159 lines
26 KiB
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159 lines
26 KiB
XML
<document ID-CLB-Dataset="6712" ID-DOI="http://dx.doi.org/10.3897/dez.65.21000" ID-GBIF-Dataset="03cce405-5887-4474-b040-35de18e8d020" ID-Pensoft-Pub="1860-1324-1-1" ID-Pensoft-UUID="77D7AEC9294554E5BBDCABA37901022A" ID-Zenodo-Dep="1166397" ID-ZooBank="73DEE0F32BB04A21B4455E168FE50F54" ModsDocID="1860-1324-65-1-1" checkinTime="1621372489409" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2018" docId="E8893B80A39B47D3090E4754C6154251" docLanguage="en" docName="DeutEntomolZeit 65(1): 1-63" docOrigin="Deutsche Entomologische Zeitschrift 65 (1)" docPubDate="2018-01-18" docSource="http://dx.doi.org/10.3897/dez.65.21000" docTitle="Qecyclothorax (Meonochilus) lewisensis subsp. subgenus Liebherr & Marris, 2009 [new status] 2009" docType="treatment" docVersion="5" id="77D7AEC9294554E5BBDCABA37901022A" lastPageNumber="12" masterDocId="77D7AEC9294554E5BBDCABA37901022A" masterDocTitle="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel" masterLastPageNumber="63" masterPageNumber="1" pageNumber="12" updateTime="1732929906348" updateUser="ExternalLinkService">
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<mods:title id="CFFA9915DB5C09AF843593F3B024BAF2">Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel</mods:title>
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<mods:namePart id="EEBA05FC9C9FEA88C49354E13B2D0230">Liebherr, James K.</mods:namePart>
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<mods:affiliation id="A08D90B505CAE2B5637630E0D00D2EF5">Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<mods:nameIdentifier id="1278698B0D60969A9B3A175A8E64ABF1" type="email">jkl5@cornell.edu</mods:nameIdentifier>
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<mods:title id="AF10D68A8F5F4162EFB8B685343C94E6">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="5912567BD9531DCE4A0EAB2EFA87A70A">2018</mods:date>
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<mods:number id="B85251976CA598023753A680CC753AA9">65</mods:number>
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<mods:classification id="069085A44A30ACF9067E56E17C504852">journal article</mods:classification>
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<mods:identifier id="B9C9382347FCAF7CD8CDB9124444E178" type="DOI">http://dx.doi.org/10.3897/dez.65.21000</mods:identifier>
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<treatment id="E8893B80A39B47D3090E4754C6154251" ID-GBIF-Taxon="182223680" LSID="urn:lsid:plazi:treatment:E8893B80A39B47D3090E4754C6154251" httpUri="http://treatment.plazi.org/id/E8893B80A39B47D3090E4754C6154251" lastPageId="12" lastPageNumber="12" pageId="11" pageNumber="12" scope_family="Carabidae" scope_genus="Mecyclothorax" scope_order="Coleoptera" scope_tribe="Moriomorphini">
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<subSubSection id="BB200FCC6234B6076DEDAD9D413B049E" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="49C20C1B5608BD3DCA5BDC8A7829755C" pageId="11" pageNumber="12">
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<taxonomicName id="BA6C866236458A45A8FD8C3DF8D5C129" authority="Meonochilus Liebherr & Marris, 2009 [new status]" authorityName="Liebherr & Marris, 2009 [new status]" authorityYear="2009" class="Insecta" family="Carabidae" genus="Qecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="(Meonochilus) subsp. subgenus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subGenus" subGenus="Meonochilus">subgenus Meonochilus Liebherr & Marris, 2009 [new status]</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="F4C9BE2335F7750A11A9F8B6A26A5D9F" pageId="11" pageNumber="12" type="diagnosis">
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<paragraph id="3B63220B441E1BE3241109386C7363CC" pageId="11" pageNumber="12">Diagnosis.</paragraph>
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<paragraph id="96B328D1ACA822000233C2FB62C31983" pageId="11" pageNumber="12">
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Species of this subgenus share a robust, convex body shape with
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<taxonomicName id="8152CD3DFD0AC52A05D9751920B588D9" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Qecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qecyclothorax" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="2488129724796E006D15EAA2BF0F3D04" italics="true" pageId="11" pageNumber="12">Qecyclothorax</emphasis>
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</taxonomicName>
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spp. (Fig.
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<figureCitation id="1F612CC6FE6062BD8294D99B0B2097A1" captionStart="Figure 8" captionStartId="F9" captionText="Figure 8. Mecyclothorax spp., dorsal view, with subgeneric assignments: A, M. (Eucyclothorax, subgen. n.) Mecyclothorax blackburni (type species); B, M. (Qecyclothorax, subgen. n.) Mecyclothorax storeyi (type species); C, M. (Meonochilus, status n.) Meonochilus amplipennis (type species); D, M. (Mecyclothorax) montivagus (type species); E, M. (Mecyclothorax) lissus." figureDoi="10.3897/dez.65.21000.figure8" httpUri="https://binary.pensoft.net/fig/179964" pageId="11" pageNumber="12">8B-C</figureCitation>
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), though in all
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<taxonomicName id="114A993419CB556A96C85DE854CDAFF4" authorityName="Liebherr & Marris" authorityYear="2009" class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Meonochilus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="298F606830807742D6E22D4DF4D6E7EA" italics="true" pageId="11" pageNumber="12">Meonochilus</emphasis>
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</taxonomicName>
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spp. the elytral striae are well developed and punctate, with striae 1-7 deep at elytral midlength. The pronotal median base is depressed relative to the convex disc (
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<bibRefCitation id="4AA3EC020BEF4E1E2CAAD3BAECAEFB8F" DOI="https://doi.org/10.3897/zookeys.147.1898" author="Liebherr, JK" journalOrPublisher="from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae ZooKeys" pageId="51" pageNumber="52" pagination="277 - 335" refId="B57" refString="Liebherr, JK, 2011. Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n. from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae ZooKeys 147: 277 - 335, DOI: https://doi.org/10.3897/zookeys.147.1898" title="Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n." url="https://doi.org/10.3897/zookeys.147.1898" volume="147" year="2011">Liebherr 2011</bibRefCitation>
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, figs 9-10). The broad elytra are associated with a rounded humerus, a condition also observed among some New Caledonian
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<taxonomicName id="BCA5E05AA54117A9E6E1CC95FAFB526C" class="Insecta" family="Carabidae" genus="Phacothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phacothorax" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="8411D8F2227D5B7C0A21707BE2FB1212" italics="true" pageId="11" pageNumber="12">Phacothorax</emphasis>
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</taxonomicName>
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and various species of subgenus
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<taxonomicName id="D6AF24C479FFBB25BF6DF42A2434436E" authority="Mecyclothorax" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Mecyclothorax)" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subGenus" subGenus="Mecyclothorax">Mecyclothorax Mecyclothorax</taxonomicName>
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. The head is transversely depressed behind the eyes, and the mentum is broad: over 3
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<normalizedToken id="866232D76C8C785CAB95BC0E18FDCC04" originalValue="×">x</normalizedToken>
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broad as long. Opposed to New Zealand species of subgenus
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<taxonomicName id="3746C828886EA03421D242ABB6642C39" authority="Mecyclothorax" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Mecyclothorax)" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subGenus" subGenus="Mecyclothorax">Mecyclothorax Mecyclothorax</taxonomicName>
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(as in
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<normalizedToken id="85131CA86D6D7F83E62BC1141358F681" originalValue="Hawaii’s">Hawaii's</normalizedToken>
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<taxonomicName id="F7D1A2E3829A73CC3768E02F0653DFAE" lsidName="M. montivagus" pageId="11" pageNumber="12" rank="species" species="montivagus">
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<emphasis id="49BFA5786873AB742E0281CD7EF1245F" italics="true" pageId="11" pageNumber="12">M. montivagus</emphasis>
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</taxonomicName>
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, Fig.
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<figureCitation id="9168CCE980D779A2BE9CAC715A6679B7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Head and mouthpart structures of Mecyclothorax spp. A-C. Head capsule, dorsal view: A, M. goweri; B, M. fleutiauxi; C, M. laterorectus. D-I, Mentum, labial palps and ligula: D, M. peryphoides, ventral view; E, M. storeyi, ventral view; F, M. amplipennis, dorsal view; G, M. montivagus, dorsal view; H, M. kanak, ventral view; I, M. fleutiauxi, ventral view. Abbreviations include: aso, anterior supraorbital seta; pso, posterior supraorbital seta." figureDoi="10.3897/dez.65.21000.figure1" httpUri="https://binary.pensoft.net/fig/179957" pageId="11" pageNumber="12">1G</figureCitation>
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), the ligula is rounded apically with the ligular setal sockets separated by only 2 diameters. The paraglossae extend beyond the ligular margin 1-2
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<normalizedToken id="8280EB6307AF682E38D5450C98071F5F" originalValue="×">x</normalizedToken>
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the distance from their base to the
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<normalizedToken id="42430893A94D79F5C71BBF189969F824" originalValue="ligula’s">ligula's</normalizedToken>
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margin (Fig.
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<figureCitation id="1779C91327CD90D80D627747D6BECF9A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Head and mouthpart structures of Mecyclothorax spp. A-C. Head capsule, dorsal view: A, M. goweri; B, M. fleutiauxi; C, M. laterorectus. D-I, Mentum, labial palps and ligula: D, M. peryphoides, ventral view; E, M. storeyi, ventral view; F, M. amplipennis, dorsal view; G, M. montivagus, dorsal view; H, M. kanak, ventral view; I, M. fleutiauxi, ventral view. Abbreviations include: aso, anterior supraorbital seta; pso, posterior supraorbital seta." figureDoi="10.3897/dez.65.21000.figure1" httpUri="https://binary.pensoft.net/fig/179957" pageId="11" pageNumber="12">1F</figureCitation>
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). The male parameres also differ from those occurring among taxa of other
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<taxonomicName id="53F913D41F5A98FB648D2EE0EDEE2A59" authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="FF26EFA522C22F943AA777C469BA6AB4" italics="true" pageId="11" pageNumber="12">Mecyclothorax</emphasis>
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</taxonomicName>
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subgenera, with the right paramere club-shaped and broadly setose apically (Fig.
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<figureCitation id="C44C263D0E353C311D85CB897B5A8A2B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paired left (above) and right (below) parameres of Mecyclothorax spp. (ectal view): A, M. blackburni; B, M. storeyi; C, M. amplipennis; D, M. montivagus; E, M. laterobustus. As the aedeagal assembly of M. storeyi males is inverted relative to other Mecyclothorax species (Moore 1984), the configuration of the upper, or right paramere of M. storeyi should be compared to the ventral, or left parameres of the other species, and vice versa." figureDoi="10.3897/dez.65.21000.figure5" httpUri="https://binary.pensoft.net/fig/179961" pageId="11" pageNumber="12">5C</figureCitation>
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), and the left paramere broadly conchoid.
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</paragraph>
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</subSubSection>
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<subSubSection id="9D4876D3A7AB66110B8468C77FFEECED" pageId="11" pageNumber="12" type="member species">
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<paragraph id="CAF4E5B65A51973201005F9A7FFEC0DA" pageId="11" pageNumber="12">Member species.</paragraph>
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<paragraph id="F8A8EACE916624C6EFB04F888C9A8DBF" pageId="11" pageNumber="12">
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The six species of this subgenus were revised by
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<bibRefCitation id="998763B0E2A30074FCE07B930C393124" DOI="https://doi.org/10.3897/zookeys.147.1898" author="Liebherr, JK" journalOrPublisher="from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae ZooKeys" pageId="51" pageNumber="52" pagination="277 - 335" refId="B57" refString="Liebherr, JK, 2011. Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n. from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae ZooKeys 147: 277 - 335, DOI: https://doi.org/10.3897/zookeys.147.1898" title="Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n." url="https://doi.org/10.3897/zookeys.147.1898" volume="147" year="2011">Liebherr (2011)</bibRefCitation>
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, and comprise the five species in this analysis (Fig.
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<figureCitation id="BC708AD10554468630A2962231599F20" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Strict consensus cladogram of 16 equally most-parsimonious trees derived from cladistic analysis of 65 moriomorphine taxa, including the secondary outgroup Neonomius laevicollis, six primary outgroup taxa in Amblytelus and allied genera, and 58 ingroup terminals; 57 Mecyclothorax species plus two genitalic forms of M. andersoni. Species terminals are labeled with species name and three-letter abbreviation of relevant generic or subgeneric name. Full listing of included taxa along with subgeneric assignments is presented in Table 2. Cladistic analysis is based on 137 characters (see character descriptions for nature of each character Areas occupied by the included taxa indicated by abbreviations following species epithets: Bo, Borneo; EOZ, eastern Australia, i. e. east of the Nullarbor Plain; FP, French Polynesia, Tahiti; HI, Hawaiian Islands, Maui; Jv, Java; LH, Lord Howe Island; NC, New Caledonia; Nf, Norfolk Island; NNZ, North Island of New Zealand; NZ, generally distributed across New Zealand; OZ, generally distributed across Australia; PNG, Papua New Guinea; QOZ, restricted to Queensland, Australia; SNZ, South Island of New Zealand plus Chatham Islands; SP & A, St. Paul and Amsterdam Islands; WOZ, western Australia, i. e. west of the Nullarbor Plain." figureDoi="10.3897/dez.65.21000.figure7" httpUri="https://binary.pensoft.net/fig/179963" pageId="11" pageNumber="12">7</figureCitation>
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) plus
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<taxonomicName id="2DED22012FCBB8BDEA32D7EC6E65E02B" lsidName="M. placens" pageId="11" pageNumber="12" rank="species" species="placens">
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<emphasis id="13C1E01E7DB4802A844683666109921B" italics="true" pageId="11" pageNumber="12">M. placens</emphasis>
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</taxonomicName>
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(Broun). All species are restricted to North Island, New Zealand.
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</paragraph>
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</subSubSection>
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<subSubSection id="6688C89F024355DCBA26EF6C85A22E38" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="new taxonomic status">
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<paragraph id="1BA54DC95F5A936B48D8148D141D4A1D" pageId="11" pageNumber="12">New taxonomic status.</paragraph>
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<paragraph id="5FC0C1E44386F3AB287A608D7F1FB395" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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<bibRefCitation id="5801581AF3463755B638408232DA2DDD" author="Liebherr, JK" journalOrPublisher="(Psydrinae: Meonini New Zealand Entomologist" pageId="51" pageNumber="52" pagination="5 - 22" refId="B65" refString="Liebherr, JK, Marris, JWM, 2009. Revision of the New Zealand species of Mecyclothorax Sharp (Coleoptera: Carabidae: Psydrinae, Mecyclothoracini) and the consequent removal of several species to Meonochilus gen. n. (Psydrinae: Meonini New Zealand Entomologist 32: 5 - 22" title="Revision of the New Zealand species of Mecyclothorax Sharp (Coleoptera: Carabidae: Psydrinae, Mecyclothoracini) and the consequent removal of several species to Meonochilus gen. n." volume="32" year="2009">Liebherr and Marris (2009)</bibRefCitation>
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proposed this clade as a distinct genus based on the great disparity between
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<taxonomicName id="2AC9ACD4B1DAE0835553255427E08993" authorityName="Liebherr & Marris" authorityYear="2009" class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Meonochilus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="939EBE4FFB2378E74785D21F81CFDED3" italics="true" pageId="11" pageNumber="12">Meonochilus</emphasis>
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</taxonomicName>
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spp. and other New Zealand species currently assigned to subgenus
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<taxonomicName id="9A2B17D7A2FF90ABBE2CF515E0A56ADD" authority="Mecyclothorax" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Mecyclothorax)" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subGenus" subGenus="Mecyclothorax">Mecyclothorax Mecyclothorax</taxonomicName>
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. A subsequent cladistic analysis (
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<bibRefCitation id="0597B3806F66D6C4C78404D791BD721D" DOI="https://doi.org/10.3897/zookeys.147.1898" author="Liebherr, JK" journalOrPublisher="from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae ZooKeys" pageId="51" pageNumber="52" pagination="277 - 335" refId="B57" refString="Liebherr, JK, 2011. Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n. from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae ZooKeys 147: 277 - 335, DOI: https://doi.org/10.3897/zookeys.147.1898" title="Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n." url="https://doi.org/10.3897/zookeys.147.1898" volume="147" year="2011">Liebherr 2011</bibRefCitation>
|
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) with taxon representation inadequate to display the phylogenetic structure within
|
||
<taxonomicName id="998918694AFCF6621B45F6E81B7754F4" authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3743A9D8795922A6B9FD02EB60152CBB" italics="true" pageId="11" pageNumber="12">Mecyclothorax</emphasis>
|
||
</taxonomicName>
|
||
affirmed that generic status. The current analysis including a much more comprehensive sampling of
|
||
<taxonomicName id="8CF446671AD5271888C851E28FFC7C9D" authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F1F406850B4FC30F38A2B252EF586683" italics="true" pageId="11" pageNumber="12">Mecyclothorax</emphasis>
|
||
</taxonomicName>
|
||
spp. requires revision of that decision, with
|
||
<taxonomicName id="E4D193CE44935E0435849B6E920EFEDE" authorityName="Liebherr & Marris" authorityYear="2009" class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Meonochilus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Meonochilus</taxonomicName>
|
||
now treated as a subsidiary monophyletic subgenus of
|
||
<taxonomicName id="874B983AC478B6E558B46D6C7F2FCAAD" authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="248244FE01E01C3CA935990438347308" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Strict consensus cladogram of 16 equally most-parsimonious trees derived from cladistic analysis of 65 moriomorphine taxa, including the secondary outgroup Neonomius laevicollis, six primary outgroup taxa in Amblytelus and allied genera, and 58 ingroup terminals; 57 Mecyclothorax species plus two genitalic forms of M. andersoni. Species terminals are labeled with species name and three-letter abbreviation of relevant generic or subgeneric name. Full listing of included taxa along with subgeneric assignments is presented in Table 2. Cladistic analysis is based on 137 characters (see character descriptions for nature of each character Areas occupied by the included taxa indicated by abbreviations following species epithets: Bo, Borneo; EOZ, eastern Australia, i. e. east of the Nullarbor Plain; FP, French Polynesia, Tahiti; HI, Hawaiian Islands, Maui; Jv, Java; LH, Lord Howe Island; NC, New Caledonia; Nf, Norfolk Island; NNZ, North Island of New Zealand; NZ, generally distributed across New Zealand; OZ, generally distributed across Australia; PNG, Papua New Guinea; QOZ, restricted to Queensland, Australia; SNZ, South Island of New Zealand plus Chatham Islands; SP & A, St. Paul and Amsterdam Islands; WOZ, western Australia, i. e. west of the Nullarbor Plain." figureDoi="10.3897/dez.65.21000.figure7" httpUri="https://binary.pensoft.net/fig/179963" pageId="11" pageNumber="12">7</figureCitation>
|
||
). The member species include: 1,
|
||
<taxonomicName id="E5633F77A7708C2F1E08582460E3CEA5" baseAuthorityName="Broun" baseAuthorityYear="1912" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Meonochilus) amplipennis" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="amplipennis" subGenus="Meonochilus">Mecyclothorax (Meonochilus) amplipennis</taxonomicName>
|
||
(Broun) comb. n.; 2,
|
||
<taxonomicName id="B127D566F143375D868D09930ADA7609" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Meonochilus) bellorum" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bellorum" subGenus="Meonochilus">
|
||
<pageBreakToken id="DA66D11068DE1C0BD646825ACB253C0E" pageId="12" pageNumber="13" start="start">Mecyclothorax</pageBreakToken>
|
||
(Meonochilus) bellorum
|
||
</taxonomicName>
|
||
(Liebherr) comb. n.; 3,
|
||
<taxonomicName id="09C765BB86829D2805538351285458FE" baseAuthorityName="Broun" baseAuthorityYear="1923" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Meonochilus) eplicatus" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="eplicatus" subGenus="Meonochilus">Mecyclothorax (Meonochilus) eplicatus</taxonomicName>
|
||
(Broun) comb. n.; 4,
|
||
<taxonomicName id="39E28050DFB8DC59BE787EB0CA7E589C" baseAuthorityName="Broun" baseAuthorityYear="1880" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Meonochilus) placens" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="placens" subGenus="Meonochilus">Mecyclothorax (Meonochilus) placens</taxonomicName>
|
||
(Broun) comb. n.; 5,
|
||
<taxonomicName id="DBBD2808E253B71B5999DF0EC555A146" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Meonochilus) rectus" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="rectus" subGenus="Meonochilus">Mecyclothorax (Meonochilus) rectus</taxonomicName>
|
||
(Liebherr) comb. n.; 6,
|
||
<taxonomicName id="7538DAD61CAA3497823DA10B7EADDD4C" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Meonochilus) spiculatus" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="spiculatus" subGenus="Meonochilus">Mecyclothorax (Meonochilus) spiculatus</taxonomicName>
|
||
(Liebherr) comb. n.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |