262 lines
21 KiB
XML
262 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.131.36548" ID-GBIF-Dataset="6fe68ec7-4966-41aa-8e1d-f91a68be1a0d" ID-PMC="PMC6742672" ID-Pensoft-Pub="1314-2003-131-69" ID-Pensoft-UUID="04D6BB3D36805DE6A593FC9390C287AC" ID-PubMed="31565025" ID-Zenodo-Dep="3406905" ModsDocID="1314-2003-131-69" checkinTime="1567745835896" checkinUser="pensoft" docAuthor="Chao, Yi-Shan, Huang, Yu-Fang, Dong, Shi-Yong, Huang, Yao-Moan & Liu, Ho-Yih" docDate="2019" docId="DA4F28BC607A5FE28156C1F219A0C87A" docLanguage="en" docName="PhytoKeys 131: 69-81" docOrigin="PhytoKeys 131" docSource="http://dx.doi.org/10.3897/phytokeys.131.36548" docTitle="Bolbitis lianhuachihensis Y. S. Chao, Y. F. Huang, & H. Y. Liu 2019, sp. nov." docType="treatment" docVersion="6" id="04D6BB3D36805DE6A593FC9390C287AC" lastPageNumber="75" masterDocId="04D6BB3D36805DE6A593FC9390C287AC" masterDocTitle="Bolbitis lianhuachihensis (Dryopteridaceae), a new species from Taiwan" masterLastPageNumber="81" masterPageNumber="69" pageNumber="74" updateTime="1668139486795" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Bolbitis lianhuachihensis (Dryopteridaceae), a new species from Taiwan</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chao, Yi-Shan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huang, Yu-Fang</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dong, Shi-Yong</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huang, Yao-Moan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liu, Ho-Yih</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>131</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>69</mods:start>
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<mods:end>81</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.131.36548</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.131.36548</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-131-69</mods:identifier>
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<mods:identifier type="Pensoft-UUID">04D6BB3D36805DE6A593FC9390C287AC</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3406905</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159413351" LSID="urn:lsid:plazi:treatment:DA4F28BC607A5FE28156C1F219A0C87A" httpUri="http://treatment.plazi.org/id/DA4F28BC607A5FE28156C1F219A0C87A" lastPageId="6" lastPageNumber="75" pageId="5" pageNumber="74">
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<subSubSection pageId="5" pageNumber="74" type="nomenclature">
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<paragraph pageId="5" pageNumber="74">
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<taxonomicName LSID="DA4F28BC-607A-5FE2-8156-C1F219A0C87A" authority="Y. S. Chao, Y. F. Huang, & H. Y. Liu" authorityName="Y. S. Chao, Y. F. Huang, & H. Y. Liu" authorityYear="2019" class="Polypodiopsida" family="Dryopteridaceae" genus="Bolbitis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bolbitis lianhuachihensis" order="Polypodiales" pageId="5" pageNumber="74" phylum="Pteridophyta" rank="species" species="lianhuachihensis" status="sp. nov.">Bolbitis lianhuachihensis Y.S.Chao, Y.F.Huang, & H.Y.Liu</taxonomicName>
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<taxonomicNameLabel pageId="5" pageNumber="74">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bolbitis lianhuachihensis A habit; note taller fertile fronds (photographed by Y. - S. Chao) B acrostichoid sori (photographed by P. - F. Lu) C venation of sterile frond (photographed by Y. - F. Huang)." figureDoi="10.3897/phytokeys.131.36548.figure1" httpUri="https://binary.pensoft.net/fig/331240" pageId="5" pageNumber="74" tableDoi="10.3897/phytokeys.131.36548.figure1">Fig. 1</figureCitation>
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and Suppl. material 1: Figures S1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="74" type="reference_group">
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<paragraph pageId="5" pageNumber="74">
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<taxonomicName authorityName="Hennipman" authorityYear="1970" class="Polypodiopsida" family="Dryopteridaceae" genus="Bolbitis" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Plantae" lsidName="Bolbitis virens var. compacta" order="Polypodiales" pageId="5" pageNumber="74" phylum="Pteridophyta" rank="variety" species="virens" variety="compacta">Bolbitis virens var. compacta</taxonomicName>
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auct. non Hennipman: Knapp, Ferns Fern Allies Taiwan: 440. 2011.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="74" type="materials_examined">
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<paragraph pageId="5" pageNumber="74">Type.</paragraph>
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<paragraph pageId="5" pageNumber="74">
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TAIWAN. Nantou County: Lienhuachih, 700 m a.s.l., 11 March 2018,
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<emphasis italics="true" pageId="5" pageNumber="74">Yi-Shan Chao 3006</emphasis>
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(holotype TAIF!, isotype TAIF!).
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</paragraph>
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<paragraph pageId="5" pageNumber="74">
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<taxonomicName authorityName="Y. S. Chao, Y. F. Huang, & H. Y. Liu" authorityYear="2019" class="Polypodiopsida" family="Dryopteridaceae" genus="Bolbitis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bolbitis lianhuachihensis" order="Polypodiales" pageId="5" pageNumber="74" phylum="Pteridophyta" rank="species" species="lianhuachihensis">
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<emphasis bold="true" italics="true" pageId="5" pageNumber="74">Bolbitis lianhuachihensis</emphasis>
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</taxonomicName>
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morphologically resembles
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<taxonomicName infraspecific-rank="var." lsidName="B. virens var. compacta" pageId="5" pageNumber="74" rank="variety" species="virens" variety="compacta">B. virens var. compacta</taxonomicName>
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and
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<taxonomicName lsidName="B. hainanensis" pageId="5" pageNumber="74" rank="species" species="hainanensis">
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<emphasis italics="true" pageId="5" pageNumber="74">B. hainanensis</emphasis>
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</taxonomicName>
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, from which it is distinguished in having lanceolate sterile pinnae and no free veinlets in areoles of sterile fronds.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="75" pageId="5" pageNumber="74" type="description">
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<paragraph pageId="5" pageNumber="74">Description.</paragraph>
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<paragraph lastPageId="6" lastPageNumber="75" pageId="5" pageNumber="74">
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Terrestrial or lithophytic. Rhizome short-creeping, thick, densely scaly; scales concolorous, black, lanceolate, 3-5 mm long, 0.5 mm wide, margin entire. Fronds clustered, 40-80 cm long, dimorphic, pinnate. Sterile fronds with stipes 18-50 cm long, near base 2-4 mm diam., scaly, glabrous upwards; lamina broad-ovate, 21-34
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<normalizedToken originalValue="×">x</normalizedToken>
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21-35 cm, chartaceous, conform; lateral pinnae 1-5 pairs, alternate, lanceolate,
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<pageBreakToken pageId="6" pageNumber="75" start="start">14</pageBreakToken>
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-24
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<normalizedToken originalValue="×">x</normalizedToken>
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3-4.5 cm, bases cuneate, margins entire, undulate, apices caudate or acuminate, basal two pairs of pinnae with winged petiolules <8 mm; terminal pinna larger or similar to lateral pinnae, sometimes with a bulbil near the apex; veins reticulate, 2-4 rows, no or very few areoles with free veinlets in the largest pinnae (wider than 3.5 cm). Fertile fronds longer than or as long as the sterile ones; stipes 35-50 cm long; lamina oblong-ovate, 7.5-14.5
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<normalizedToken originalValue="×">x</normalizedToken>
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12-18 cm; pinnae 3-4 pairs, alternate, lanceolate, 5-7.5
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<normalizedToken originalValue="×">x</normalizedToken>
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0.8-1.2 cm, base narrow-cuneate, apex acuminate, stalked. Sporangia acrostichoid.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="75" type="materials_examined">
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<paragraph pageId="6" pageNumber="75">Additional specimens examined.</paragraph>
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<paragraph pageId="6" pageNumber="75">
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<emphasis bold="true" pageId="6" pageNumber="75">TAIWAN. Chiayi County</emphasis>
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: Tsenwen Dam, 24 Jan. 1987,
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<emphasis italics="true" pageId="6" pageNumber="75">Bi-Jao Wang 10039</emphasis>
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,
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<emphasis italics="true" pageId="6" pageNumber="75">11009</emphasis>
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(HAST, TAIF).
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<emphasis bold="true" pageId="6" pageNumber="75">Nantou County</emphasis>
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: Lienhuachih, 700 m a.s.l.,
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<emphasis italics="true" pageId="6" pageNumber="75">Yih-Han Chang 20070317-008</emphasis>
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(TAIF), 29 Aug. 2009,
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<emphasis italics="true" pageId="6" pageNumber="75">Cheng-Wei Chen Wade 955</emphasis>
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(TAIF), 22 Mar. 2015,
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<emphasis italics="true" pageId="6" pageNumber="75">Cheng-Wei Chen Wade 4181</emphasis>
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(TAIF), 29 Jan. 2016,
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<emphasis italics="true" pageId="6" pageNumber="75">Yu-Fang Huang 115</emphasis>
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,
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<emphasis italics="true" pageId="6" pageNumber="75">116</emphasis>
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,
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<emphasis italics="true" pageId="6" pageNumber="75">117</emphasis>
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(TAIF), 10 Oct. 2005,
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<emphasis italics="true" pageId="6" pageNumber="75">Ralf Knapp 697</emphasis>
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(P), 22 Oct. 2011,
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<emphasis italics="true" pageId="6" pageNumber="75">
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Ralf
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<bibRefCitation author="Knapp, R" journalOrPublisher="KBCC Press, Pingtung and Yuan-Liou Pub, Taipei" pageId="8" pageNumber="77" refId="B9" refString="Knapp, R, 2011. Ferns and Fern Allies of Taiwan. KBCC Press, Pingtung and Yuan-Liou Pub, Taipei" title="Ferns and Fern Allies of Taiwan." year="2011">Knapp 2011</bibRefCitation>
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1022-4
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</emphasis>
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(HAST), 4 Dec. 2012,
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<emphasis italics="true" pageId="6" pageNumber="75">Pi-Fong Lu 24940</emphasis>
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(TAIF), 23 Apr. 2006,
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<emphasis italics="true" pageId="6" pageNumber="75">Wei-Hsiu Wu s.n.</emphasis>
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(TAIF); Tiandi, 830 m a.s.l., 8 Aug. 2006,
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<emphasis italics="true" pageId="6" pageNumber="75">Ralf Knapp 20060806-18</emphasis>
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(HAST, TAIF), 900 m a.s.l., 26 Sep. 2016,
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<emphasis italics="true" pageId="6" pageNumber="75">Yu-Fang Huang 172</emphasis>
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,
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<emphasis italics="true" pageId="6" pageNumber="75">173</emphasis>
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,
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<emphasis italics="true" pageId="6" pageNumber="75">174</emphasis>
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(TAIF).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="75" type="distribution">
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<paragraph pageId="6" pageNumber="75">Distribution.</paragraph>
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<paragraph pageId="6" pageNumber="75">
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Taiwan (
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of Bolbitis lianhuachihensis (black circles) in Taiwan." figureDoi="10.3897/phytokeys.131.36548.figure4" httpUri="https://binary.pensoft.net/fig/331243" pageId="6" pageNumber="75" tableDoi="10.3897/phytokeys.131.36548.figure4">Fig. 4</figureCitation>
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).
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</paragraph>
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<caption ID-Zenodo-Dep="3487965" doi="10.3897/phytokeys.131.36548.figure4" httpUri="https://binary.pensoft.net/fig/331243" pageId="6" pageNumber="75" start="Figure 4" startId="F4">
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<paragraph pageId="6" pageNumber="75">
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<emphasis bold="true" pageId="6" pageNumber="75">Figure 4.</emphasis>
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Distribution of
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<taxonomicName authorityName="Y. S. Chao, Y. F. Huang, & H. Y. Liu" authorityYear="2019" class="Polypodiopsida" family="Dryopteridaceae" genus="Bolbitis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bolbitis lianhuachihensis" order="Polypodiales" pageId="6" pageNumber="75" phylum="Pteridophyta" rank="species" species="lianhuachihensis">
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<emphasis italics="true" pageId="6" pageNumber="75">Bolbitis lianhuachihensis</emphasis>
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</taxonomicName>
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(black circles) in Taiwan.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="75" type="ecology">
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<paragraph pageId="6" pageNumber="75">Ecology.</paragraph>
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<paragraph pageId="6" pageNumber="75">Evergreen, broad-leaf forests, often near streams, below 1000 m a.s.l.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="75" type="etymology">
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<paragraph pageId="6" pageNumber="75">Etymology.</paragraph>
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<paragraph pageId="6" pageNumber="75">
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The specific epithet '
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<taxonomicName authorityName="Y. S. Chao, Y. F. Huang, & H. Y. Liu" authorityYear="2019" class="Polypodiopsida" family="Dryopteridaceae" genus="Bolbitis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="lianhuachihensis" order="Polypodiales" pageId="6" pageNumber="75" phylum="Pteridophyta" rank="species" species="lianhuachihensis">lianhuachihensis</taxonomicName>
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' refers to the type locality.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="75" type="common name (assigned here)">
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<paragraph pageId="6" pageNumber="75">Common name (assigned here).</paragraph>
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<paragraph pageId="6" pageNumber="75">Lian Hua Chih Shih Jyue (蓮華池實蕨; Chinese name).</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="75" type="preliminary conservation assessment">
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<paragraph pageId="6" pageNumber="75">Preliminary conservation assessment.</paragraph>
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<paragraph pageId="6" pageNumber="75">
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To date, only three small populations of
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<taxonomicName authorityName="Y. S. Chao, Y. F. Huang, & H. Y. Liu" authorityYear="2019" class="Polypodiopsida" family="Dryopteridaceae" genus="Bolbitis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bolbitis lianhuachihensis" order="Polypodiales" pageId="6" pageNumber="75" phylum="Pteridophyta" rank="species" species="lianhuachihensis">
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<emphasis italics="true" pageId="6" pageNumber="75">Bolbitis lianhuachihensis</emphasis>
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</taxonomicName>
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Y.S.Chao, Y.F.Huang, & H.Y.Liu have been recorded in Taiwan. However, the estimated number of individuals is smaller than 250. It meets the category Endangered (EN D1) based on the
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<bibRefCitation author="IUCN" journalOrPublisher="Standards and Petitions Subcommittee, Gland and Cambridge" pageId="8" pageNumber="77" refId="B7" refString="IUCN, 2017. Guidelines for using the IUCN red list categories and criteria. version 13. Standards and Petitions Subcommittee, Gland and Cambridge" title="Guidelines for using the IUCN red list categories and criteria. version 13." year="2017">IUCN (2017)</bibRefCitation>
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criteria.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="75" type="notes">
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<paragraph pageId="6" pageNumber="75">Note.</paragraph>
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<paragraph pageId="6" pageNumber="75">
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<taxonomicName authorityName="Y. S. Chao, Y. F. Huang, & H. Y. Liu" authorityYear="2019" class="Polypodiopsida" family="Dryopteridaceae" genus="Bolbitis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bolbitis lianhuachihensis" order="Polypodiales" pageId="6" pageNumber="75" phylum="Pteridophyta" rank="species" species="lianhuachihensis">
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<emphasis italics="true" pageId="6" pageNumber="75">Bolbitis lianhuachihensis</emphasis>
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</taxonomicName>
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can be delimited by the combination of anastomosing venation and fewer sterile pinnae than any other species of
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<taxonomicName class="Polypodiopsida" family="Dryopteridaceae" genus="Bolbitis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bolbitis" order="Polypodiales" pageId="6" pageNumber="75" phylum="Pteridophyta" rank="genus">
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<emphasis italics="true" pageId="6" pageNumber="75">Bolbitis</emphasis>
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</taxonomicName>
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in Taiwan. It was thought to be related to two similar taxa with "thick laminae" in nearby regions,
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<taxonomicName infraspecific-rank="var." lsidName="B. virens var. compacta" pageId="6" pageNumber="75" rank="variety" species="virens" variety="compacta">B. virens var. compacta</taxonomicName>
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and
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<taxonomicName lsidName="B. hainanensis" pageId="6" pageNumber="75" rank="species" species="hainanensis">
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<emphasis italics="true" pageId="6" pageNumber="75">B. hainanensis</emphasis>
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</taxonomicName>
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(
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<bibRefCitation author="Knapp, R" journalOrPublisher="KBCC Press, Pingtung and Yuan-Liou Pub, Taipei" pageId="8" pageNumber="77" refId="B9" refString="Knapp, R, 2011. Ferns and Fern Allies of Taiwan. KBCC Press, Pingtung and Yuan-Liou Pub, Taipei" title="Ferns and Fern Allies of Taiwan." year="2011">Knapp 2011</bibRefCitation>
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). However, based on our study, only
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<taxonomicName lsidName="B. hainanensis" pageId="6" pageNumber="75" rank="species" species="hainanensis">
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<emphasis italics="true" pageId="6" pageNumber="75">B. hainanensis</emphasis>
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</taxonomicName>
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has coriaceous laminae, thicker than chartaceous laminar of
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<taxonomicName lsidName="B. lianhuachihensis" pageId="6" pageNumber="75" rank="species" species="lianhuachihensis">
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<emphasis italics="true" pageId="6" pageNumber="75">B. lianhuachihensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName infraspecific-rank="var." lsidName="B. virens var. compacta" pageId="6" pageNumber="75" rank="variety" species="virens" variety="compacta">B. virens var. compacta</taxonomicName>
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. Moreover,
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<taxonomicName lsidName="B. lianhuachihensis" pageId="6" pageNumber="75" rank="species" species="lianhuachihensis">
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<emphasis italics="true" pageId="6" pageNumber="75">B. lianhuachihensis</emphasis>
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</taxonomicName>
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has lanceolate sterile pinnae and very few free veinlets in areoles of sterile pinnae wider than 3.5 cm, differing from the two similar taxa.
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</paragraph>
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<paragraph pageId="6" pageNumber="75">
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In this study, we revealed the venation diversity in the Taiwanese
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<taxonomicName class="Polypodiopsida" family="Dryopteridaceae" genus="Bolbitis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bolbitis" order="Polypodiales" pageId="6" pageNumber="75" phylum="Pteridophyta" rank="genus">
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<emphasis italics="true" pageId="6" pageNumber="75">Bolbitis</emphasis>
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</taxonomicName>
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species and related taxa. Based on the illustration of venation, it is found that every taxon possesses its own venation morphology, supporting the taxonomic value of venation (
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<bibRefCitation author="Hennipman, E" journalOrPublisher="Leiden University Press, Leiden" pageId="8" pageNumber="77" refId="B6" refString="Hennipman, E, 1977. A Monograph of the Fern Genus Bolbitis (Lomariopsidaceae). Leiden University Press, Leiden" title="A Monograph of the Fern Genus Bolbitis (Lomariopsidaceae)." year="1977">Hennipman 1977</bibRefCitation>
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;
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<bibRefCitation author="Moran, RC" journalOrPublisher="International Journal of Plant Sciences" pageId="9" pageNumber="78" pagination="547 - 559" refId="B13" refString="Moran, RC, Labiak, PH, Sundue, M, 2010a. Phylogeny and character evolution of the Bolbitidoid ferns (Dryopteridaceae). International Journal of Plant Sciences 171 (5): 547 - 559" title="Phylogeny and character evolution of the Bolbitidoid ferns (Dryopteridaceae)." volume="171" year="2010 a">Moran et al. 2010a</bibRefCitation>
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). We have applied several characters, including angles of veinlets to costae, free or anastomosing, row number of areoles between the costae and margins in sterile fronds and free veinlets in areoles or not for sterile fronds. We also found that venation patterns are more complicated, composed of multiple areoles, sub-areoles and free veinlets, which is also reported by
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<bibRefCitation author="Hennipman, E" journalOrPublisher="Leiden University Press, Leiden" pageId="8" pageNumber="77" refId="B6" refString="Hennipman, E, 1977. A Monograph of the Fern Genus Bolbitis (Lomariopsidaceae). Leiden University Press, Leiden" title="A Monograph of the Fern Genus Bolbitis (Lomariopsidaceae)." year="1977">Hennipman (1977)</bibRefCitation>
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. A character-state change from free venation to anastomosing venation is reported by
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<bibRefCitation author="Moran, RC" journalOrPublisher="International Journal of Plant Sciences" pageId="9" pageNumber="78" pagination="547 - 559" refId="B13" refString="Moran, RC, Labiak, PH, Sundue, M, 2010a. Phylogeny and character evolution of the Bolbitidoid ferns (Dryopteridaceae). International Journal of Plant Sciences 171 (5): 547 - 559" title="Phylogeny and character evolution of the Bolbitidoid ferns (Dryopteridaceae)." volume="171" year="2010 a">Moran et al. (2010a)</bibRefCitation>
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. The venation characteristics in
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<taxonomicName class="Polypodiopsida" family="Dryopteridaceae" genus="Bolbitis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bolbitis" order="Polypodiales" pageId="6" pageNumber="75" phylum="Pteridophyta" rank="genus">
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<emphasis italics="true" pageId="6" pageNumber="75">Bolbitis</emphasis>
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</taxonomicName>
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species are worthy of further investigation and application to the systematic and evolutionary study. To classify and describe the diverse venation morphology in detail would contribute to further studies of
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<taxonomicName class="Polypodiopsida" family="Dryopteridaceae" genus="Bolbitis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bolbitis" order="Polypodiales" pageId="6" pageNumber="75" phylum="Pteridophyta" rank="genus">
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<emphasis italics="true" pageId="6" pageNumber="75">Bolbitis</emphasis>
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</taxonomicName>
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systematics.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |