422 lines
44 KiB
XML
422 lines
44 KiB
XML
<document ID-CLB-Dataset="6050" ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.60072" ID-GBIF-Dataset="a8375122-971c-4804-92a6-fe16e10e3224" ID-PMC="PMC8222295" ID-Pensoft-Pub="1313-2970-1044-427" ID-Pensoft-UUID="8320077CEFA5571D971EFEA66B0C6268" ID-PubMed="34183882" ID-ZooBank="9AA72654B6CC4081B557CD0BAE3939A3" ModsDocID="1313-2970-1044-427" checkinTime="1623875398087" checkinUser="pensoft" docAuthor="Allegro, Gianni & Giachino, Pier Mauro" docDate="2021" docId="DC82C0062D235FA08F17B1D83A9DD9EB" docLanguage="en" docName="ZooKeys 1044: 427-448" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.60072" docTitle="Diplocheila (Diplocheila) erwini Allegro & Giachino 2021, sp. nov." docType="treatment" docUuid="108A8C27-E70A-4322-AF94-73E2CE64DEAB" docUuidSource="ZooBank" docVersion="6" id="8320077CEFA5571D971EFEA66B0C6268" lastPageNumber="427" masterDocId="8320077CEFA5571D971EFEA66B0C6268" masterDocTitle="The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini)" masterLastPageNumber="448" masterPageNumber="427" pageNumber="427" updateTime="1732756944497" updateUser="ExternalLinkService">
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<mods:title id="3C1CC819EE235CB966AE1EC9A0F3C76B">The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini)</mods:title>
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<mods:name id="6B51176F87919E474B85863CB17C5890" type="personal">
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<mods:roleTerm id="06E640F37E51BCCD47AA8FAF1981FB42">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="17D7DCE0E8702FAF3ED7B335BAC0D38D">Allegro, Gianni</mods:namePart>
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<mods:nameIdentifier id="847652DE4F1191C88E183FF982D84400" type="ORCID">https://orcid.org/0000-0002-5079-6671</mods:nameIdentifier>
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<mods:affiliation id="07596B4A591D9B8C335469A8843B29D3">Strada Patro 11, I- 14036 Moncalvo (AT), Italy</mods:affiliation>
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<mods:nameIdentifier id="50C297768CC38B15F3ACCDA55EAA9A44" type="email">gianni.allegro54@gmail.com</mods:nameIdentifier>
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<mods:name id="4FA4B436647B3DEF1DC3B691785AD238" type="personal">
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<mods:roleTerm id="1E4CA08D06964DEFC18C2710A0CA188E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5CACE3A36A07E1440654AFAF48F4A84E">Giachino, Pier Mauro</mods:namePart>
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<mods:nameIdentifier id="E984D4F9E03450DA0C05F9E7BB1FB1DD" type="ORCID">https://orcid.org/0000-0002-1167-5447</mods:nameIdentifier>
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<mods:affiliation id="0CF86795D121FD72D568F573BE44BE11">Via della Trinita 13, I- 10010 San Martino Canavese (TO), Italy</mods:affiliation>
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<mods:typeOfResource id="65CB2F54ADD4C6C2CF4C655827A332E8">text</mods:typeOfResource>
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<mods:title id="054E4E56A753AC14A39E24731445C884">ZooKeys</mods:title>
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<mods:part id="D4215A967E1243BD4E6859A47D0ECCC3">
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<mods:date id="9D36AC48D3D00EB55CEF9208293D36CE">2021</mods:date>
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<mods:number id="266B543F28F6A3FE096458489AFFABEC">2021-06-16</mods:number>
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<mods:detail id="CAAAD519ADAE6ECDD67C32DD5487BB80" type="volume">
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<mods:number id="80DC85BB11F96551915EFB88BEB71ED8">1044</mods:number>
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<mods:start id="94CE3DFC399E7C3AEE65C55085D6AAF9">427</mods:start>
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<mods:url id="B7783038EDBAB98F7929FC1293BB8229">http://dx.doi.org/10.3897/zookeys.1044.60072</mods:url>
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</mods:location>
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<mods:classification id="9649E610FF08B0EF8ACA26CA87A4BB95">journal article</mods:classification>
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<mods:identifier id="B93728BB9E179F0B7FD4D6F32E52955D" type="DOI">http://dx.doi.org/10.3897/zookeys.1044.60072</mods:identifier>
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<mods:identifier id="925FA70C8AA643000DAA23E4AE453899" type="Pensoft-Pub">1313-2970-1044-427</mods:identifier>
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<mods:identifier id="CD25A8CC9E0CD93C617FF4F833368F1E" type="ZooBank">9AA72654B6CC4081B557CD0BAE3939A3</mods:identifier>
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<mods:identifier id="5EEEEBDF645ED276D14D148108194D9D" type="Pensoft-UUID">8320077CEFA5571D971EFEA66B0C6268</mods:identifier>
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<treatment id="DC82C0062D235FA08F17B1D83A9DD9EB" ID-GBIF-Taxon="182754008" LSID="urn:lsid:zoobank.org:act:108A8C27-E70A-4322-AF94-73E2CE64DEAB" httpUri="http://treatment.plazi.org/id/DC82C0062D235FA08F17B1D83A9DD9EB" lastPageNumber="427" pageId="0" pageNumber="427" scope_family="Carabidae" scope_order="Coleoptera" scope_tribe="Licinini">
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<subSubSection id="B516455F95D4AE6A46E98AD0C2D4F02E" pageId="0" pageNumber="427" type="nomenclature">
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||
<paragraph id="2ED978399B452D9638673C9CC0EB9777" pageId="0" pageNumber="427">
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||
<taxonomicName id="C4032CFD98699A3C453E0512B580EBFC" ID-CoL="8M57S" LSID="http://zoobank.org/108A8C27-E70A-4322-AF94-73E2CE64DEAB" authority="Allegro & Giachino, 2021" authorityName="Allegro & Giachino" authorityYear="2021" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila (Diplocheila) erwini" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="erwini" status="sp. nov." subGenus="Diplocheila">Diplocheila (Diplocheila) erwini</taxonomicName>
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<taxonomicNameLabel id="CEAA4826379F3E31847B666A602B1E51" pageId="0" pageNumber="427">sp. nov.</taxonomicNameLabel>
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<figureCitation id="DC419303D15F6E14298BEA3A02CE7D06" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Diplocheila erwini sp. nov.: habitus of holotype." figureDoi="10.3897/zookeys.1044.60072.figure5" httpUri="https://binary.pensoft.net/fig/554096" pageId="0" pageNumber="427">Figures 5</figureCitation>
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<figureCitation id="F3843CAAA40175948FEF7976E03C445B" captionStart="Figures 6–11" captionStartId="F6" captionText="Figures 6 - 11. Pronotum of Diplocheila species 6 D. laevigata HT 7 D. laevigotoides HT 8 D. laevigata from Cambodia 9 Diplocheila sp. from Thailand 10 D. walterrossii sp. nov. HT 11 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427">, 11</figureCitation>
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<figureCitation id="87E60A606A1A1F1C4C65A6AAEADF9D85" captionStart="Figures 12–17" captionStartId="F7" captionText="Figures 12 - 17. Labrum of Diplocheila species 12 D. laevigata HT 13 D. laevigotoides HT 14 D. laevigata from Cambodia 15 Diplocheila sp. from Thailand 16 D. walterrossii sp. nov. HT 17 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures12-17" httpUri="https://binary.pensoft.net/fig/554098" pageId="0" pageNumber="427">, 17</figureCitation>
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<figureCitation id="AAB3CAA5530B60A23E05B90CFD1A7911" captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Aedeagus in lateral view of Diplocheila species 20 D. laevigata from Cambodia 21 Diplocheila sp. from Thailand 22 D. walterrossii sp. nov. HT 23 D. erwini sp. nov. HT 24 D. laevis from Cambodia 25 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures20-25" httpUri="https://binary.pensoft.net/fig/554100" pageId="0" pageNumber="427">, 23</figureCitation>
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<figureCitation id="5A43C23F70C2AADDCE356926F8A8DFCF" captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">, 30</figureCitation>
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<figureCitation id="775FD73AC6C84902323C366478B20E1C" captionStart="Figure 33" captionStartId="F11" captionText="Figure 33. Distribution map of Diplocheila walterrossii sp. nov. and D. erwini sp. nov. in Cambodia." figureDoi="10.3897/zookeys.1044.60072.figure33" httpUri="https://binary.pensoft.net/fig/554102" pageId="0" pageNumber="427">, 33</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="D431D910D4A3D699D3CB4C453FF6CFED" pageId="0" pageNumber="427" type="type locality">
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||
<paragraph id="36354E47E5E61C71901D4B26C04E5307" pageId="0" pageNumber="427">Type locality.</paragraph>
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<paragraph id="12E82119AA8818BE0FA30DEA955D0C4E" pageId="0" pageNumber="427">Cambodia, Kampong Chhnang, Khom Domnatpopol, Tonle Sap Lake.</paragraph>
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</subSubSection>
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||
<subSubSection id="45445E139A4491EC123DB2ED904D5C15" pageId="0" pageNumber="427" type="materials_examined">
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||
<paragraph id="F843F8AE6E1E56B02E82C400FED154D6" pageId="0" pageNumber="427">Material examined.</paragraph>
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<paragraph id="88EB334E8B5D395A2CA6566B2BDA66C7" pageId="0" pageNumber="427">
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||
<materialsCitation id="CC03BFD2DFF55AC5770E636A9042ADA4" collectingDate="2018-05-21" collectingMethod="light trap" collectorName="Rossi, Bernardi, Kong" country="Cambodia" latitude="12.237223" location="Tonle Sap Lake" longLatPrecision="21" longitude="104.6875" municipality="Khom Domnatpopol" specimenCount="♂" stateProvince="Kampong Chhnang Province" typeStatus="Holotype">
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||
<emphasis id="30AAD93A938FE787D7E96133B4BBBA20" bold="true" italics="true" pageId="0" pageNumber="427">
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||
<typeStatus id="9E4300A39F327361D3B54284835D4AC5">Holotype</typeStatus>
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||
</emphasis>
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||
:
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||
<specimenCount id="CC8A30FB6372A42F08F755FD49D79BED">♂</specimenCount>
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,
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<collectingCountry id="6A48858B9862BE39A345DBA2B76D4EDA" name="Cambodia">Cambodia</collectingCountry>
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||
,
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<collectingRegion id="BD1C95698F2A30506168D473DDD69FDA" country="Cambodia" name="Kampong Chhnang">Kampong Chhnang Province</collectingRegion>
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||
,
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||
<collectingMunicipality id="26937CCEF5D81BBA34E7D093F178ECB2">Khom Domnatpopol</collectingMunicipality>
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||
,
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||
<location id="4A7326C6677D03AB590E5636C48D3351" LSID="urn:lsid:plazi:treatment:DC82C0062D235FA08F17B1D83A9DD9EB:4A7326C6677D03AB590E5636C48D3351" country="Cambodia" latitude="12.237223" longLatPrecision="21" longitude="104.6875" municipality="Khom Domnatpopol" name="Tonle Sap Lake" stateProvince="Kampong Chhnang Province">Tonle Sap Lake</location>
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||
,
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||
<geoCoordinate id="191EFFD698E373EC3956B62A5CDBA310" degrees="12" direction="north" minutes="14" orientation="latitude" precision="15" seconds="14" value="12.237223">12°14'14"N</geoCoordinate>
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||
,
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||
<geoCoordinate id="C1C0ACE38C6875DF67D48E0B02AEE883" degrees="104" direction="east" minutes="41" orientation="longitude" precision="15" seconds="15" value="104.6875">104°41'15"E</geoCoordinate>
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||
, (
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<collectingMethod id="DF2A24A2585C35C96C9151C2C54CF4AC">light trap</collectingMethod>
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||
),
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<collectingDate id="6F8CF247219A92D1D2014F42A36EC438" value="2018-05-21">21.V.2018</collectingDate>
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,
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<collectorName id="36F72D68B7F52DD4698EE1A152AC0F8C">Rossi</collectorName>
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,
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<collectorName id="7CC15CAF39EA57C84D56BE1DDEB89E98">Bernardi</collectorName>
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and
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<collectorName id="B3365F353F90DD73AD99836B277C2B55">Kong</collectorName>
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leg. (CAl).
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="D0F6EB169E22C91F00011A51569DD877" pageId="0" pageNumber="427">
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||
<emphasis id="3C00A44E52C6F1DEA43AE0C4662D95ED" bold="true" italics="true" pageId="0" pageNumber="427">
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||
<typeStatus id="B66E7F78468DDAB1FFF711DDB0BFEA8B">Paratypes</typeStatus>
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||
</emphasis>
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||
: (
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<specimenCount id="7B151D87AE4A749BF175538B6F2FB4DD" type="male">1 ♂</specimenCount>
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and
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<specimenCount id="46E0D5CC7FB68BCD0C4A48AE23508652" type="female">3 ♀♀</specimenCount>
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);
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<materialsCitation id="BF1CFD5EF4EAEF119690DD9C0DD80211" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Rossi & Bernardi & Kong" country="Cambodia" latitude="12.237223" location="Tonle Sap Lake" longLatPrecision="21" longitude="104.6875" municipality="Khom Domnatpopol" specimenCount="1" specimenCount-female="1" stateProvince="Kampong Chhnang" typeStatus="paratype">
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<specimenCount id="9021AF40F52004375D41CB74BF10D19C" type="female">1 ♀</specimenCount>
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same data as holotype
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</materialsCitation>
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;
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<materialsCitation id="A33408E2BBB72617F98B411CA4ECF3D8" collectingDate="2018-11-07" collectingMethod="light trap" collectorName="W. Rossi, V. Kong" country="Cambodia" county="Rolea" latitude="12.225277" location="Toulkrolanh Village" longLatPrecision="21" longitude="104.66389" municipality="District" specimenCount="1" specimenCount-male="1" stateProvince="Kampong Chhnang" typeStatus="paratype">
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<specimenCount id="2ED15766BBEFB2CC2372D32DEF88AB17" type="male">1 ♂</specimenCount>
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<collectingCountry id="8EDDEDAA4DEA6BC1DC755A1F85A6C245" name="Cambodia">Cambodia</collectingCountry>
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,
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<collectingRegion id="BFF5B9E6BA4C21453719B376DC163499" country="Cambodia" name="Kampong Chhnang">Kampong Chhnang</collectingRegion>
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||
,
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||
<collectingCounty id="F2AB86FB6DD63AEBC6BD2465CF956A21">Rolea</collectingCounty>
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||
<normalizedToken id="0638120FEBCE29A4DB3EA3F2ADB4C65E" originalValue="B’ier">B'ier</normalizedToken>
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||
<collectingMunicipality id="8A0259921F6EE9E2002E44F7F6CD2DA5">District</collectingMunicipality>
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||
,
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||
<location id="B511AA93BB25AF21D0B8B2631DEF3FAE" LSID="urn:lsid:plazi:treatment:DC82C0062D235FA08F17B1D83A9DD9EB:B511AA93BB25AF21D0B8B2631DEF3FAE" country="Cambodia" county="Rolea" latitude="12.225277" longLatPrecision="21" longitude="104.66389" municipality="District" name="Toulkrolanh Village" stateProvince="Kampong Chhnang">Toulkrolanh Village</location>
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||
,
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||
<geoCoordinate id="CBE9BF92D4DA510D7AD243F3D3AACA99" degrees="12" direction="north" minutes="13" orientation="latitude" precision="15" seconds="31" value="12.225277">12°13'31"N</geoCoordinate>
|
||
,
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||
<geoCoordinate id="179AA4C5AA1CE9A983B1E55A54AAAEBB" degrees="104" direction="east" minutes="39" orientation="longitude" precision="15" seconds="50" value="104.66389">104°39'50"E</geoCoordinate>
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||
,
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||
<collectingMethod id="72769702F1BDC7D2680C16AE0515F700">light trap</collectingMethod>
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||
,
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||
<collectingDate id="05EA2A6C9F58F5F60EAEFC8F1BB33D71" value="2018-11-07">7.XI.2018</collectingDate>
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||
,
|
||
<collectorName id="028D853A76B614F6EC4A7AAD814D834E">W. Rossi</collectorName>
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||
and
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||
<collectorName id="F11FBE61B1FA5272F4C835DEA3CEA4AD">V. Kong</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="E36C8AC5899259115BB45E2BA2C53F4E" collectingDate="2019-05-17" collectingMethod="light trap" collectorName="W. Rossi, V. Kong" country="Cambodia" location="Tonle Sap Lake" specimenCount="2" specimenCount-female="2" stateProvince="Kampong Chhnang" typeStatus="paratype">
|
||
<specimenCount id="342AC196F76BF10A2717EF9A3B22BD1A" type="female">2 ♀♀</specimenCount>
|
||
,
|
||
<collectingCountry id="653BC833DACD74B36CA548702893A228" name="Cambodia">Cambodia</collectingCountry>
|
||
,
|
||
<collectingRegion id="2097A424444B5B9CBB23BC4C91E8A4CB" country="Cambodia" name="Kampong Chhnang">Kampong Chhnang</collectingRegion>
|
||
, banks of
|
||
<location id="9712BFCB0D04CF69E72E5185E37AA987" LSID="urn:lsid:plazi:treatment:DC82C0062D235FA08F17B1D83A9DD9EB:9712BFCB0D04CF69E72E5185E37AA987" country="Cambodia" name="Tonle Sap Lake" stateProvince="Kampong Chhnang">Tonle Sap Lake</location>
|
||
,
|
||
<collectingMethod id="7F96D26BED46B4C933E99FDD70E9623F">light trap</collectingMethod>
|
||
,
|
||
<collectingDate id="633FF45AD2B8D2C3066B782E78472D88" value="2019-05-17">17.V.2019</collectingDate>
|
||
,
|
||
<collectorName id="1E1E034729791DDA5418FF6CCA6E4B18">W. Rossi</collectorName>
|
||
and
|
||
<collectorName id="74CAD78388215D370FF9D8349C444A06">V. Kong</collectorName>
|
||
leg. (CAl, CGi)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8A497A688FEA1AFF0F79B4873EFE89C" pageId="0" pageNumber="427" type="diagnosis">
|
||
<paragraph id="B76704DF320192AA9F10ABF6F8AAC378" pageId="0" pageNumber="427">Diagnosis.</paragraph>
|
||
<paragraph id="E806496E47AA1E7E428ECF82DE830F9C" pageId="0" pageNumber="427">
|
||
A medium-sized to small
|
||
<taxonomicName id="D52D804CC85F3451399486F71C57D3E3" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4158463FDDF471BE53EED636717F7A51" italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
|
||
</taxonomicName>
|
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(ABL: 12-14 mm) of the
|
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<taxonomicName id="682E32301CF75DAE6634BB252107D65D" baseAuthorityName="Fabricius" baseAuthorityYear="1792" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila polita" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="polita">
|
||
<emphasis id="1DBC1331A25A1F47EE9E0B6F2B850BA4" italics="true" pageId="0" pageNumber="427">Diplocheila polita</emphasis>
|
||
</taxonomicName>
|
||
group in the subgenus
|
||
<taxonomicName id="BB8A2BA828DCD35DA502E69EB894AD61" authority="Diplocheila" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila (Diplocheila)" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="subGenus" subGenus="Diplocheila">Diplocheila Diplocheila</taxonomicName>
|
||
(sensu
|
||
<bibRefCitation id="D11FFBA4BA803A54D74436375C69AD52" author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
|
||
). It is easily distinguished from the other species of the group with a sexsetose labrum (
|
||
<taxonomicName id="33D6E947A7C0CE01C3A9A5C85A303755" lsidName="D. indus" pageId="0" pageNumber="427" rank="species" species="indus">
|
||
<emphasis id="349912496F32EF09D3E3760756C19145" italics="true" pageId="0" pageNumber="427">D. indus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7E1A9FE42E1E661B6C627F9FDF63640E" lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis id="6980494CEA69676500A8D2044297F98D" italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2F808AD0DC1DA0B5077D03DA4FA27995" lsidName="D. laevigotoides" pageId="0" pageNumber="427" rank="species" species="laevigotoides">
|
||
<emphasis id="6F9078C73EA7DBFA81561109014FE88B" italics="true" pageId="0" pageNumber="427">D. laevigotoides</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="18D2DF42FD62942376402C72A046584C" lsidName="D. walterrossii" pageId="0" pageNumber="427" rank="species" species="walterrossii">
|
||
<emphasis id="210E26C6FE3C1D748EF5275EBF0427ED" italics="true" pageId="0" pageNumber="427">D. walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.) by the smaller body size (≤14 mm), the narrower and almost quadrangular pronotum (transverse in the four other species), with hind angles not protruding (protruding in
|
||
<taxonomicName id="B9CF5F0D8C7ABFF3FC12F1E3EBABF556" lsidName="D. indus" pageId="0" pageNumber="427" rank="species" species="indus">
|
||
<emphasis id="F679846759B8514830EE2BDAADE1BD0D" italics="true" pageId="0" pageNumber="427">D. indus</emphasis>
|
||
</taxonomicName>
|
||
), and by the morphology of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A8C9CFF240F8325D8171819BE6554A34" pageId="0" pageNumber="427" type="description">
|
||
<paragraph id="29A623FFA431F8DCED21322B65A8DB23" pageId="0" pageNumber="427">Description.</paragraph>
|
||
<paragraph id="EB0D35520DF85AEBEFEE824973977724" pageId="0" pageNumber="427">
|
||
<emphasis id="86257C95EFE1F77B8A19B7E7E3938DEB" bold="true" italics="true" pageId="0" pageNumber="427">Habitus</emphasis>
|
||
: ABL: 12-14 mm (HT ♂ 13.5 mm). Body parallel-sided, moderately shiny, black with antennae and palpi piceous-brown (Fig.
|
||
<figureCitation id="687E203801DF91A156029CBE70FDB6FD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Diplocheila erwini sp. nov.: habitus of holotype." figureDoi="10.3897/zookeys.1044.60072.figure5" httpUri="https://binary.pensoft.net/fig/554096" pageId="0" pageNumber="427">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="20589FDCD30A51309C3ADF024620CFEF" doi="10.3897/zookeys.1044.60072.figure5" httpUri="https://binary.pensoft.net/fig/554096" pageId="0" pageNumber="427" start="Figure 5" startId="F5">
|
||
<paragraph id="706393F9E7C496C3B9FD93225C41B8D2" pageId="0" pageNumber="427">
|
||
<emphasis id="16F5CB600C372224FDF6682EDD3CA93D" bold="true" pageId="0" pageNumber="427">Figure 5.</emphasis>
|
||
<taxonomicName id="C92F15A772280080B984AA7D05CF616C" authorityName="Allegro & Giachino" authorityYear="2021" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila erwini" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="erwini">
|
||
<emphasis id="99F4D38818BF41001C55238355C5738A" italics="true" pageId="0" pageNumber="427">Diplocheila erwini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.: habitus of holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A89FB5C63639449C186570CE18EB6060" pageId="0" pageNumber="427">
|
||
<emphasis id="BED121769176950814D7683E3D96FB61" bold="true" italics="true" pageId="0" pageNumber="427">Head</emphasis>
|
||
: almost quadrangular, glabrous except for the supraorbital setae, narrow in comparison with pronotum. Eyes markedly convex; a single supraorbital seta on each side. Dorsum with microsculpture not evident, only with scattered punctures visible at>100
|
||
<normalizedToken id="765555722BA9D2644475EC30F90CBB73" originalValue="×">x</normalizedToken>
|
||
magnification; frontal impressions short and superficial. Labrum symmetrically and deeply (LR = 0.78) emarginate, with six setigerous punctures on anterior margin (4 medial equidistant + 2 lateral on lobes). Clypeus trapezoid, distinctly concave anteriorly, with 1 seta on each side at anterolateral corner. Antennae moderately long, densely pubescent from segment 4, with terminal 2 articles surpassing base of pronotum; segments elongate, the second one short, as long as a half of first. Mandibles elongate, broad, approximately similar each another (the left with apical cutting edge more concave), with scrobe well-defined and glabrous and apex blunt; terebral tooth triangular and prominent. Labial and maxillary palps fusiform, with apices narrowly truncate.
|
||
</paragraph>
|
||
<paragraph id="C9C24D9088CD2D425AF0764F6AEA5E49" pageId="0" pageNumber="427">
|
||
<emphasis id="D290D628373E40E4A7C9E25BF49BE686" bold="true" italics="true" pageId="0" pageNumber="427">Thorax</emphasis>
|
||
: pronotum smooth, with very faint microsculpture evident at>200
|
||
<normalizedToken id="0192AA4E576D2F5EEBDC06069A9410F2" originalValue="×">x</normalizedToken>
|
||
magnification and with scattered punctures, subquadrate (PW/PL = 1.18), widest at middle (Fig.
|
||
<figureCitation id="48309A897C01A7D5495FE3E865AA5AAD" captionStart="Figures 6–11" captionStartId="F6" captionText="Figures 6 - 11. Pronotum of Diplocheila species 6 D. laevigata HT 7 D. laevigotoides HT 8 D. laevigata from Cambodia 9 Diplocheila sp. from Thailand 10 D. walterrossii sp. nov. HT 11 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427">11</figureCitation>
|
||
). Disk moderately convex. Sides from rectilinear to hardly rounded in anterior half; rectilinear or very slightly sinuate backwards. Hind angles rounded obtuse, provided with a postero-lateral seta. Posterior margin rectilinear between basal impressions, which are linear and markedly impressed; anterior margin with front angles nearly obsolete. A single lateral seta on each side just above middle. Lateral bead continuous, separated from the discal area by a narrow groove, only scarcely dilated before hind angles. Medial longitudinal impression fine, nearly reaching anterior and posterior margins; anterior transversal impression absent.
|
||
</paragraph>
|
||
<paragraph id="C008DAF609B2C769AEA7BDF8C8C1FB59" pageId="0" pageNumber="427">
|
||
<emphasis id="9ED72FA24E915E00390A8947F1B24A04" bold="true" italics="true" pageId="0" pageNumber="427">Elytra</emphasis>
|
||
: moderately long (EL/EW = 1.59), parallel-sided, slightly convex and flattened on disk, widest at middle, with rounded shoulders and sides delicately sinuate before apex. Surface moderately shiny; microsculpture evident only at high magnification (>100
|
||
<normalizedToken id="6895171EDE5DBE4E2420A319D82E3075" originalValue="×">x</normalizedToken>
|
||
), consisting of fine, slightly transverse meshes. Epipleura without any distinct external plicae ("uncrossed epipleura"). Intervals moderately convex, smooth; striae deeply impressed on the whole length, distinctly punctate. Parascutellar stria present; scutellar setigerous pore present at base of stria 1, just before conjunction with stria 2. Basal margin complete. Discal setigerous punctures absent; umbilicate series of setigerous punctures continuous, not interrupted at middle. Hind wings fully developed.
|
||
</paragraph>
|
||
<paragraph id="E8EF77510BA306DEEDA2F1943A523C48" pageId="0" pageNumber="427">
|
||
<emphasis id="9F91AC3441051DE29B9FC23D6F492C43" bold="true" pageId="0" pageNumber="427">
|
||
<emphasis id="BC02323E5C2121B95041AF072230F86B" italics="true" pageId="0" pageNumber="427">Ventral surface</emphasis>
|
||
(thorax and abdomen)
|
||
</emphasis>
|
||
: prosternum and proepisterna glabrous and impunctate (only with very fine punctures). Metepisterna as long as twice the width of anterior side; metepimera large, broadly rounded. Prosternal intercoxal process widely rounded and bordered at apex. Abdominal ventrites IV-VI shiny but shagreened at sides, glabrous except one pair subapical central setae; males with 2, females with 4 marginal setae at apex of ventrite VII.
|
||
</paragraph>
|
||
<paragraph id="5038A4EB16046009BB3A08169F90C5F6" pageId="0" pageNumber="427">
|
||
<emphasis id="49693E124AD573CC3407C55F8A9A29C1" bold="true" italics="true" pageId="0" pageNumber="427">Legs</emphasis>
|
||
: moderately slender. Posterior face of femora with 1 seta in profemora, 2 in mesofemora and metafemora. Metatrochanters glabrous and slightly shorter than half length of metafemora. Protibial antennal cleaning organ well developed, with 2 clip setae. Protibiae robust, with 4 or 5 outer apical spines; mesotibiae with a group of setae at middle of inner face; metatibiae longitudinally furrowed at inner face. Dorsal face of tarsomeres smooth. Male protarsomeres 1-3 distinctly dilated, slightly asymmetrical; meso- and metatarsomeres not dilated in both sexes; tarsomere 5 ventrally glabrous, dorsally with 2 apical setae; claws smooth.
|
||
</paragraph>
|
||
<paragraph id="47A301755553866384BD5F67FCAEE45E" pageId="0" pageNumber="427">
|
||
<emphasis id="2D962467F6C7B36E88A7B14E0764B81D" bold="true" italics="true" pageId="0" pageNumber="427">Male genitalia</emphasis>
|
||
: median lobe of aedeagus short and markedly swollen before apex in lateral view (Fig.
|
||
<figureCitation id="6F2F116CF2E34D222C53847A994498FB" captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Aedeagus in lateral view of Diplocheila species 20 D. laevigata from Cambodia 21 Diplocheila sp. from Thailand 22 D. walterrossii sp. nov. HT 23 D. erwini sp. nov. HT 24 D. laevis from Cambodia 25 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures20-25" httpUri="https://binary.pensoft.net/fig/554100" pageId="0" pageNumber="427">23</figureCitation>
|
||
); the apical lamella shortly triangular in dorsal view, with blunt tip (Fig.
|
||
<figureCitation id="FECDE7883E5BCEC9F79E778358CC6E1E" captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">30</figureCitation>
|
||
), apex very slightly bent downwards in lateral view. Ostium long, in dorsal position. Right paramere oval; left paramere conchoid.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AF08C21B39FE4615603630D31DCFECFA" pageId="0" pageNumber="427" type="etymology">
|
||
<paragraph id="EF79D65C600C15ACE50FB70B63A23150" pageId="0" pageNumber="427">Etymology.</paragraph>
|
||
<paragraph id="B5426D0FC524C7297B88BE57A4E6A6E1" pageId="0" pageNumber="427">
|
||
This species is named, as a token of our esteem, after our late colleague Terry Erwin, a world-renowned specialist in world and tropical
|
||
<taxonomicName id="D7987DDD348A9716C381B9BD8CA35911" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1FE0780725886C6CFD0FE5F17633C8D7" pageId="0" pageNumber="427" type="distribution">
|
||
<paragraph id="C0196F0EA263E854DEB0FF105CFE4668" pageId="0" pageNumber="427">Distribution and ecology.</paragraph>
|
||
<paragraph id="83A50A5909A1A2BCE51A66BB2B3A4E51" pageId="0" pageNumber="427">
|
||
<taxonomicName id="0AACFEA4D02550830331CEF5B4E90C32" genus="Geographical" lsidName="Geographical distribution" pageId="0" pageNumber="427" rank="species" species="distribution">
|
||
<emphasis id="1D0EAFD698448165FB019678F29C2645" italics="true" pageId="0" pageNumber="427">Geographical distribution</emphasis>
|
||
</taxonomicName>
|
||
: this species is recorded only from the extreme south banks of the Tonle Sap Lake, Kampong Chhnang Province, Cambodia. It seems to have a more restricted
|
||
<taxonomicName id="2DBD7D1136703896C82D20642A83D799" genus="Geographical" lsidName="Geographical distribution" pageId="0" pageNumber="427" rank="species" species="distribution">Geographical distribution</taxonomicName>
|
||
than
|
||
<taxonomicName id="673ABF203771256DB41D33A7FC70C002" lsidName="D. walterrossii" pageId="0" pageNumber="427" rank="species" species="walterrossii">
|
||
<emphasis id="33B91EDEB8A554D44987B3C9030D62C8" italics="true" pageId="0" pageNumber="427">D. walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., which has been recorded from the same site as well as from other two localities in north-western Cambodia (Fig.
|
||
<figureCitation id="9937BF4D2027D9D8BBF8F33D12F59D7E" captionStart="Figure 33" captionStartId="F11" captionText="Figure 33. Distribution map of Diplocheila walterrossii sp. nov. and D. erwini sp. nov. in Cambodia." figureDoi="10.3897/zookeys.1044.60072.figure33" httpUri="https://binary.pensoft.net/fig/554102" pageId="0" pageNumber="427">33</figureCitation>
|
||
).
|
||
<emphasis id="D8268A337047FD537490469E113209B0" italics="true" pageId="0" pageNumber="427">Life habits</emphasis>
|
||
: the specimens of the type series were collected on lake banks by light trapping. No other data are available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="385C7DA50366D2163D14CA2185BF4024" pageId="0" pageNumber="427" type="remarks">
|
||
<paragraph id="88A834D395919C59930B807FE1EE4F86" pageId="0" pageNumber="427">Remarks.</paragraph>
|
||
<paragraph id="F8EAB45FAF5C262F0396B1321ED9F993" pageId="0" pageNumber="427">
|
||
It is difficult to assess the closest relatives of
|
||
<taxonomicName id="A695D2AE8B023C320D4262D830B57371" lsidName="D. erwini" pageId="0" pageNumber="427" rank="species" species="erwini">
|
||
<emphasis id="E3B99DFED72977A6EAF2A6D573118F54" italics="true" pageId="0" pageNumber="427">D. erwini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. It belongs to the
|
||
<taxonomicName id="ED8DA897C333E0D21B1C8B43BFCA4E40" lsidName="D. polita" pageId="0" pageNumber="427" rank="species" species="polita">
|
||
<emphasis id="7F85FD506C137FE13C46B0462D81C6ED" italics="true" pageId="0" pageNumber="427">D. polita</emphasis>
|
||
</taxonomicName>
|
||
group (sensu
|
||
<bibRefCitation id="7A385AB4D25AC4C88207298FADD2D992" author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
|
||
), as it shares the characters distinguishing this group and, moreover, its aedeagus is very similar to that of
|
||
<taxonomicName id="F8DD851AA0FE664652F898FA697FB3AA" lsidName="D. polita" pageId="0" pageNumber="427" rank="species" species="polita">
|
||
<emphasis id="7D861D37B59249C16EF901D6AE30F55C" italics="true" pageId="0" pageNumber="427">D. polita</emphasis>
|
||
</taxonomicName>
|
||
(see pl. V, fig. 72a in
|
||
<bibRefCitation id="4FF7C86452ED90CE8CEB4600EC3ACCAA" author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
|
||
).
|
||
<taxonomicName id="2F423A6F6F72ABE8B215ABD2FB47C37A" authorityName="Allegro & Giachino" authorityYear="2021" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila erwini" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="erwini">
|
||
<emphasis id="1C3D01792754400061B828B7557589EC" italics="true" pageId="0" pageNumber="427">Diplocheila erwini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., on the other hand, is easily distinguished from
|
||
<taxonomicName id="9875582E105A96119D80C6699C799D93" lsidName="D. polita" pageId="0" pageNumber="427" rank="species" species="polita">
|
||
<emphasis id="981F8B363671ABDFDEF6CA04984FE926" italics="true" pageId="0" pageNumber="427">D. polita</emphasis>
|
||
</taxonomicName>
|
||
which has a quadrisetose labrum and pronotum more transverse (PW/PL = 1.31 according to Fig.
|
||
<figureCitation id="ED3BB54D1530113E58AE243BB4CCB947" captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">26</figureCitation>
|
||
in Plate II in
|
||
<bibRefCitation id="F9FD7649FB3BAB0A2F65E077D2687387" author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
|
||
), as well as a larger body size (ABL = 13.4-18.4) (
|
||
<bibRefCitation id="5CE05B895CC36686743FD6BDD03A5CD0" author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="0B2D1F05F73165DEDAA8BA405F09D544" doi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427" start="Figures 6–11" startId="F6">
|
||
<paragraph id="5924A57DA06F54BD7E7888A0BD69DE36" pageId="0" pageNumber="427">
|
||
<emphasis id="87CF0B3B4BCC1AC3C3562DF445A37A9D" bold="true" pageId="0" pageNumber="427">Figures 6-11.</emphasis>
|
||
Pronotum of
|
||
<taxonomicName id="36AA6512D4D2B239C070833BEC2720D9" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C423B398DCD9B6A6161F24F26578427B" italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
|
||
</taxonomicName>
|
||
species
|
||
<emphasis id="4D81C037480979735C9712D4D700E247" bold="true" pageId="0" pageNumber="427">6</emphasis>
|
||
<taxonomicName id="0873A238D52A10A5E7454F9F6F56452F" lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis id="1A1A3ABA1E7C7827525FACF27C8204D7" italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
HT
|
||
<emphasis id="CE6E365A1C65DD607E6D5A39172B10B9" bold="true" pageId="0" pageNumber="427">7</emphasis>
|
||
<taxonomicName id="01F75060D062DFAB902F525DD6E9673D" lsidName="D. laevigotoides" pageId="0" pageNumber="427" rank="species" species="laevigotoides">
|
||
<emphasis id="B5681CAB58CADD9F0C25DAFB8AF76698" italics="true" pageId="0" pageNumber="427">D. laevigotoides</emphasis>
|
||
</taxonomicName>
|
||
HT
|
||
<emphasis id="13B6161D65FBF6AEBCE5A4CD5DCE17D5" bold="true" pageId="0" pageNumber="427">8</emphasis>
|
||
<taxonomicName id="90A4D464CEA0755EA99209880B9CCCC7" lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis id="074533F96FF69CFC55A4E2B75A6454A7" italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
from Cambodia
|
||
<emphasis id="BEE49EA94F3CF67D73D267D9A07F2013" bold="true" pageId="0" pageNumber="427">9</emphasis>
|
||
<taxonomicName id="F55468DFD600E992F4BF17BCF58FF884" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="95E0EAFC5B48C2115CC02946A8C82129" italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
|
||
</taxonomicName>
|
||
sp. from Thailand
|
||
<emphasis id="BA3A4AAAC6075342CE680B28DAA4EEA1" bold="true" pageId="0" pageNumber="427">10</emphasis>
|
||
<taxonomicName id="1984453E9AEBBD59CEEE5E7C8CC904A2" lsidName="D. walterrossii" pageId="0" pageNumber="427" rank="species" species="walterrossii">
|
||
<emphasis id="81019D8D31FDAB986120200D47D92691" italics="true" pageId="0" pageNumber="427">D. walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. HT
|
||
<emphasis id="740331EAC82567E2516A02336E9942B6" bold="true" pageId="0" pageNumber="427">11</emphasis>
|
||
<taxonomicName id="78A344283D72FAB5E17CA981C9DAD2D5" lsidName="D. erwini" pageId="0" pageNumber="427" rank="species" species="erwini">
|
||
<emphasis id="0C192C2CBEBDEFB599884E526CFE2E3C" italics="true" pageId="0" pageNumber="427">D. erwini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. HT.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="5D2A36548198A7421928E1AC2EC381B7" doi="10.3897/zookeys.1044.60072.figures12-17" httpUri="https://binary.pensoft.net/fig/554098" pageId="0" pageNumber="427" start="Figures 12–17" startId="F7">
|
||
<paragraph id="C822EA4A5E314904A75DA3DF61FB2668" pageId="0" pageNumber="427">
|
||
<emphasis id="BA698E85CC7E957823026CC0B98AFCB6" bold="true" pageId="0" pageNumber="427">Figures 12-17.</emphasis>
|
||
Labrum of
|
||
<taxonomicName id="B166495E38544E37AD0DBE960C25135E" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6DFC4997375D4EB581DBA10A0FA4084A" italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
|
||
</taxonomicName>
|
||
species
|
||
<emphasis id="A751AAB92A59BDD1AC86EA103083B03B" bold="true" pageId="0" pageNumber="427">12</emphasis>
|
||
<taxonomicName id="6223F7E8A71B42FC68EC27F5902A4B46" lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis id="B9D6C3D50922F60D279749645336CFFA" italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
HT
|
||
<emphasis id="BF58AF6F1D67C49CA8AC3D6738E69EFF" bold="true" pageId="0" pageNumber="427">13</emphasis>
|
||
<taxonomicName id="1BB62F4277126AA50E1EA9B1614F35C4" lsidName="D. laevigotoides" pageId="0" pageNumber="427" rank="species" species="laevigotoides">
|
||
<emphasis id="81D745C448F288ECE3E73FBF07B73BFD" italics="true" pageId="0" pageNumber="427">D. laevigotoides</emphasis>
|
||
</taxonomicName>
|
||
HT
|
||
<emphasis id="24FC35BEDB52DBAFCB608D373327FFE3" bold="true" pageId="0" pageNumber="427">14</emphasis>
|
||
<taxonomicName id="69D593E7468F8C5FD406446BD4395C66" lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis id="00DD2B6914F5E9AE75490764BAD10309" italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
from Cambodia
|
||
<emphasis id="AD81D1D5BD5459A5A6A9489189A5BD5A" bold="true" pageId="0" pageNumber="427">15</emphasis>
|
||
<taxonomicName id="0D61AF695069FE2E28B8E3B20FBD2878" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="30D7830A63CBD076275BFED82067263B" italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
|
||
</taxonomicName>
|
||
sp. from Thailand
|
||
<emphasis id="9842420551220C7C0FFF4D6B873F3F41" bold="true" pageId="0" pageNumber="427">16</emphasis>
|
||
<taxonomicName id="E6B2008368C8CAFCC1DF600D27CF8989" lsidName="D. walterrossii" pageId="0" pageNumber="427" rank="species" species="walterrossii">
|
||
<emphasis id="6171CC2F5D0D391A2D268B50ECBF72E0" italics="true" pageId="0" pageNumber="427">D. walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. HT
|
||
<emphasis id="249D293E9C1FF4AFBA31082C75C874E1" bold="true" pageId="0" pageNumber="427">17</emphasis>
|
||
<taxonomicName id="3AA4B79E79DF4DEE8BCCBC26EBB544AC" lsidName="D. erwini" pageId="0" pageNumber="427" rank="species" species="erwini">
|
||
<emphasis id="55A4FBC7BE3BAE2D4EE648BD1D58D87C" italics="true" pageId="0" pageNumber="427">D. erwini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. HT.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="25E41963F94869BC18C4D52B3E781F06" doi="10.3897/zookeys.1044.60072.figures18-19" httpUri="https://binary.pensoft.net/fig/554099" pageId="0" pageNumber="427" start="Figures 18–19" startId="F8">
|
||
<paragraph id="44D5DD924ECE3461F43A16331203909F" pageId="0" pageNumber="427">
|
||
<emphasis id="E0F58AEFFE92F67CFAD31E1BD173D4F1" bold="true" pageId="0" pageNumber="427">Figures 18-19.</emphasis>
|
||
Male protarsi of
|
||
<taxonomicName id="8E20B0EE484A0C042DBE92760A4EF395" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F83899FD9BD76851C02C0BA514B02A13" italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
|
||
</taxonomicName>
|
||
species
|
||
<emphasis id="3DC9E587B5ECFB357278EF1A85E29A41" bold="true" pageId="0" pageNumber="427">18</emphasis>
|
||
<taxonomicName id="4B2662F0218179376568881A0A0491DC" lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis id="D9078909B243E73BB8D199511ED76B2C" italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
from Cambodia
|
||
<emphasis id="4AABABED3D2E752269E36DF4DFA3BD02" bold="true" pageId="0" pageNumber="427">19</emphasis>
|
||
<taxonomicName id="66A4B645BCD5D872396C2616552D824C" lsidName="D. walterrossii" pageId="0" pageNumber="427" rank="species" species="walterrossii">
|
||
<emphasis id="8ECC8B11C4A506B982E7763D51A7E059" italics="true" pageId="0" pageNumber="427">D. walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. HT.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |