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<document id="7F1EF1191EA99301E6019B0439A54FC5" ID-DOI="10.3897/zookeys.1221.135026" ID-ZooBank="0845BEEE-7645-4D58-8FC9-C972F1B2696E" ID-publisher-id="135026" URI-arpha="681FB59F-557D-5A82-B742-EB988338828A" URI-zoobank="http://zoobank.org/0845BEEE-7645-4D58-8FC9-C972F1B2696E" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1735636897127" checkinUser="pensoft" docAuthor="Souma, Jun, Le, Cuong Viet Canh &amp; Pham, Thai-Hong" docDate="2024" docId="E74F79CF237F5458A096E650B3BCDD0E" docLanguage="en" docName="ZooKeys 1221: 363-375" docOrigin="ZooKeys 1221" docSource="https://zookeys.pensoft.net/article/135026/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Bannacoris hyalinus Souma &amp; Le &amp; Pham 2024, comb. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="375" masterDocId="681FB59F557D5A82B742EB988338828A" masterDocTitle="First formal record of the feeding habits of Saileriolidae (Hemiptera, Heteroptera, Pentatomomorpha, Pentatomoidea), with redescription of Bannacoris hyalinus (Schaefer &amp; Ashlock, 1970), comb. nov. endemic to Vietnam" masterLastPageNumber="375" masterPageNumber="363" pageNumber="363" updateTime="1735667628419" updateUser="admin">
<mods:mods id="EA29A5BB23787E4B4E9FFB926236A6EE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E6F140471E5E529A9E69A9D8E1E55AC5">First formal record of the feeding habits of Saileriolidae (Hemiptera, Heteroptera, Pentatomomorpha, Pentatomoidea), with redescription of Bannacoris hyalinus (Schaefer &amp; Ashlock, 1970), comb. nov. endemic to Vietnam</mods:title>
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<mods:namePart id="729787CCD8D6D007CB4E14BE0E6C7306">Souma, Jun</mods:namePart>
<mods:nameIdentifier id="08F862D7C9310081A2FA287ECF7C9F64" type="ORCID">0000-0002-2238-5015</mods:nameIdentifier>
<mods:affiliation id="D6F25D20CF781B0622DA34BC52448A2D">Shirakami Research Center for Environmental Sciences, Faculty of Agriculture and Life Science, Hirosaki University, Aomori, Japan</mods:affiliation>
</mods:name>
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<mods:namePart id="212A64BB93E590FA547D639F6912D2BB">Le, Cuong Viet Canh</mods:namePart>
<mods:nameIdentifier id="AD68AEAE7A7A4F4C670A38ED91935FD2" type="ORCID">0000-0002-1430-6305</mods:nameIdentifier>
<mods:affiliation id="CB701ACD60968D440628A4C400D0B65C">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, Hue, Vietnam</mods:affiliation>
</mods:name>
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<mods:namePart id="AE860C6A0544FFD352D0FCAED88132B1">Pham, Thai-Hong</mods:namePart>
<mods:nameIdentifier id="D4D0D6EE6C2E475087E61A32FA9AC61B" type="ORCID">0000-0002-4763-3679</mods:nameIdentifier>
<mods:affiliation id="379ECFE38D5D97FDA69CB2C015571141">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, Hue, Vietnam &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, Hanoi, Vietnam</mods:affiliation>
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<mods:title id="07500DED3C92EC10F72F66851785EE58">ZooKeys</mods:title>
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<mods:date id="DCDE1A3713FA0D3FAD6D433743468FB9">2024</mods:date>
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<mods:number id="CBDE31220DB55FC262294BDFEA38A5E4">2024-12-31</mods:number>
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<treatment id="E74F79CF237F5458A096E650B3BCDD0E" ID-DOI="http://doi.org/10.5281/zenodo.14581185" ID-Zenodo-Dep="14581185" ID-arpha="E74F79CF-237F-5458-A096-E650B3BCDD0E" LSID="urn:lsid:plazi:treatment:E74F79CF237F5458A096E650B3BCDD0E" httpUri="http://treatment.plazi.org/id/E74F79CF237F5458A096E650B3BCDD0E" scope_order="Hemiptera" scope_superFamily="Pentatomoidea">
<subSubSection id="DCA5D6BA56DD23D9EFA1B33FE162B82F" type="nomenclature">
<paragraph id="787DEB84730D2ADF4A6C4C013EF52CB0">
<taxonomicName id="7D75470FE04F8361B4C15C7045F67C99" authority="(Schaefer &amp; Ashlock, 1970)" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Saileriolidae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="58C63C2B369C7EC2420C6AD7C0A1B467" italics="true">Bannacoris hyalinus</emphasis>
(
<bibRefCitation id="0E092D8E34EC98A8CDDAEDB0B7415B4B" author="Schaefer CW &amp; Ashlock PD" issue="3" journalOrPublisher="Pacific Insects" pagination="629-639" refId="B11" refString="Schaefer CW, Ashlock PD (1970) A new genus and new species of Saileriolinae (Hemiptera: Urostylidae). Pacific Insects 12 (3): 629 - 639." title="A new genus and new species of Saileriolinae (Hemiptera: Urostylidae)." volume="12" year="1970">Schaefer &amp; Ashlock, 1970</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="B2ABFE1D7175518D5586AAE4BA72B951" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="3751AA4E998E1D4675D1B5964909BFB3" type="description">
<paragraph id="0EDBD8E7B7442E2078A42EE67C9A2833">
<figureCitation id="E4F07A48E9182A7A3F3E4A14AC00BABC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Bannacoris hyalinus comb. nov. from Vietnam. A male B female." figureDoi="10.3897/zookeys.1221.135026.figure1" httpUri="https://binary.pensoft.net/fig/1210690">Figs 1 A, B</figureCitation>
,
<figureCitation id="23212B08B082F0420BC755F7C7413744" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Detailed morphological images of Bannacoris hyalinus comb. nov. from Vietnam A head, pronotum, and scutellum, dorsal view B head, cranial view C head and thorax, ventral view D forewing, dorsal view E male terminalia, ventral view F female terminalia, ventral view G phallus, ventral view." figureDoi="10.3897/zookeys.1221.135026.figure2" httpUri="https://binary.pensoft.net/fig/1210691">2 A G</figureCitation>
,
<figureCitation id="8BEF74FD2BC52BEC9AD13687210C9DC1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of Bannacoris hyalinus comb. nov. from Vietnam A genital capsule, dorsal view B paramere, dorsal view C paramere ventral view D female terminalia, ventral view. Abbreviations: abs 6, abs 7, abdominal sternites VI, VII; lt 9, laterotergite IX; vmp, ventromedian process." figureDoi="10.3897/zookeys.1221.135026.figure3" httpUri="https://binary.pensoft.net/fig/1210692">3 A D</figureCitation>
,
<figureCitation id="E54785D21B7D5476DA9129FC5FF06F52" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Photographs regarding Bannacoris hyalinus comb. nov. from B ạch Mã National Park, Th ừa Thiên Hu ế Province, Vietnam A living adult B living nymph C host plant (Litsea sp.) D surrounding habitat." figureDoi="10.3897/zookeys.1221.135026.figure4" httpUri="https://binary.pensoft.net/fig/1210693">4 A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8E7513AE324544AE880E0F3B264227B0" type="reference_group">
<paragraph id="C68A05B021A1A2373788303D7601636D">
<materialsCitation id="D338A82D1B0D9307D29D16400ECC3299" country="Vietnam" county="Di Linh District" elevation="990" location="Dilinh" municipality="Bao Thuan" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong Province" typeStatus="holotype">
<treatmentCitationGroup id="40400CF2A3AF8D2E1833C778AE08598B">
<treatmentCitation id="877EC8C457A6C80F414356FFF715F107" author="Schaefer CW &amp; Ashlock PD" page="631" year="1970">
<taxonomicName id="2669B76C01AA0D8894E8D8BA7CF900F7" authority="Schaefer &amp; Ashlock, 1970: 631" authorityName="Schaefer &amp; Ashlock" authorityPageNumber="631" authorityYear="1970" class="Insecta" family="Urostylididae" genus="Saileriola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalina">
<originalDetermination id="FE0B6CB7754B3A6DD3AD02DC07925EBF">
<emphasis id="C2013B8380E0A2C036B023F992019389" italics="true">Saileriola hyalina</emphasis>
<comment id="C0440EA855066CF086B0CF2B56CE9335">
<bibRefCitation id="1AA9E3F7E79D3A0458CB2AABFDCFEA18" author="Schaefer CW &amp; Ashlock PD" issue="3" journalOrPublisher="Pacific Insects" pagination="629-639" refId="B11" refString="Schaefer CW, Ashlock PD (1970) A new genus and new species of Saileriolinae (Hemiptera: Urostylidae). Pacific Insects 12 (3): 629 - 639." title="A new genus and new species of Saileriolinae (Hemiptera: Urostylidae)." volume="12" year="1970">Schaefer &amp; Ashlock, 1970: 631</bibRefCitation>
.
<typeStatus id="D81B7C66FDBB5B1C046545D66DA9F3F4" type="holotype">Holotype</typeStatus>
:
<specimenCount id="B1EE20270BB7B4E6012253D0BC85E2BB" count="1" type="male"></specimenCount>
,
<collectingCountry id="315539C21E3EA6BEA68297B6F5733D71" name="Vietnam">Vietnam</collectingCountry>
:
<locationDeviation id="3F34F3E63216051CCE1B4099E51FD486">
<quantity id="DAB2D713A6F4443DBCC51FA58A695C71" metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
SE of
</locationDeviation>
<location id="CD715F3AB9EB3E93FAF5C20D8098B9CD" LSID="urn:lsid:plazi:treatment:E74F79CF237F5458A096E650B3BCDD0E:CD715F3AB9EB3E93FAF5C20D8098B9CD" country="Vietnam" county="Di Linh District" municipality="B ao Thu an" name="Dilinh" stateProvince="Lam D ong Province">Dilinh</location>
(Djiring),
<quantity id="BF04BA5C52222530578BE19A8D0A39C4" metricMagnitude="2" metricUnit="m" metricValue="9.9" unit="m" value="990.0">
<elevation id="F7F4C092D9D7CB24CFF841852FAC66DC" metricMagnitude="2" metricUnit="m" metricValue="9.9" unit="m" value="990.0">990 m</elevation>
</quantity>
[=
<collectingRegion id="A2C335891416D8A8B90C75EE21774181">Lâm Đ ồng Province</collectingRegion>
,
<collectingCounty id="5C71128FFE991169B16CA7164DEE8F2F">Di Linh District</collectingCounty>
,
<collectingMunicipality id="E2FB97B7F4E66C37896D9B839A277E6E">B ảo Thu ận</collectingMunicipality>
?]; Bernice P. Bishop Museum, Honolulu, Hawaii, USA.
</comment>
</originalDetermination>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4MAE" type="material">
<paragraph id="DCAF08397DF8186DC1F15EBD4E4B13EA">
<heading id="74527F856046A8DEE464232C7ADC3BB7" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="690A057F3DFF94CF2C08722C195016D9">
<emphasis id="A677FD397A99FC949E4711D5188F94ED" bold="true">Non-types</emphasis>
<materialsCitation id="C396693FED2B0CC2BD9A50175737F0D9" collectingDate="2024-06-06" collectionCode="VNMN" collectorName="J. Souma." country="Vietnam" county="Phu L oc District" latitude="16.192778" location="Du ong mon D o Quyen" longLatPrecision="21" longitude="107.84778" municipality="ach Ma National Park" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="Th ua Thien Hu e Province">
(
<specimenCount id="5943161FA8A9E99A3C54DAD352F64874" count="5" type="male">5 ♂♂</specimenCount>
<specimenCount id="CA422795BA48C1A4EE329BDFEB9DC1AE" count="4" type="female">4 ♀♀</specimenCount>
,
<named-content id="614A693EC8481B4A1FA4683D806780D3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/vietnam-national-museum-nature" xlink_title="Vietnam National Museum of Nature">
<collectionCode id="1B0DCEEA76CA02D71A717754BD8D0097" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" type="">VNMN</collectionCode>
</named-content>
):
<collectingCountry id="E094BE51E852CC04597A83067545AE04" name="Vietnam">Vietnam</collectingCountry>
<collectingRegion id="1C4798B9F2DB066DC43B7AD19EE67126">Th ừa Thiên Hu ế Province</collectingRegion>
,
<collectingCounty id="5FA248ED393798C822B2E64FEF91597A">Phú L ộc District</collectingCounty>
, B
<collectingMunicipality id="D65794907A9AD01D85AF37B978FD4289">ạch Mã National Park</collectingMunicipality>
,
<collectingMunicipality id="D686E3A5ACC17F23FDB60C69AEEF4E32">L ộc Trì</collectingMunicipality>
,
<location id="1EE6489B612E2EB77B58858570F90A10" LSID="urn:lsid:plazi:treatment:E74F79CF237F5458A096E650B3BCDD0E:1EE6489B612E2EB77B58858570F90A10" country="Vietnam" county="Phu L oc District" latitude="16.192778" longLatPrecision="21" longitude="107.84778" municipality="ach Ma National Park" name="Du ong mon D o Quyen" stateProvince="Th ua Thien Hu e Province">Đư ờng mòn Đ ỗ Quyên</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMNAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.847778,16.192778]}">
<geoCoordinate id="3055C1B8B9B7B8E025E3B63B0FB1B04F" degrees="16" direction="north" minutes="11" orientation="latitude" precision="15" seconds="34" value="16.192778">16°11'34&quot;N</geoCoordinate>
,
<geoCoordinate id="086245B879D8E5C38D092CF1BE8F1225" degrees="107" direction="east" minutes="50" orientation="longitude" precision="15" seconds="52" value="107.84778">107°50'52&quot;E</geoCoordinate>
</named-content>
;
<date id="E451BA5FB79FC582ECF9066F9A237602" value="2024-06-06">
<collectingDate id="67931E0509ACC31B57162421773D636C" value="2024-06-06">6.vi.2024</collectingDate>
</date>
; leg.
<collectorName id="47D2B0C4D9339BECF70DFD62AB634CDC">J. Souma.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERNAE" type="diagnosis">
<paragraph id="AF1A176350E39264DFE75F8F1BDD6862">
<heading id="9D1AB34088BA1984BE67450E084DCB71" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="F8A925869A80A86BB9B4C844E662B54E">
<taxonomicName id="BB0A0A7B890A1BEF46FAD2E829478919" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="A4D7BC6E721DDB460063BA007255FC09" italics="true">Bannacoris hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5DAA1BDECED3722EFE68AF996098BF5" rank="species">comb. nov.</taxonomicNameLabel>
can be distinguished from the only other congener,
<taxonomicName id="683945227747C7BEF1B68D0362133379" authorityName="Hsiao" authorityYear="1964" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="arboreus">
<emphasis id="D2D58919CD960CA30F2ECF03B6246CF2" italics="true">B. arboreus</emphasis>
</taxonomicName>
, by the following characters: head, pronotum, and scutellum mostly yellowish brown (Figs
<figureCitation id="73316149A52725B2B451DC858F6F4BCB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Bannacoris hyalinus comb. nov. from Vietnam. A male B female." figureDoi="10.3897/zookeys.1221.135026.figure1" httpUri="https://binary.pensoft.net/fig/1210690">1 A, B</figureCitation>
,
<figureCitation id="2F1F1CCC4E1419E2B7303072CDCEC1DD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Detailed morphological images of Bannacoris hyalinus comb. nov. from Vietnam A head, pronotum, and scutellum, dorsal view B head, cranial view C head and thorax, ventral view D forewing, dorsal view E male terminalia, ventral view F female terminalia, ventral view G phallus, ventral view." figureDoi="10.3897/zookeys.1221.135026.figure2" httpUri="https://binary.pensoft.net/fig/1210691">2 A</figureCitation>
) (mostly reddish to dark brown in
<taxonomicName id="3860F435B925F936AC8DCED3D7D9E6DB" authorityName="Hsiao" authorityYear="1964" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="arboreus">
<emphasis id="0F9586FB997270F5AFF70CDD7A7A2800" italics="true">B. arboreus</emphasis>
</taxonomicName>
); corium of forewing mostly hyaline (Fig.
<figureCitation id="175B7372A85FE744F131A23E085A8B54" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Detailed morphological images of Bannacoris hyalinus comb. nov. from Vietnam A head, pronotum, and scutellum, dorsal view B head, cranial view C head and thorax, ventral view D forewing, dorsal view E male terminalia, ventral view F female terminalia, ventral view G phallus, ventral view." figureDoi="10.3897/zookeys.1221.135026.figure2" httpUri="https://binary.pensoft.net/fig/1210691">2 D</figureCitation>
) (reddish to dark brown in middle part in
<taxonomicName id="8667B96CFF5E982F34E2A51ACA2E8A63" authorityName="Hsiao" authorityYear="1964" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="arboreus">
<emphasis id="E520C90B9CC928654125982C79D50FA3" italics="true">B. arboreus</emphasis>
</taxonomicName>
), not punctate in an area enclosed by Sc (subcostal) vein and medial furrow (entirely punctate in
<taxonomicName id="22B766DEF3E1952CE74203E2E03B1AA7" authorityName="Hsiao" authorityYear="1964" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="arboreus">
<emphasis id="47ADFA355D5621035FCC167EEE1A32F0" italics="true">B. arboreus</emphasis>
</taxonomicName>
); dorsolateral process of genital capsule undeveloped (Figs
<figureCitation id="F789EBD3448B861C9589B02AC9D4F751" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Detailed morphological images of Bannacoris hyalinus comb. nov. from Vietnam A head, pronotum, and scutellum, dorsal view B head, cranial view C head and thorax, ventral view D forewing, dorsal view E male terminalia, ventral view F female terminalia, ventral view G phallus, ventral view." figureDoi="10.3897/zookeys.1221.135026.figure2" httpUri="https://binary.pensoft.net/fig/1210691">2 E</figureCitation>
,
<figureCitation id="C438A0A134C72E11449046A5DBB99DFE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of Bannacoris hyalinus comb. nov. from Vietnam A genital capsule, dorsal view B paramere, dorsal view C paramere ventral view D female terminalia, ventral view. Abbreviations: abs 6, abs 7, abdominal sternites VI, VII; lt 9, laterotergite IX; vmp, ventromedian process." figureDoi="10.3897/zookeys.1221.135026.figure3" httpUri="https://binary.pensoft.net/fig/1210692">3 A</figureCitation>
) (protruding posteriad in
<taxonomicName id="EC1952BFD492F3BB6DF0D6FE8BBA1076" authorityName="Hsiao" authorityYear="1964" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="arboreus">
<emphasis id="36C64656D60AB1B11E915A0DBF527884" italics="true">B. arboreus</emphasis>
</taxonomicName>
); and ventromedian process concave in posterior margin (gently curved outward in
<taxonomicName id="92B8A213CCC54349C25675E5FF6F6C2C" authorityName="Hsiao" authorityYear="1964" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="arboreus">
<emphasis id="AB4BB348D3861544FE14776D172CD611" italics="true">B. arboreus</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="93A7F7316EB36BE6A8D503706906AF43" ID-DOI="10.3897/zookeys.1221.135026.figure1" ID-arpha="775B7619-0B93-5E42-8A36-73CF40CD689B" httpUri="https://binary.pensoft.net/fig/1210690" startId="F1">
<paragraph id="CFE754226EC6472A3C5FB312AE1BAAEA">
<label id="F6619090238FC32099916D3D7A81ED3A">Figure 1.</label>
</paragraph>
<paragraph id="95D70402F6C078225394425EB609C8C2">
Dorsal habitus of
<taxonomicName id="0C6DAE45B7345A7597BB14DBA8CED615" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="C4200CF09B675C0ED6B59C638D8078E2" italics="true">Bannacoris hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1583122A180ECA453E35EF3E55E90E3C" rank="species">comb. nov.</taxonomicNameLabel>
from Vietnam.
<emphasis id="3D62623DE86B855753EEC678C7BC909D" bold="true">A</emphasis>
male
<emphasis id="B5578060719AF1BF1771D64984E9485C" bold="true">B</emphasis>
female.
</paragraph>
</caption>
<caption id="ABE740B2199376F7222CAD973482589D" ID-DOI="10.3897/zookeys.1221.135026.figure2" ID-arpha="CD8AD80B-039E-5EEF-A961-64577AFD4D1A" httpUri="https://binary.pensoft.net/fig/1210691" startId="F2">
<paragraph id="46CFAB7BC5CBD5C8D5D387BC87D5AFAF">
<label id="307E3E611AB460BD8C237231D510759B">Figure 2.</label>
</paragraph>
<paragraph id="F9DA3F63FCB5FC12E37516BB83E5C765">
Detailed morphological images of
<taxonomicName id="F4F530462E1F7E41AB30F6CFCB895B9E" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="6C73968D049613284393001BE9184CB6" italics="true">Bannacoris hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAB49F0D630F64CC656A5E187BD356FF" rank="species">comb. nov.</taxonomicNameLabel>
from Vietnam
<emphasis id="DD38ADFBAC8F2BF73A6B31F8E40B6422" bold="true">A</emphasis>
head, pronotum, and scutellum, dorsal view
<emphasis id="C8BD9AA0D682FC8BC4D9F82DFC4E848A" bold="true">B</emphasis>
head, cranial view
<emphasis id="C1EA0BE54F3D6F91CD421EBB5CA1C467" bold="true">C</emphasis>
head and thorax, ventral view
<emphasis id="1581EE88A90531435E42DB0072299C74" bold="true">D</emphasis>
forewing, dorsal view
<emphasis id="568855FF43EF2A024E91BAFC185F3B75" bold="true">E</emphasis>
male terminalia, ventral view
<emphasis id="D13A55466D8720ACB279AA7469E243D0" bold="true">F</emphasis>
female terminalia, ventral view
<emphasis id="F7544577C2955DCDB92B00B6AD4842E8" bold="true">G</emphasis>
phallus, ventral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E1SAE" type="redescription">
<paragraph id="37224605B1E52FB04562B1BD9C913410">
<heading id="24551349ABE6D33929BB80506BF352F2" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="EF7FCAF7C71BF5770AF5EB35431048FF">
Most parts of head, antennomeres I IV, pronotum except for anterior and lateral margins, ventral surface of thoracic parts, most parts of scutellum, legs except for claws, and abdomen yellowish brown; antennomere V, compound eye, ocellus, anterior margin of scutellum, Sc (subcostal) vein of forewing, claws, and punctures on body dark brown; anterior and lateral margins of pronotum, and outer part of semi-elliptical ridge in anteromedial part of scutellum whitish brown; forewing except for Sc vein and punctures hyaline; setae on body yellowish (Figs
<figureCitation id="53FC08E42F3EBA1602F08D3CCF64010E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Bannacoris hyalinus comb. nov. from Vietnam. A male B female." figureDoi="10.3897/zookeys.1221.135026.figure1" httpUri="https://binary.pensoft.net/fig/1210690">1 A, B</figureCitation>
,
<figureCitation id="B969821A9B7C2FDADB8DCA0FD69382EC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Detailed morphological images of Bannacoris hyalinus comb. nov. from Vietnam A head, pronotum, and scutellum, dorsal view B head, cranial view C head and thorax, ventral view D forewing, dorsal view E male terminalia, ventral view F female terminalia, ventral view G phallus, ventral view." figureDoi="10.3897/zookeys.1221.135026.figure2" httpUri="https://binary.pensoft.net/fig/1210691">2 A F</figureCitation>
,
<figureCitation id="6611BFB659D6B94F73B2757FD0C5B505" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Photographs regarding Bannacoris hyalinus comb. nov. from B ạch Mã National Park, Th ừa Thiên Hu ế Province, Vietnam A living adult B living nymph C host plant (Litsea sp.) D surrounding habitat." figureDoi="10.3897/zookeys.1221.135026.figure4" httpUri="https://binary.pensoft.net/fig/1210693">4 A, B</figureCitation>
).
</paragraph>
<paragraph id="FF22EEAC01BD2D8704C66E86BE39E6BF">
Body (Fig.
<figureCitation id="35599FA33A42681E3D14CF9604FA64CB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Bannacoris hyalinus comb. nov. from Vietnam. A male B female." figureDoi="10.3897/zookeys.1221.135026.figure1" httpUri="https://binary.pensoft.net/fig/1210690">1 A, B</figureCitation>
) ovate, 1.51.6 times as long as maximum width across abdomen. Head (Fig.
<figureCitation id="507260A91803FBAD564E3387CF053F95" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Detailed morphological images of Bannacoris hyalinus comb. nov. from Vietnam A head, pronotum, and scutellum, dorsal view B head, cranial view C head and thorax, ventral view D forewing, dorsal view E male terminalia, ventral view F female terminalia, ventral view G phallus, ventral view." figureDoi="10.3897/zookeys.1221.135026.figure2" httpUri="https://binary.pensoft.net/fig/1210691">2 A C</figureCitation>
) declivent, mostly punctate, longer than maximum width across compound eyes in cranial view, with a median sulcus on vertex, sparsely bearing minute setae. Antenniferous tubercle annulate, placed anterior to compound eye. Clypeus distinctly surpassing mandibular plate at both apices. Compound eye round, separated from anterior margin of pronotum. A pair of ocelli placed along midline of vertex, closer together than a diameter of ocellus. Antenna smooth on surface; antennomere I longest among antennomeres, more than 1.5 times as long as antennomere II, bearing minute setae throughout its length; antennomere II longer than antennomere IV, bearing minute setae throughout its length; antennomere III shortest among antennomeres, bearing minute setae throughout its length; antennomere IV as long as antennomere V, bearing minute and long setae throughout its length; antennomere V bearing minute and long setae throughout its length. Labium reaching anterior part of abdominal sternite III. Buccula semi-elliptical in lateral view, highest in middle part.
</paragraph>
<paragraph id="C5C47C4B9D9C8D7A691492C3D984D5E9">
Thorax (Figs
<figureCitation id="E90769F2B4D4FBD9726ADDFC80549FAD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Bannacoris hyalinus comb. nov. from Vietnam. A male B female." figureDoi="10.3897/zookeys.1221.135026.figure1" httpUri="https://binary.pensoft.net/fig/1210690">1 A, B</figureCitation>
,
<figureCitation id="5DBBDAF3456ECC1AB0AAC9A609B870FD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Detailed morphological images of Bannacoris hyalinus comb. nov. from Vietnam A head, pronotum, and scutellum, dorsal view B head, cranial view C head and thorax, ventral view D forewing, dorsal view E male terminalia, ventral view F female terminalia, ventral view G phallus, ventral view." figureDoi="10.3897/zookeys.1221.135026.figure2" httpUri="https://binary.pensoft.net/fig/1210691">2 A, C</figureCitation>
) mostly punctate. Pronotum trapezoidal in dorsal view, shorter than its maximum width, punctate except for callus; lateral margin serrate in anterior part, without distinct spine, bearing minute setae throughout its length; humeral angle rounded. Scutellum triangular, shorter than its maximum width, semi-elliptically raised in anteromedial part, punctate except for midline and semi-elliptical ridge. Forewing (Fig.
<figureCitation id="0D260DB7B42BA5A921B44B5F0CAB33B7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Detailed morphological images of Bannacoris hyalinus comb. nov. from Vietnam A head, pronotum, and scutellum, dorsal view B head, cranial view C head and thorax, ventral view D forewing, dorsal view E male terminalia, ventral view F female terminalia, ventral view G phallus, ventral view." figureDoi="10.3897/zookeys.1221.135026.figure2" httpUri="https://binary.pensoft.net/fig/1210691">2 D</figureCitation>
) oblong; anterior margin gently curved outward; clavus shorter than membrane, with 2 rows of punctures throughout its length; corium punctate except in an area enclosed by Sc vein and medial furrow, bearing minute setae in anterior part; membrane provided with several indistinct longitudinal veins; Sc vein, basal part of Cu (cubital) vein, and claval and medial furrows distinct. Epimera and episterna punctate except metepimeron. Sterna smooth on surface. Legs smooth on surface; femora and tibiae cylindrical, bearing setae throughout their length.
</paragraph>
<paragraph id="7310EE952166CCE0FA6AF5045A8E3F08">
Abdomen (Figs
<figureCitation id="398241A789949607A8B3E15435FB7822" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Bannacoris hyalinus comb. nov. from Vietnam. A male B female." figureDoi="10.3897/zookeys.1221.135026.figure1" httpUri="https://binary.pensoft.net/fig/1210690">1 A, B</figureCitation>
,
<figureCitation id="BBFCFBC877CDB7D41664DA25531D50FC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Detailed morphological images of Bannacoris hyalinus comb. nov. from Vietnam A head, pronotum, and scutellum, dorsal view B head, cranial view C head and thorax, ventral view D forewing, dorsal view E male terminalia, ventral view F female terminalia, ventral view G phallus, ventral view." figureDoi="10.3897/zookeys.1221.135026.figure2" httpUri="https://binary.pensoft.net/fig/1210691">2 D F</figureCitation>
) longer than combined length of head and pronotum; posterior margin of sternite VI concave in male, undulate in female; sternite VII concave in posterior margin of male, with a longitudinal cleft in female. Genital capsule (Fig.
<figureCitation id="CB55D34B8E5F3655F6E2D31AE34FEC0F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of Bannacoris hyalinus comb. nov. from Vietnam A genital capsule, dorsal view B paramere, dorsal view C paramere ventral view D female terminalia, ventral view. Abbreviations: abs 6, abs 7, abdominal sternites VI, VII; lt 9, laterotergite IX; vmp, ventromedian process." figureDoi="10.3897/zookeys.1221.135026.figure3" httpUri="https://binary.pensoft.net/fig/1210692">3 A</figureCitation>
) elliptical in dorsal and ventral views, smooth on surface, bearing setae in posterior part; lateral margin gently curved outward; dorsolateral process undeveloped; ventromedian process widened apically, concave in posterior margin. Paramere (Fig.
<figureCitation id="879751CC7FA070C00F59C33E9068BF6F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of Bannacoris hyalinus comb. nov. from Vietnam A genital capsule, dorsal view B paramere, dorsal view C paramere ventral view D female terminalia, ventral view. Abbreviations: abs 6, abs 7, abdominal sternites VI, VII; lt 9, laterotergite IX; vmp, ventromedian process." figureDoi="10.3897/zookeys.1221.135026.figure3" httpUri="https://binary.pensoft.net/fig/1210692">3 B, C</figureCitation>
) elongate; crown widened apically, bearing three setae from cuticular sockets along outer margin of dorsum; neck constricted; stem widened apically. Phallus (Fig.
<figureCitation id="D96499C177EA3D10FF29D4B984EEA6A1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Detailed morphological images of Bannacoris hyalinus comb. nov. from Vietnam A head, pronotum, and scutellum, dorsal view B head, cranial view C head and thorax, ventral view D forewing, dorsal view E male terminalia, ventral view F female terminalia, ventral view G phallus, ventral view." figureDoi="10.3897/zookeys.1221.135026.figure2" httpUri="https://binary.pensoft.net/fig/1210691">2 G</figureCitation>
) oblong; basal plate and phallotheca coriaceous; conjunctiva with two pairs of sclerites. Female terminalia (Fig.
<figureCitation id="BEE2D215784EAB8FAA5B3573AC6EBF5D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of Bannacoris hyalinus comb. nov. from Vietnam A genital capsule, dorsal view B paramere, dorsal view C paramere ventral view D female terminalia, ventral view. Abbreviations: abs 6, abs 7, abdominal sternites VI, VII; lt 9, laterotergite IX; vmp, ventromedian process." figureDoi="10.3897/zookeys.1221.135026.figure3" httpUri="https://binary.pensoft.net/fig/1210692">3 D</figureCitation>
) semicircular in ventral view, protruding posteriad in middle part; laterotergite VIII reduced; valvifer VIII reduced; laterotergite IX rounded in outer margin.
</paragraph>
<caption id="5F5680C58A9CB226B8AEB3025F0003A6" ID-DOI="10.3897/zookeys.1221.135026.figure3" ID-arpha="BA426162-A79B-5B7D-98D4-404AE43AEBD0" httpUri="https://binary.pensoft.net/fig/1210692" startId="F3">
<paragraph id="C75E3BF0A015ADBDF187167C77B2CC53">
<label id="C5F0AC82E9555D8A4F36FF8DAD6E0C7E">Figure 3.</label>
</paragraph>
<paragraph id="5EF65A36719894B77E8A57F06EE180FB">
Line drawings of
<taxonomicName id="BB4E7A268EB8844FB2201747325BF2E4" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="9FCA4E20B9703C9B21D77BC53C4FF6BF" italics="true">Bannacoris hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E1D33E18750E22E67C6632C786A2CBA4" rank="species">comb. nov.</taxonomicNameLabel>
from Vietnam
<emphasis id="548053B12BA87A581EA9FCA41F05CCFE" bold="true">A</emphasis>
genital capsule, dorsal view
<emphasis id="B2136F4525584A9EAEE3CF819BB20BF8" bold="true">B</emphasis>
paramere, dorsal view
<emphasis id="E92B9AEA0C9A6F9728B73014D657444F" bold="true">C</emphasis>
paramere ventral view
<emphasis id="3A78F5B826040B2D978769DBADAAA7C6" bold="true">D</emphasis>
female terminalia, ventral view. Abbreviations: abs 6, abs 7, abdominal sternites VI, VII; lt 9, laterotergite IX; vmp, ventromedian process.
</paragraph>
</caption>
<paragraph id="28186F06589AE19BA9A2A90FCC9312E0">
Measurements (male:
<emphasis id="3871C62ACB3F604CD7C4686B823DBAC7" italics="true">n</emphasis>
= 5; female:
<emphasis id="E1AE51A2B0DDF0E6A75071086E12AEFA" italics="true">n</emphasis>
= 4). Body length with forewing
<quantity id="243F7E2D843E1616740AEE8716E9CDDF" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" unit="mm" value="3.65" valueMax="3.8" valueMin="3.5">3.53.8 mm</quantity>
in male and 3.94.0 mm in female, maximum width across forewings
<quantity id="BA1517DD8E85AAD4D2A86FEE90395CC5" metricMagnitude="-3" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" unit="mm" value="2.3499999999999996" valueMax="2.4" valueMin="2.3">2.32.4 mm</quantity>
in male and
<quantity id="C348DDEA7A3D8190F6049304FE183FAA" metricMagnitude="-3" metricUnit="m" metricValue="2.5" unit="mm" value="2.5">2.5 mm</quantity>
in female; head length in cranial view
<quantity id="D0619141A2B660BEA7B4BDBBACF50CA0" metricMagnitude="-4" metricUnit="m" metricValue="9.0" unit="mm" value="0.9">0.9 mm</quantity>
in both sexes, maximum width across compound eyes
<quantity id="45314AC640C846A4F23342C94F6E3EDC" metricMagnitude="-4" metricUnit="m" metricValue="8.0" unit="mm" value="0.8">0.8 mm</quantity>
in both sexes; length of antennomeres I V in both sexes
<quantity id="F2016DE8BF7066A6C564845F7A85E88F" metricMagnitude="-3" metricUnit="m" metricValue="1.5" unit="mm" value="1.5">1.5 mm</quantity>
,
<quantity id="E0210480C34AB5ADBC3A7D408D03DDA8" metricMagnitude="-4" metricUnit="m" metricValue="8.0" unit="mm" value="0.8">0.8 mm</quantity>
,
<quantity id="BC8937D2EFDF81F32525FE292F9B7B67" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
,
<quantity id="367A470FF5BE815FE3D1E4863797FA2E" metricMagnitude="-4" metricUnit="m" metricValue="6.0" unit="mm" value="0.6">0.6 mm</quantity>
, and
<quantity id="87E4B629404616F4DF32FFD495B74BDD" metricMagnitude="-4" metricUnit="m" metricValue="6.0" unit="mm" value="0.6">0.6 mm</quantity>
, respectively; length of labial segments I IV in both sexes
<quantity id="91E1DA57F3153D8D452218BA7B632D08" metricMagnitude="-4" metricUnit="m" metricValue="3.0" unit="mm" value="0.3">0.3 mm</quantity>
,
<quantity id="B91A78F2C54CD4F68851DDAE80F51D8B" metricMagnitude="-4" metricUnit="m" metricValue="3.0" unit="mm" value="0.3">0.3 mm</quantity>
,
<quantity id="94CA6345FACAD28EAC6A5FFFF5C77E13" metricMagnitude="-4" metricUnit="m" metricValue="3.0" unit="mm" value="0.3">0.3 mm</quantity>
, and
<quantity id="6BD01FBC563EFE1EA88609C380E71206" metricMagnitude="-4" metricUnit="m" metricValue="4.0" unit="mm" value="0.4">0.4 mm</quantity>
, respectively; pronotum length
<quantity id="71C73259211E0864E945CDE8BD099F29" metricMagnitude="-4" metricUnit="m" metricValue="7.0" unit="mm" value="0.7">0.7 mm</quantity>
in both sexes, maximum width
<quantity id="70508CC22D37E8394709166A6D1D0FCA" metricMagnitude="-3" metricUnit="m" metricValue="1.8" unit="mm" value="1.8">1.8 mm</quantity>
in male and 1.92.0 mm in female; scutellum length
<quantity id="D8B1488DDD3BFB0A0F16D8F1BE54397B" metricMagnitude="-4" metricUnit="m" metricValue="8.0" unit="mm" value="0.8">0.8 mm</quantity>
in male and
<quantity id="AE9FDC59E00BD8808BF8B72ED9E1FDFE" metricMagnitude="-4" metricUnit="m" metricValue="9.0" unit="mm" value="0.9">0.9 mm</quantity>
in female, maximum width 1.0 mm in male and
<quantity id="96AF5C322821DC33725A64767C7D452C" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
in female; forewing length
<quantity id="370CF791FF072122DF4AF3A43434A1F4" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" unit="mm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.82.9 mm</quantity>
in male and 3.0
<quantity id="197D556626AC4B852FB993EA695D4311" metricMagnitude="-3" metricUnit="m" metricValue="3.1" unit="mm" value="3.1">3.1 mm</quantity>
in female, maximum width
<quantity id="AB66F76EE3A3E316C90534061DEC199F" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
in male and
<quantity id="B313F294949D849371B79BC9A606306E" metricMagnitude="-3" metricUnit="m" metricValue="1.3" unit="mm" value="1.3">1.3 mm</quantity>
in female.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEWAE" type="remarks">
<paragraph id="F0A0DAE63F6D4ECEE830B2FAC057C883">
<heading id="36C6D63DE07348599F7991897E29CCB8" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="2DB7F0FFA0A207DFBB733D1925558374">
The
<specimenCount id="F9ECA8CDECAFC149E98DB7BAF022A1F3" count="9" type="generic">nine specimens</specimenCount>
recorded above (Fig.
<figureCitation id="86F6A7A91CAACA3B3C7CE6B397EDB699" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Bannacoris hyalinus comb. nov. from Vietnam. A male B female." figureDoi="10.3897/zookeys.1221.135026.figure1" httpUri="https://binary.pensoft.net/fig/1210690">1 A, B</figureCitation>
) matched well with the original description and illustrations of
<taxonomicName id="0728B5A021923FA99205E78312C8B2CE" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="DFACE6B7EE2F88D356C740E5F74E9B61" italics="true">Bannacoris hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A105A4169BA13A86941086C2F651F1D" rank="species">comb. nov.</taxonomicNameLabel>
(
<bibRefCitation id="F009EB9DC59641F07CD425CB695BDEB3" author="Schaefer and Ashlock" firstAuthor="Schaefer" issue="3" journalOrPublisher="Pacific Insects" pagination="629-639" refId="B11" refString="Schaefer CW, Ashlock PD (1970) A new genus and new species of Saileriolinae (Hemiptera: Urostylidae). Pacific Insects 12 (3): 629639." title="A new genus and new species of Saileriolinae (Hemiptera: Urostylidae)." volume="12" year="1970">Schaefer and Ashlock 1970</bibRefCitation>
) in terms of morphological characteristics, especially the structure of the head (Fig.
<figureCitation id="1BA6B91811275949697784062FE3DE04" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Detailed morphological images of Bannacoris hyalinus comb. nov. from Vietnam A head, pronotum, and scutellum, dorsal view B head, cranial view C head and thorax, ventral view D forewing, dorsal view E male terminalia, ventral view F female terminalia, ventral view G phallus, ventral view." figureDoi="10.3897/zookeys.1221.135026.figure2" httpUri="https://binary.pensoft.net/fig/1210691">2 B</figureCitation>
) and the shape of the genital capsule (Figs
<figureCitation id="956AE7088E59C6022274E92C47F51290" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Detailed morphological images of Bannacoris hyalinus comb. nov. from Vietnam A head, pronotum, and scutellum, dorsal view B head, cranial view C head and thorax, ventral view D forewing, dorsal view E male terminalia, ventral view F female terminalia, ventral view G phallus, ventral view." figureDoi="10.3897/zookeys.1221.135026.figure2" httpUri="https://binary.pensoft.net/fig/1210691">2 E</figureCitation>
,
<figureCitation id="F73CD15297204B571DC7C655803B381A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of Bannacoris hyalinus comb. nov. from Vietnam A genital capsule, dorsal view B paramere, dorsal view C paramere ventral view D female terminalia, ventral view. Abbreviations: abs 6, abs 7, abdominal sternites VI, VII; lt 9, laterotergite IX; vmp, ventromedian process." figureDoi="10.3897/zookeys.1221.135026.figure3" httpUri="https://binary.pensoft.net/fig/1210692">3 A</figureCitation>
) and paramere (Fig.
<figureCitation id="6331AE748A128D991999209783146002" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Line drawings of Bannacoris hyalinus comb. nov. from Vietnam A genital capsule, dorsal view B paramere, dorsal view C paramere ventral view D female terminalia, ventral view. Abbreviations: abs 6, abs 7, abdominal sternites VI, VII; lt 9, laterotergite IX; vmp, ventromedian process." figureDoi="10.3897/zookeys.1221.135026.figure3" httpUri="https://binary.pensoft.net/fig/1210692">3 B, C</figureCitation>
). Therefore, we identified the specimens studied as
<taxonomicName id="F0D9FC896525B1DBE3C73E9C362EFC15" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="CB99AE70C5CE64F1D6DF39BC15712D11" italics="true">B. hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0E3CB241A11E593F0111E1F5373FA2B3" rank="species">comb. nov.</taxonomicNameLabel>
and redescribed this species in the above section.
</paragraph>
<paragraph id="673527274283DC8CDCA332E8547A1F81">
In the original description (
<bibRefCitation id="30EF24A47E2109BB490767C925C1CFD2" author="Schaefer and Ashlock" firstAuthor="Schaefer" issue="3" journalOrPublisher="Pacific Insects" pagination="629-639" refId="B11" refString="Schaefer CW, Ashlock PD (1970) A new genus and new species of Saileriolinae (Hemiptera: Urostylidae). Pacific Insects 12 (3): 629639." title="A new genus and new species of Saileriolinae (Hemiptera: Urostylidae)." volume="12" year="1970">Schaefer and Ashlock 1970</bibRefCitation>
),
<taxonomicName id="6F0FE5C14E431F564BFEDECCCA7D8A01" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="4EB840A296583E398A3BA20AFA0DB33A" italics="true">B. hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35774E8BAF0DDA790ADC78794C69367F" rank="species">comb. nov.</taxonomicNameLabel>
was not compared with the only other congener,
<taxonomicName id="0E03AE9E89E7E80F7B4DA90A539131BC" authorityName="Hsiao" authorityYear="1964" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="arboreus">
<emphasis id="2DF4E98B2C00A24F6F70AF414F9E9261" italics="true">B. arboreus</emphasis>
</taxonomicName>
, making the identification of the two species difficult. However, based on the comparison among the
<specimenCount id="7F7DBC386856C8836349624078A0732F" count="9" type="generic">nine specimens</specimenCount>
of
<taxonomicName id="72DF811A1A6A2AADCE3C039EDC177EE9" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="0F180955053492961B743530DBB04CB3" italics="true">B. hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F17D6AF4A2E716F3B20096A5695CA274" rank="species">comb. nov.</taxonomicNameLabel>
and the illustrations (
<bibRefCitation id="47E2113DF820BA9968012A371473DF11" author="Hsiao" firstAuthor="Hsiao" issue="2" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="283-292" refId="B4" refString="Hsiao TY (1964) New species and new record of Hemiptera - Heteroptera from China. Acta Zootaxonomica Sinica 1 (2): 283292. [in Chinese with English summary]" title="New species and new record of Hemiptera - Heteroptera from China." volume="1" year="1964">Hsiao 1964</bibRefCitation>
;
<bibRefCitation id="B71755C37D01D938697B9F69C2D47DD6" author="Hsiao and Ching" firstAuthor="Hsiao" refId="B5" refString="Hsiao TY, Ching HL (1977) Urostylidae. In: Hsiao TY, Jen SC, Cheng LI, Liu SL, Ching HL (Eds) A Handbook for the Determination of the Chinese Hemiptera - Heteroptera. Vol. I. Science Press, Beijing, 181197 + 302304. [pls. 3032] [in Chinese with English summary]" year="1977">Hsiao and Ching 1977</bibRefCitation>
), photographs (
<bibRefCitation id="67D9BE99253DA8FB5EF251A4DE122BDB" author="Hsiao and Ching" firstAuthor="Hsiao" refId="B5" refString="Hsiao TY, Ching HL (1977) Urostylidae. In: Hsiao TY, Jen SC, Cheng LI, Liu SL, Ching HL (Eds) A Handbook for the Determination of the Chinese Hemiptera - Heteroptera. Vol. I. Science Press, Beijing, 181197 + 302304. [pls. 3032] [in Chinese with English summary]" year="1977">Hsiao and Ching 1977</bibRefCitation>
;
<bibRefCitation id="7241188F430E849ED5BF8ECB6341C74C" DOI="10.1201/9781315371221-2" author="Rider" editor="McPherson JE" etAl="et al." firstAuthor="Rider" journalOrPublisher="CRC Press, Boca Raton" pagination="25-201" refId="B10" refString="Rider DA, Schwertner CF, Vilímová J, Rédei D, Kment P, Thomas DB (2018) Higher systematics of the Pentatomoidea. In: McPherson JE (Ed.) Invasive Stink Bugs and Related Species (Pentatomoidea): Biology, Higher Systematics, Semiochemistry, and Management. CRC Press, Boca Raton, 25201. [pls 2.252.32] https://doi.org/10.1201/9781315371221-2" title="Higher systematics of the Pentatomoidea." volumeTitle="Invasive Stink Bugs and Related Species (Pentatomoidea): Biology, Higher Systematics, Semiochemistry, and Management." year="2018">Rider et al. 2018</bibRefCitation>
;
<ext-link id="25F61FFB6F4768666CE36A0B603E7B0A" ext-link-type="uri" xlink_href="https://spain.inaturalist.org/taxa/1360964-Bannacoris-arboreus" xlink_type="simple">https://spain.inaturalist.org/taxa/1360964-Bannacoris-arboreus</ext-link>
), and original description (
<bibRefCitation id="1AC7C8A1D0BA8A9DCB9796E0AE6B7DE4" author="Hsiao" firstAuthor="Hsiao" issue="2" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="283-292" refId="B4" refString="Hsiao TY (1964) New species and new record of Hemiptera - Heteroptera from China. Acta Zootaxonomica Sinica 1 (2): 283292. [in Chinese with English summary]" title="New species and new record of Hemiptera - Heteroptera from China." volume="1" year="1964">Hsiao 1964</bibRefCitation>
) of
<taxonomicName id="165110066329783A8AD77822F537540F" authorityName="Hsiao" authorityYear="1964" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="arboreus">
<emphasis id="207D7B41B465453D284FF293BD3B4163" italics="true">B. arboreus</emphasis>
</taxonomicName>
, the five characters described in the Diagnosis section above were recognized to easily differentiate
<taxonomicName id="8AEF4859C5ADB58AAC2F9F3A05978EC7" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="1FA20B7825B1EE14B22F1D019BFB7571" italics="true">B. hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889767A6C72DA873F983A4EC90BCDD11" rank="species">comb. nov.</taxonomicNameLabel>
from
<taxonomicName id="BE3E48BFA653C18856A1C57BA4B4027C" authorityName="Hsiao" authorityYear="1964" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="arboreus">
<emphasis id="8CB0356E95687A557458407FAFA6A048" italics="true">B. arboreus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ER2AE" type="distribution">
<paragraph id="F74B8DB5E70A73D1471FC4D7FCE7E9DB">
<heading id="658DBD3FE4892DE938A805598B09482F" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="A51687EB02EC91245A3BE2E9FCE3297B">
<collectingCountry id="4FCAD1A581B222107AD277C882730F39" name="Vietnam">Vietnam</collectingCountry>
(Th ừa Thiên Hu ế Province, Lâm Đ ồng Province) (Fig.
<figureCitation id="18B38A46A2628E7A13E5E57E46CC4C2B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Collection sites of Bannacoris hyalinus comb. nov.: red circle = new record; green circle = known record." figureDoi="10.3897/zookeys.1221.135026.figure5" httpUri="https://binary.pensoft.net/fig/1210694">5</figureCitation>
) (
<bibRefCitation id="73C044A281D44985D440AF08B9E8FDA5" author="Schaefer and Ashlock" firstAuthor="Schaefer" issue="3" journalOrPublisher="Pacific Insects" pagination="629-639" refId="B11" refString="Schaefer CW, Ashlock PD (1970) A new genus and new species of Saileriolinae (Hemiptera: Urostylidae). Pacific Insects 12 (3): 629639." title="A new genus and new species of Saileriolinae (Hemiptera: Urostylidae)." volume="12" year="1970">Schaefer and Ashlock 1970</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E62AE" type="host">
<paragraph id="711504B32D594A533BCB674CCC6D0BEB">
<heading id="4270990022DF75B78418483F6044F734" reason="title">Host plant.</heading>
</paragraph>
<paragraph id="8C3689C8084905FD38A7F2C3F32EC7DC">
Adults and nymphs of
<taxonomicName id="A21088BA0AF7A597B928CF1970D8B550" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="F2D23FFEED0BADCFBFB511BEDB0F04AF" italics="true">Bannacoris hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="670190CD75B55ED6F246963848B7CBE5" rank="species">comb. nov.</taxonomicNameLabel>
were observed to congregate in groups of several on the abaxial side of leaves of
<taxonomicName id="1A76E6A462A86AB0E876FEF82D0062B6" class="Magnoliopsida" family="Lauraceae" genus="Litsea" kingdom="Plantae" order="Laurales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="9FAB807DC593840E48F14B7578110D46" italics="true">Litsea</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="786B32FBE91155FEB8FD677A65E8A3CF" class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" order="Laurales" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
) (Fig.
<figureCitation id="230BFF465882FF15A58D716F20232EEB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Photographs regarding Bannacoris hyalinus comb. nov. from B ạch Mã National Park, Th ừa Thiên Hu ế Province, Vietnam A living adult B living nymph C host plant (Litsea sp.) D surrounding habitat." figureDoi="10.3897/zookeys.1221.135026.figure4" httpUri="https://binary.pensoft.net/fig/1210693">4 C</figureCitation>
) in B ạch Mã National Park, Th ừa Thiên Hu ế Province,
<collectingCountry id="B8085C3BA5E2FF8D9FC86144010BAB60" name="Vietnam">Vietnam</collectingCountry>
(Fig.
<figureCitation id="E367F70074007D23AC82FC89FE8A3EBB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Photographs regarding Bannacoris hyalinus comb. nov. from B ạch Mã National Park, Th ừa Thiên Hu ế Province, Vietnam A living adult B living nymph C host plant (Litsea sp.) D surrounding habitat." figureDoi="10.3897/zookeys.1221.135026.figure4" httpUri="https://binary.pensoft.net/fig/1210693">4 D</figureCitation>
), by the first author. In addition, nymphs were observed sucking sap from the abaxial side of the leaves of this lauraceous tree in captivity. Thus,
<taxonomicName id="4A629975FEAD26B878DB4E4A13FE4235" class="Magnoliopsida" family="Lauraceae" genus="Litsea" kingdom="Plantae" order="Laurales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="BDACBE1CFFA31EA34962E5A33441C3E8" italics="true">Litsea</emphasis>
sp.
</taxonomicName>
is considered the host plant of
<taxonomicName id="9E7F3E1C2E50C94570CB47B3B3C731B4" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="FECCDC48F8379F6EDE81560D2EA7E56B" italics="true">B. hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3A79A7899E49382283E13E5C9E721F82" rank="species">comb. nov.</taxonomicNameLabel>
, the biological information of which was unknown in the original description (
<bibRefCitation id="734BB1CFB9105700FF7D79C26EBF2F74" author="Schaefer and Ashlock" firstAuthor="Schaefer" issue="3" journalOrPublisher="Pacific Insects" pagination="629-639" refId="B11" refString="Schaefer CW, Ashlock PD (1970) A new genus and new species of Saileriolinae (Hemiptera: Urostylidae). Pacific Insects 12 (3): 629639." title="A new genus and new species of Saileriolinae (Hemiptera: Urostylidae)." volume="12" year="1970">Schaefer and Ashlock 1970</bibRefCitation>
).
</paragraph>
<caption id="BA81A35311AAFDB87BE2B6748E857FA8" ID-DOI="10.3897/zookeys.1221.135026.figure4" ID-arpha="7500D07D-00CA-57FE-A5D0-98A05C09817B" httpUri="https://binary.pensoft.net/fig/1210693" startId="F4">
<paragraph id="B253ADDE78A3BAF3CC3D84C152037C87">
<label id="B6C765D5D0CB72A409048FEBFC00A46C">Figure 4.</label>
</paragraph>
<paragraph id="7612BB85A5CDAECCB644472C4FDB68CD">
Photographs regarding
<taxonomicName id="C642097E9D0BC2D4424C29BF0C4C1EDC" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="8894D8C3086A835FDB508761FF7A4FAF" italics="true">Bannacoris hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C96CCA4DF2A768FD9E527C821AD35FCF" rank="species">comb. nov.</taxonomicNameLabel>
from B ạch Mã National Park, Th ừa Thiên Hu ế Province, Vietnam
<emphasis id="7241A685A60B1769998042606D91329E" bold="true">A</emphasis>
living adult
<emphasis id="98576A5CF2704EBB70032486E0DE3A3D" bold="true">B</emphasis>
living nymph
<emphasis id="15791E6DFF179A5B0B5AB2F7C38AFB8A" bold="true">C</emphasis>
host plant (
<taxonomicName id="0ADE7FE29F7F97126F1A8388C25F6B21" class="Magnoliopsida" family="Lauraceae" genus="Litsea" kingdom="Plantae" order="Laurales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="6091E9C9E33097B9A950CE4740AF23CC" italics="true">Litsea</emphasis>
sp.
</taxonomicName>
)
<emphasis id="ABF5BD6852086F474BFD6AE5A7208C94" bold="true">D</emphasis>
surrounding habitat.
</paragraph>
</caption>
<caption id="BF5D7AC6D645B2F00CB70A651154162D" ID-DOI="10.3897/zookeys.1221.135026.figure5" ID-arpha="C808DC1F-1A99-51F4-AB3F-C313E18375E1" httpUri="https://binary.pensoft.net/fig/1210694" startId="F5">
<paragraph id="1EB9BD94E5AD8069F1ACBE025084A894">
<label id="08DB9129E839A96033182022E70CB5FD">Figure 5.</label>
</paragraph>
<paragraph id="721A992D7783839C76F144FE02EB3730">
Collection sites of
<taxonomicName id="6FFA8F9D499E3DCAEF48C0BE855857CA" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="9926E2840B1B11E56F0D9F9078CC681D" italics="true">Bannacoris hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FD0D6A01B0E43DC9F27A880BA84F4E0B" rank="species">comb. nov.</taxonomicNameLabel>
: red circle = new record; green circle = known record.
</paragraph>
</caption>
<paragraph id="A86E6B846E9E5033A73FC8DD1E3E050E">
The adaxial side of the leaves apparently damaged by this saileriolid species was irregularly yellowed in the field and in captivity, suggesting the possibility that
<taxonomicName id="21045ACEBB654E54BBC4AD9487ED3CB8" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="6F741A7CDC59FB9897935108EF8B1B66" italics="true">B. hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72BDAA3E41A49052CC10D990FDFACED3" rank="species">comb. nov.</taxonomicNameLabel>
feeds on leaf chlorophyll.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHAAG" type="Bionomics">
<paragraph id="66F7161632E57D3B6ACED79B6866D1D6">
<heading id="11083AEBBAEB3C4C92C2DE41A04CB13E" reason="title">Bionomics.</heading>
</paragraph>
<paragraph id="3340CE810CD44928ED50DB968E7DC0A9">
<taxonomicName id="847B40DD0F4FD5A5B3C0A527310B089A" authorityName="Souma &amp; Le &amp; Pham" authorityYear="2024" baseAuthorityName="Schaefer &amp; Ashlock" baseAuthorityYear="1970" class="Insecta" family="Urostylididae" genus="Bannacoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="hyalinus" status="comb. nov.">
<emphasis id="68498E5B5337B8580D37310951C1BB24" italics="true">Bannacoris hyalinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFE2EC2AAFF1AA86FEBD6E125375C2F7" rank="species">comb. nov.</taxonomicNameLabel>
inhabits evergreen broad-leaved forests in the mountainous areas of
<collectingCountry id="ED8D675F88D4C3ECC261BEE7889D6EBE" name="Vietnam">Vietnam</collectingCountry>
with a subtropical climate.
</paragraph>
<paragraph id="33AEF697F05D80ACA237A0F93F7B73C2">
Adults were collected in
<date id="52E8A45A59F1D594C0A0D20D5DC60C6E" value="1960-05">May 1960</date>
and
<date id="68A610A281C27060B0AF7A5CB8309F54" value="2024-06">June 2024</date>
(
<bibRefCitation id="AA82D27C72AD48EE7D37EC03C1D8534F" author="Schaefer and Ashlock" firstAuthor="Schaefer" issue="3" journalOrPublisher="Pacific Insects" pagination="629-639" refId="B11" refString="Schaefer CW, Ashlock PD (1970) A new genus and new species of Saileriolinae (Hemiptera: Urostylidae). Pacific Insects 12 (3): 629639." title="A new genus and new species of Saileriolinae (Hemiptera: Urostylidae)." volume="12" year="1970">Schaefer and Ashlock 1970</bibRefCitation>
; present study), and nymphs were observed in
<date id="6C07381A7F548196A70EAFC586FA92BA" value="2024-06">June 2024</date>
(present study).
</paragraph>
</subSubSection>
</treatment>
</document>