strong outer brownish-yellow ventral setae, with following chaetotaxy: 20 anteroventral denticles, 6 anteroventral spine-like setae, 21 posteroventral denticles, 6 posteroventral spine-like setae; denticles closely spaced and rows converging distally; posteroventral spine-like setae shorter distally. Fore tibia 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as fore femur, evenly curved with anteroventral row of short, spine-like setae; with apicoventral dark spur-like seta, longer than width of tibia. Mid and hind femora with anteroventral row of short, slender setae.
</paragraph>
<paragraph pageId="9" pageNumber="62">Abdomen yellow ventrally, brown dorsally, with pale setae most conspicuous on hind margin of posterior sternites. Terminalia (Fig. 11): cercus dark brown, thick, with narrow, elongate process at mid-length with forked apex (process folded horizontally in non-macerated condition); anterior end of cercus pointed and curved medially, with long setae, posterior end of cercus rounded; cercus wider then epandrium. Epandrium yellowish-brown, concave medially, posteriorly pointed with 5 stout setae on inner apical margin directed medially; entire epandrium covered in numerous setae. Hypandrium yellow, quadrate, with posteroapical lobe and concave posterior margin; pale setae on posteroventral face. Postgonite slender, sickle-shaped. Distiphallus membranous, expanded into two elongate lobes; apex of posterior lobe with pigmented arch-shaped sclerotization.</paragraph>
<paragraph pageId="9" pageNumber="62">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="62" type="etymology">
<paragraph pageId="9" pageNumber="62">Etymology.</paragraph>
<paragraph pageId="9" pageNumber="62">
The new species is named after the late Dr Bogdan Horvat, mentor of the first author, colleague and during his life a leading expert on the genus
<taxonomicName class="Insecta" family="Empididae" genus="Chelifera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chelifera" order="Diptera" pageId="9" pageNumber="62" phylum="Arthropoda" rank="genus">Chelifera</taxonomicName>
Macquart.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="62" type="remarks">
<paragraph pageId="9" pageNumber="62">Remarks.</paragraph>
<paragraph pageId="9" pageNumber="62">
<taxonomicName class="Insecta" family="Empididae" genus="Chelifera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chelifera horvati" order="Diptera" pageId="9" pageNumber="62" phylum="Arthropoda" rank="species" species="horvati">Chelifera horvati</taxonomicName>
sp. n. is known only from one site in Greece. The narrow pigmented and sclerotized apex of the distiphallus of
<taxonomicName lsidName="C. horvati" pageId="9" pageNumber="62" rank="species" species="horvati">C. horvati</taxonomicName>
sp. n. is similar in
<taxonomicName lsidName="C. concinnicauda" pageId="9" pageNumber="62" rank="species" species="concinnicauda">C. concinnicauda</taxonomicName>
Collin, 1927,
<taxonomicName lsidName="C. diversicauda" pageId="9" pageNumber="62" rank="species" species="diversicauda">C. diversicauda</taxonomicName>
Collin, 1927,
<taxonomicName lsidName="C. giraudae" pageId="9" pageNumber="62" rank="species" species="giraudae">C. giraudae</taxonomicName>
Vaillant, 1982 and
<taxonomicName lsidName="C. subangusta" pageId="9" pageNumber="62" rank="species" species="subangusta">C. subangusta</taxonomicName>
Collin, 1961 (see
<bibRefCitation author="Collin, JE" journalOrPublisher="University Press, Cambridge" pageId="32" pageNumber="85" title="Empididae. British flies. Volume 6." year="1961">Collin 1961</bibRefCitation>
and
<bibRefCitation author="Vaillant, F" journalOrPublisher="Bonner Zoologische Beitraege" pageId="34" pageNumber="87" pagination="351 - 408" title="Dipteres EmpididaeHemerodromiinae nouveaux ou per connus de la region palearctique (premiere partie)." volume="32" year="1982">Vaillant 1982</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>