339 lines
36 KiB
XML
339 lines
36 KiB
XML
<document ID-CLB-Dataset="8822" ID-DOI="http://dx.doi.org/10.3897/zookeys.1011.59449" ID-GBIF-Dataset="03523ad0-0923-4dbc-9842-914b124243fe" ID-PMC="PMC7835194" ID-Pensoft-Pub="1313-2970-1011-41" ID-Pensoft-UUID="4593F2AF44DB5B85848EF850C01AF05B" ID-PubMed="33551648" ID-ZooBank="E8A208E41FE648558FFC34671103342E" ModsDocID="1313-2970-1011-41" checkinTime="1611008234243" checkinUser="pensoft" docAuthor="Wrase, David W. & Assmann, Thorsten" docDate="2021" docId="D8870D7027695E25886F2841BFA5ABA5" docLanguage="en" docName="ZooKeys 1011: 41-50" docOrigin="ZooKeys 1011" docPubDate="2021-01-18" docSource="http://dx.doi.org/10.3897/zookeys.1011.59449" docTitle="Parazuphium weigeli Wrase & Assmann 2021, sp. nov." docType="treatment" docUuid="69A63791-3CDF-4CCF-A0AE-B359EC350BF7" docUuidSource="ZooBank" docVersion="6" id="4593F2AF44DB5B85848EF850C01AF05B" lastPageNumber="41" masterDocId="4593F2AF44DB5B85848EF850C01AF05B" masterDocTitle="A new Parazuphium Jeannel, 1942 species (Coleoptera, Carabidae) from the Zagros Mountains in Iran" masterLastPageNumber="50" masterPageNumber="41" pageNumber="41" updateTime="1732865050421" updateUser="ExternalLinkService">
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<mods:titleInfo id="5745E828430B38C586B51C865C2EE19E">
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<mods:title id="1F68F46D8F2E7A60408A39CF706C6E56">A new Parazuphium Jeannel, 1942 species (Coleoptera, Carabidae) from the Zagros Mountains in Iran</mods:title>
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<mods:name id="AEB9CD9B6832CC7DEFDBEBE431D8D58A" type="personal">
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<mods:roleTerm id="13B314EBB5B6D60E8A1FFD59F69D1FB8">Author</mods:roleTerm>
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<mods:namePart id="A4324134B5C8E0880FD146B3CD7AA0A3">Wrase, David W.</mods:namePart>
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<mods:affiliation id="399F261102AB767FC511E5F494557EEF">Oderstrasse 2, D- 15306, Gusow-Platkow, Germany</mods:affiliation>
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<mods:roleTerm id="B2391FC8D09471863DD2D036A1C47B6B">Author</mods:roleTerm>
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<mods:namePart id="68C6AD278E783E1A994456547EC3E3F5">Assmann, Thorsten</mods:namePart>
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<mods:nameIdentifier id="17E268E5543B5AF1E67857FF71AD4790" type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
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<mods:affiliation id="2EC73A7F77DF5901F676327CD34DB2D0">Institute of Ecology, Leuphana University Lueneburg, Universitaetsallee 1, D- 21331, Lueneburg, Germany</mods:affiliation>
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<mods:nameIdentifier id="9291E0D4D693AB21EDB3CE8D0624D7B3" type="email">assmann@uni.leuphana.de</mods:nameIdentifier>
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<mods:typeOfResource id="285AD604AD8E18562DF6007869E20713">text</mods:typeOfResource>
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<mods:titleInfo id="341511BEE42033103361975906CC17DA">
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<mods:title id="9AB4727A64EFEF72990E79B188B8E9C4">ZooKeys</mods:title>
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<mods:part id="D277A84212DDDF797ACC8596FCF1193A">
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<mods:date id="AD95ED57689E77E57BF1DBF61ED1BA9A">2021</mods:date>
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<mods:detail id="BCEACFB565F9F3AEDD79181F1154E1BB" type="pubDate">
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<mods:number id="6391CB9AB61716F26D99E6C33BA6A25E">2021-01-18</mods:number>
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<mods:detail id="7D75917282A95556567E8C5079A40F44" type="volume">
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<mods:number id="981A25AF8D5883493622AF56353B4257">1011</mods:number>
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<mods:start id="4D8F20142DDB738FC7D3FE6BA8CC90E7">41</mods:start>
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<mods:location id="6E9588FA822085A7DD8EEDCBA5FB24AB">
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<mods:url id="1AC00894FA2CC6A49E9ADD24C953CEA0">http://dx.doi.org/10.3897/zookeys.1011.59449</mods:url>
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<mods:classification id="D19E5E08FA83C4BA4AAB4C3F2A80C69F">journal article</mods:classification>
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<mods:identifier id="AD6724BBDE815F39111990E2E610B5A7" type="DOI">http://dx.doi.org/10.3897/zookeys.1011.59449</mods:identifier>
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<mods:identifier id="73AB8B431473070B414D28F13B791445" type="Pensoft-Pub">1313-2970-1011-41</mods:identifier>
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<mods:identifier id="4675489CD24D6C1A0AC57CB4D8B65FC9" type="ZooBank">E8A208E41FE648558FFC34671103342E</mods:identifier>
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<treatment id="D8870D7027695E25886F2841BFA5ABA5" ID-GBIF-Taxon="177006818" LSID="urn:lsid:zoobank.org:act:69A63791-3CDF-4CCF-A0AE-B359EC350BF7" httpUri="http://treatment.plazi.org/id/D8870D7027695E25886F2841BFA5ABA5" lastPageNumber="41" pageId="0" pageNumber="41" scope_family="Carabidae" scope_order="Coleoptera">
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<subSubSection id="72CD5A3F64FBCBB5F7AB81D41B4E9F8E" pageId="0" pageNumber="41" type="nomenclature">
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<paragraph id="43809250B57A5F29410FC037B5EA41FA" pageId="0" pageNumber="41">
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<taxonomicName id="C3E8879F91C7605DF81291B2B086A12B" ID-CoL="8MDDH" LSID="http://zoobank.org/69A63791-3CDF-4CCF-A0AE-B359EC350BF7" authority="Wrase & Assmann, 2021" authorityName="Wrase & Assmann" authorityYear="2021" class="Insecta" family="Carabidae" genus="Parazuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazuphium weigeli" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="weigeli" status="sp. nov.">Parazuphium weigeli</taxonomicName>
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<taxonomicNameLabel id="3C1A9D1061A6D16B4690A1E2A5E3A2FC" pageId="0" pageNumber="41">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="4021F909F18B77C86C7190932F894A37" pageId="0" pageNumber="41" type="types">
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<paragraph id="F9F4C0C9537CCBAA37C1507A0E60DBD8" pageId="0" pageNumber="41">Types.</paragraph>
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<paragraph id="FEDA74434AC82467B423E9A300202DF8" pageId="0" pageNumber="41">
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<emphasis id="75F10380CF00D46AFBB71804D9F328EA" bold="true" italics="true" pageId="0" pageNumber="41">Holotype</emphasis>
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male: "Iran Zagros Mts., P. Chahar / Mahall va Bachtiari, Asad / Abad 5 km SW,
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<geoCoordinate id="E272503DF04A33101ECF885150DCC96A" degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30"N</geoCoordinate>
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,
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<geoCoordinate id="04545A1BF59E730969C463527DD3A7A2" degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59"E</geoCoordinate>
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, snow field, high pasture, 19.IV.2018, 2500- / 2770 m, leg. A.Weigel" (NKME; left and right antennomeres X and XI are missing). Paratype: 1 female, same data as HT (cWR).
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</paragraph>
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</subSubSection>
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<subSubSection id="C0CA14DAE8B55C221147EB1902940573" pageId="0" pageNumber="41" type="diagnosis">
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<paragraph id="92C61F6FB2D6504C5604923BD28FE62D" pageId="0" pageNumber="41">Diagnosis.</paragraph>
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<paragraph id="EFC00B7D76CC382792C21E5A004AC1FB" pageId="0" pageNumber="41">
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A microphthalmic, depigmented, brachypterous
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<taxonomicName id="E4EE421589090955270595C17D8E29E8" authorityName="Jeannel" authorityYear="1942" class="Insecta" family="Carabidae" genus="Parazuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazuphium" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis id="2B6F005308A13E214D625FD7EFD2669A" italics="true" pageId="0" pageNumber="41">Parazuphium</emphasis>
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</taxonomicName>
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species with short, robust legs and moderately long antennae. Median lobe of aedeagus with three sclerites. For habitus see Fig.
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<figureCitation id="277246DB655EA0A6C4D13605DA86E405" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Parazuphium weigeli sp. nov., paratype, female." figureDoi="10.3897/zookeys.1011.59449.figure1" httpUri="https://binary.pensoft.net/fig/497914" pageId="0" pageNumber="41">1</figureCitation>
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.
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</paragraph>
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<caption id="AA900830F42A1A69B6751A067B826327" doi="10.3897/zookeys.1011.59449.figure1" httpUri="https://binary.pensoft.net/fig/497914" pageId="0" pageNumber="41" start="Figure 1" startId="F1">
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<paragraph id="90B9E6702E8DCF99835342A30B72AED8" pageId="0" pageNumber="41">
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<emphasis id="46B2D02F9CC022A15D499C4AD3DDE483" bold="true" pageId="0" pageNumber="41">Figure 1.</emphasis>
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Habitus of
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<taxonomicName id="92D8E30BF8B0106FE0860A82430648C9" authorityName="Wrase & Assmann" authorityYear="2021" class="Insecta" family="Carabidae" genus="Parazuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazuphium weigeli" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="weigeli">
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<emphasis id="104760C6C55B14F31135FBB0CB7F8B78" italics="true" pageId="0" pageNumber="41">Parazuphium weigeli</emphasis>
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</taxonomicName>
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sp. nov., paratype, female.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="F9F4B022B48F0D4B7A7BAAC27ECD2DE5" pageId="0" pageNumber="41" type="description">
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<paragraph id="D1A5325F827D2B7EDA7BB93ECECE47A0" pageId="0" pageNumber="41">Description.</paragraph>
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<paragraph id="818879715D0D4CD9F14A706EB80538E8" pageId="0" pageNumber="41">BL 3.7-3.9 mm; EW 1.3-1.4 mm.</paragraph>
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<paragraph id="F116C97B1EED49403419C595E177301B" pageId="0" pageNumber="41">
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<emphasis id="557FE66670753B6E58C4B8BF2317909A" bold="true" italics="true" pageId="0" pageNumber="41">Colour</emphasis>
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: Light yellowish to brownish.
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</paragraph>
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<paragraph id="A92986D3DE0AAEB972DF016E0A0EB8D0" pageId="0" pageNumber="41">
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<emphasis id="7E43F948666CA2D05FD933AFC22026B9" bold="true" italics="true" pageId="0" pageNumber="41">Head</emphasis>
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large, slightly triangular, with rounded tempora. Eyes small and flat, ~ 1/3 as long as tempora (dorsal view) (Figs
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<figureCitation id="FDA1C5C3282E68AF12817D2015C7935B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Parazuphium weigeli sp. nov., paratype, female." figureDoi="10.3897/zookeys.1011.59449.figure1" httpUri="https://binary.pensoft.net/fig/497914" pageId="0" pageNumber="41">1</figureCitation>
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,
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<figureCitation id="30EE1C0FE27F8DA939D9B88627758AAB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Parazuphium weigeli sp. nov. A head in lateral view (holotype) B apex of elytra (paratype)." figureDoi="10.3897/zookeys.1011.59449.figure2" httpUri="https://binary.pensoft.net/fig/497915" pageId="0" pageNumber="41">2a</figureCitation>
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). Antennae moderately long, scapus (antennomere I) somewhat shorter than width of head (A1L/HW 0.82-0.83), ~ 5
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<normalizedToken id="5F1180FFBD058F80144BC670CB7B6166" originalValue="×">x</normalizedToken>
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as long as antennomere II (A1L/A2L 4.7-5.17), antennomere II somewhat longer than 1/2 antennomere III (A2L/A3L 0.54-0.58), antennomere III somewhat shorter than antennomere IV (A3L/A4L 0.92). Neck somewhat wider than 1/3 of head width. Surface moderately shiny, punctures somewhat indistinct and scattered, setae also fine and scattered. Microsculpture consisting of isodiametric meshes.
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</paragraph>
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<caption id="EFF27AD2AE46F9151BDC4F46F1C8BECC" doi="10.3897/zookeys.1011.59449.figure2" httpUri="https://binary.pensoft.net/fig/497915" pageId="0" pageNumber="41" start="Figure 2" startId="F2">
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<paragraph id="D44207E37BED9F6E8D7A0E2D15658446" pageId="0" pageNumber="41">
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<emphasis id="0158F9738C53037D712A1E66B22ED88A" bold="true" pageId="0" pageNumber="41">Figure 2.</emphasis>
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<taxonomicName id="274F00A76E93F95152EC18C7134477B2" authorityName="Wrase & Assmann" authorityYear="2021" class="Insecta" family="Carabidae" genus="Parazuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazuphium weigeli" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="weigeli">
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<emphasis id="A6150EB13BDF0430B94C90727D9C2D3E" italics="true" pageId="0" pageNumber="41">Parazuphium weigeli</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="C7AA4899BF065EF96F97ED41815CF290" bold="true" pageId="0" pageNumber="41">A</emphasis>
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head in lateral view (holotype)
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<emphasis id="E4DCDA9F849B652154FDA4EA2B4CB3CB" bold="true" pageId="0" pageNumber="41">B</emphasis>
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apex of elytra (paratype).
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</paragraph>
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</caption>
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<paragraph id="7E2825AACB2D4DDD139E9ED3E24E7D31" pageId="0" pageNumber="41">
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<emphasis id="2A251643082C2F74405BFE32999113D1" bold="true" italics="true" pageId="0" pageNumber="41">Pronotum</emphasis>
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somewhat wider than head (HW/PW 0.88-0.91), slightly longer than wide (PW/PL 0.92), widest at approximately the apical 1/6, where lateral seta is inserted. Anterior margin almost rectilinear, somewhat excavated at insertion of neck. Anterior angles rounded, but hardly prominent. Lateral margin in apical 2/3 continuously rounded, concavely curved at the prebasal excision. Posterior angles obtuse and weakly prominent (PBaW/PEW 1.02-1.05), distinctly shifted forward due to a strong excision of the basal margin (Fig.
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<figureCitation id="31618388B73529C797C00AD224127228" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pronotum of Parazuphium A P. weigeli sp. nov. B P. salmoni." figureDoi="10.3897/zookeys.1011.59449.figure4" httpUri="https://binary.pensoft.net/fig/497917" pageId="0" pageNumber="41">4A</figureCitation>
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). Disc flat, median line medially somewhat excavated. Median sulcus fine, impressed on the posterior margin, not reaching the basal margin. Regularly and more strongly punctate and setose than head, microsculpture with irregular transverse meshes.
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</paragraph>
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<paragraph id="30580DEA8AC4F524D928B835ACB61E85" pageId="0" pageNumber="41">
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<emphasis id="B8DC25DA747516A3226617BC30D74418" bold="true" italics="true" pageId="0" pageNumber="41">Elytra</emphasis>
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short (EL/EW 1.28-1.31), subparallel, distinctly widening to the apex. Apical margin sinuous (Fig.
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<figureCitation id="132293CE235C093F4BFC5BB17FAED723" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Parazuphium weigeli sp. nov. A head in lateral view (holotype) B apex of elytra (paratype)." figureDoi="10.3897/zookeys.1011.59449.figure2" httpUri="https://binary.pensoft.net/fig/497915" pageId="0" pageNumber="41">2b</figureCitation>
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). Striae weak, only intimated, somewhat irregular, internal intervals weakly convex. Punctuation denser than on pronotum, somewhat wrinkled. Surface with setosity formed by short fine setae (somewhat denser than on pronotum), inclined backwards, setae at apex somewhat longer. Series umbilicata consists of eight (five long and three short) humeral and five (three long and two short) apical setae. Surface moderately shiny, microsculpture with irregular meshes. Brachypterous.
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</paragraph>
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<paragraph id="3DE78FFCDCF9BE72051D290A06A962EB" pageId="0" pageNumber="41">
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<emphasis id="BECAE900E5F74631CAED2EF2E813FE72" bold="true" italics="true" pageId="0" pageNumber="41">Legs</emphasis>
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robust and short, protibia somewhat curved inwards, mesotibia bent slightly outwards, metatibia straight. Male protarsomeres I-IV enlarged and with adhesive setae beneath.
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</paragraph>
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<paragraph id="C9FA2B94EFB2FA6A9990CCD0A53336C6" pageId="0" pageNumber="41">
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<emphasis id="1602627373E838179559043CE07C2F2B" bold="true" italics="true" pageId="0" pageNumber="41">Median lobe of aedeagus</emphasis>
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~ 0.75 mm long, strongly sclerotised. Dorsal side sinuous, apex pointed. Preputial field with three sclerites; the central one large, prolonged, at the tip spoon-like rounded, but central and basal part irregularly shaped; the left one rudimentary; the right one triangular (Fig.
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<figureCitation id="20BB8CC263C510A6A4E745C2874B8199" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Median lobe of aedeagus of Parazuphium a, c, d, f lateral views b, e, g view onto oroficium (preputial field) a, b P. weigeli sp. nov. c P. damascenum d, e P. salmoni f, g P. chevrolatii." figureDoi="10.3897/zookeys.1011.59449.figure3" httpUri="https://binary.pensoft.net/fig/497916" pageId="0" pageNumber="41">3a, b</figureCitation>
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). Left paramere rounded, larger than the strongly reduced right one.
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</paragraph>
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<caption id="7D75BC85B62B667806A00DDEBF5E2D57" doi="10.3897/zookeys.1011.59449.figure3" httpUri="https://binary.pensoft.net/fig/497916" pageId="0" pageNumber="41" start="Figure 3" startId="F3">
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<paragraph id="D42878B3275B33DFE6DF5BB14F2ABABE" pageId="0" pageNumber="41">
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<emphasis id="C4859CC1A35577309E1C5F5259BE0293" bold="true" pageId="0" pageNumber="41">Figure 3.</emphasis>
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Median lobe of aedeagus of
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<taxonomicName id="201365BA09D507BF1D3F2B2EBF0CE94E" authorityName="Jeannel" authorityYear="1942" class="Insecta" family="Carabidae" genus="Parazuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazuphium" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis id="E3368C08F64FA3588193278660EB4281" italics="true" pageId="0" pageNumber="41">Parazuphium</emphasis>
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</taxonomicName>
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<emphasis id="AED4967F05637618C1C461BBDB3DC0DB" bold="true" pageId="0" pageNumber="41">a, c, d, f</emphasis>
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lateral views
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<emphasis id="5E91F958C72C910D8794217F5CB3E41B" bold="true" pageId="0" pageNumber="41">b, e, g</emphasis>
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view onto oroficium (preputial field)
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<emphasis id="EB1BF1F74FD110876A5919A235BD8D3E" bold="true" pageId="0" pageNumber="41">a, b</emphasis>
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<taxonomicName id="42F82614C2C231EEC0BF80959BC7E21D" lsidName="P. weigeli" pageId="0" pageNumber="41" rank="species" species="weigeli">
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<emphasis id="4CDAFFF219354ACE04E247D6E2B7F650" italics="true" pageId="0" pageNumber="41">P. weigeli</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="D4C67F223097F18FFBE2CD90C1664169" bold="true" pageId="0" pageNumber="41">c</emphasis>
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<taxonomicName id="7D3EAA44BE83A688C828F24A5DD75FFA" lsidName="P. damascenum" pageId="0" pageNumber="41" rank="species" species="damascenum">
|
||
<emphasis id="C7E9FB97D1FCF9F2D25294A523325205" italics="true" pageId="0" pageNumber="41">P. damascenum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1D6F433D66D0C029E0888ADD5BD0B4FF" bold="true" pageId="0" pageNumber="41">d, e</emphasis>
|
||
<taxonomicName id="7F7B35F139BF37AF2DA8D74BB0B3CC00" lsidName="P. salmoni" pageId="0" pageNumber="41" rank="species" species="salmoni">
|
||
<emphasis id="4B11B8B9D77B1D1F0DAF610BCC2AE854" italics="true" pageId="0" pageNumber="41">P. salmoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F0DACD6EBDF5BC8269C83EA4C80B3BF1" bold="true" pageId="0" pageNumber="41">f, g</emphasis>
|
||
<taxonomicName id="8B75494B29CF1776BA4913A5BADDBF3C" lsidName="P. chevrolatii" pageId="0" pageNumber="41" rank="species" species="chevrolatii">
|
||
<emphasis id="256CD6C8BEA39CBCBED2956D506B79FE" italics="true" pageId="0" pageNumber="41">P. chevrolatii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="E14F887FC468BF27EBC3E333893FD352" pageId="0" pageNumber="41" type="comparisons">
|
||
<paragraph id="E1ECD29C81658322B1A650457B3F6B64" pageId="0" pageNumber="41">Comparisons.</paragraph>
|
||
<paragraph id="107793A5C2EB9BAF70E3D8435DBAC791" pageId="0" pageNumber="41">
|
||
The new species belongs to the genus
|
||
<taxonomicName id="E8D7F6E918DC50C7B1EDCCAC92917CC9" authorityName="Jeannel" authorityYear="1942" class="Insecta" family="Carabidae" genus="Parazuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazuphium" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="58A86759C88C73CBD67D8751EC5BFA6B" italics="true" pageId="0" pageNumber="41">Parazuphium</emphasis>
|
||
</taxonomicName>
|
||
because of small, numerous, erect setae on the scapus, one pair of supraorbital setae, a sinuous apical margin of the elytra, and a tube-shaped median lobe of the aedeagus with small sclerites on the preputial field.
|
||
</paragraph>
|
||
<paragraph id="955CE4324B211EC6E021B6BA2A15F818" pageId="0" pageNumber="41">
|
||
In its characters it is similar to
|
||
<taxonomicName id="C42FC6A2B0F9618E490170507A80535A" lsidName="P. salmoni" pageId="0" pageNumber="41" rank="species" species="salmoni">
|
||
<emphasis id="39172AC7F568E3C898CF0084736D78C4" italics="true" pageId="0" pageNumber="41">P. salmoni</emphasis>
|
||
</taxonomicName>
|
||
Assmann, Renan & Wrase, 2015. It shares with it the small eyes, strong and relatively short legs, moderately long antennae, metatibia straight in both sexes, similar body indices, brachyptery, and the colour. It differs from
|
||
<taxonomicName id="94C8351095BF1458CC585D25C6655C0F" lsidName="P. salmoni" pageId="0" pageNumber="41" rank="species" species="salmoni">
|
||
<emphasis id="6060BB685376E4DF9474F39406C77FBD" italics="true" pageId="0" pageNumber="41">P. salmoni</emphasis>
|
||
</taxonomicName>
|
||
by the following features: (1) eyes larger (eye diameter in dorsal view ~ 1/3 of temple length, in
|
||
<taxonomicName id="8925E168460B24BDE15FA9E0D84E94FE" lsidName="P. salmoni" pageId="0" pageNumber="41" rank="species" species="salmoni">
|
||
<emphasis id="79609F1512F9F0304CD21A30066F55F2" italics="true" pageId="0" pageNumber="41">P. salmoni</emphasis>
|
||
</taxonomicName>
|
||
~ 1/4); (2) antennomere I shorter in relation to antennomere II, ~ 5
|
||
<normalizedToken id="6A652EF5D26AC29CE1DE68E79416E70B" originalValue="×">x</normalizedToken>
|
||
as long as antennomere II (in
|
||
<taxonomicName id="FDA03EA90D73BDE6DAFE450599752738" lsidName="P. salmoni" pageId="0" pageNumber="41" rank="species" species="salmoni">
|
||
<emphasis id="E163F41E6940221F3B3B3D7655847869" italics="true" pageId="0" pageNumber="41">P. salmoni</emphasis>
|
||
</taxonomicName>
|
||
5.5-6
|
||
<normalizedToken id="399F61BF49CD81F95A4A1C8D10037FA1" originalValue="×">x</normalizedToken>
|
||
) (measured from the basal incision of the antennomeres to the apex); (3) pronotum more slender in relation to head (HW/PW 0.88-0.91, in
|
||
<taxonomicName id="CA372EA5D01F225BDA05FE0CE9BE79B9" lsidName="P. salmoni" pageId="0" pageNumber="41" rank="species" species="salmoni">
|
||
<emphasis id="6EADF609633B41E1341604D97C764940" italics="true" pageId="0" pageNumber="41">P. salmoni</emphasis>
|
||
</taxonomicName>
|
||
0.83-0.85); (4) hind angles less prominent (Fig.
|
||
<figureCitation id="44060B580D1AFA16A907DDFF703FFF5F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pronotum of Parazuphium A P. weigeli sp. nov. B P. salmoni." figureDoi="10.3897/zookeys.1011.59449.figure4" httpUri="https://binary.pensoft.net/fig/497917" pageId="0" pageNumber="41">4</figureCitation>
|
||
); (5) punctuation of pronotum less prominent (Fig.
|
||
<figureCitation id="3D564601CBA5A4DA6A94C782542EA55F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pronotum of Parazuphium A P. weigeli sp. nov. B P. salmoni." figureDoi="10.3897/zookeys.1011.59449.figure4" httpUri="https://binary.pensoft.net/fig/497917" pageId="0" pageNumber="41">4</figureCitation>
|
||
); (6) microsculpture of pronotum weaker and transverse (in
|
||
<taxonomicName id="568B4938CB361976799148FB5ABB83C3" lsidName="P. salmoni" pageId="0" pageNumber="41" rank="species" species="salmoni">
|
||
<emphasis id="CE102CE94ED4E863B67DCFF570C0A177" italics="true" pageId="0" pageNumber="41">P. salmoni</emphasis>
|
||
</taxonomicName>
|
||
isodiametric); (7) median lobe of aedeagus not downwardly bent and with different configuration and shape of the sclerites of the preputial field (Fig.
|
||
<figureCitation id="44B9B09BE828FFD69D8EE79C9A65B526" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Median lobe of aedeagus of Parazuphium a, c, d, f lateral views b, e, g view onto oroficium (preputial field) a, b P. weigeli sp. nov. c P. damascenum d, e P. salmoni f, g P. chevrolatii." figureDoi="10.3897/zookeys.1011.59449.figure3" httpUri="https://binary.pensoft.net/fig/497916" pageId="0" pageNumber="41">3a, b</figureCitation>
|
||
vs. Fig.
|
||
<figureCitation id="E1ED16BEAEC8378B208D6E06CB341B22" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Median lobe of aedeagus of Parazuphium a, c, d, f lateral views b, e, g view onto oroficium (preputial field) a, b P. weigeli sp. nov. c P. damascenum d, e P. salmoni f, g P. chevrolatii." figureDoi="10.3897/zookeys.1011.59449.figure3" httpUri="https://binary.pensoft.net/fig/497916" pageId="0" pageNumber="41">3d, e</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="3CBAF2905ACBABCBA4DB211B1918C397" doi="10.3897/zookeys.1011.59449.figure4" httpUri="https://binary.pensoft.net/fig/497917" pageId="0" pageNumber="41" start="Figure 4" startId="F4">
|
||
<paragraph id="20CBF296DE14F9B777ED797BDF568437" pageId="0" pageNumber="41">
|
||
<emphasis id="C62859FA00E77A0AAEA2691946008D5F" bold="true" pageId="0" pageNumber="41">Figure 4.</emphasis>
|
||
Pronotum of
|
||
<taxonomicName id="122F6239242FB28130DC7B146A5DE3F9" authorityName="Jeannel" authorityYear="1942" class="Insecta" family="Carabidae" genus="Parazuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazuphium" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F85EB858D03852A68D593B2EE692F5A9" italics="true" pageId="0" pageNumber="41">Parazuphium</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B57C4738C4A69170786F4DCD628D36DA" bold="true" pageId="0" pageNumber="41">A</emphasis>
|
||
<taxonomicName id="21A4FB31FA60AE9DB3278BB9F31FF179" lsidName="P. weigeli" pageId="0" pageNumber="41" rank="species" species="weigeli">
|
||
<emphasis id="B671CF0CB4FCD21CCF0248826F417744" italics="true" pageId="0" pageNumber="41">P. weigeli</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="FC9094183FAA5F84F72384203CD357E0" bold="true" pageId="0" pageNumber="41">B</emphasis>
|
||
<taxonomicName id="B2F85C58CE5C9AD64901BCCE2B8CE526" lsidName="P. salmoni" pageId="0" pageNumber="41" rank="species" species="salmoni">
|
||
<emphasis id="1C97843E5A140D369A1FF952C8B8FA02" italics="true" pageId="0" pageNumber="41">P. salmoni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="23337EA62291294BCC505447DB2612D8" pageId="0" pageNumber="41">
|
||
The new species differs from
|
||
<taxonomicName id="AD0D2CA87D3741D344410803C91FBF22" lsidName="P. damascenum" pageId="0" pageNumber="41" rank="species" species="damascenum">
|
||
<emphasis id="1F788468D342597AC13B2399C9B7A7BC" italics="true" pageId="0" pageNumber="41">P. damascenum</emphasis>
|
||
</taxonomicName>
|
||
(Fairmaire, 1897) by (1) straight metatibia in both genders; (2) smaller and flat eyes; (3) brachyptery, (4) deviating shape of median lobe of aedeagus and its sclerites (Fig.
|
||
<figureCitation id="7FB265FFC97D55C78D7A8813D8E60356" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Median lobe of aedeagus of Parazuphium a, c, d, f lateral views b, e, g view onto oroficium (preputial field) a, b P. weigeli sp. nov. c P. damascenum d, e P. salmoni f, g P. chevrolatii." figureDoi="10.3897/zookeys.1011.59449.figure3" httpUri="https://binary.pensoft.net/fig/497916" pageId="0" pageNumber="41">3a-c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F112577ECFD30E5DF045CF1930F01CAD" pageId="0" pageNumber="41">
|
||
<taxonomicName id="9606C6CC1CDABF6ED0AA02BAFD693B73" authorityName="Wrase & Assmann" authorityYear="2021" class="Insecta" family="Carabidae" genus="Parazuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazuphium weigeli" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="weigeli">
|
||
<emphasis id="485203F5507E791716E8C7FB6E61AC52" italics="true" pageId="0" pageNumber="41">Parazuphium weigeli</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from
|
||
<taxonomicName id="44291B8702DF9BB76F870E7A66CE8ED2" lsidName="P. chevrolatii" pageId="0" pageNumber="41" rank="species" species="chevrolatii">
|
||
<emphasis id="210BF14B94B1ADC5510A67246A7397C7" italics="true" pageId="0" pageNumber="41">P. chevrolatii</emphasis>
|
||
</taxonomicName>
|
||
by: (1) smaller eyes; (2) antennomere I shorter, shorter than head width; (3) weakly developed elytral intervals; (4) slender median lobe of aedeagus without an excision at the base of prolonged apical tip (Fig.
|
||
<figureCitation id="B4D7A278480BB291DAF8281F6CBA0376" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Median lobe of aedeagus of Parazuphium a, c, d, f lateral views b, e, g view onto oroficium (preputial field) a, b P. weigeli sp. nov. c P. damascenum d, e P. salmoni f, g P. chevrolatii." figureDoi="10.3897/zookeys.1011.59449.figure3" httpUri="https://binary.pensoft.net/fig/497916" pageId="0" pageNumber="41">3a, b</figureCitation>
|
||
vs. Fig.
|
||
<figureCitation id="1FDF361A58520E18DE5722E48D9E2C16" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Median lobe of aedeagus of Parazuphium a, c, d, f lateral views b, e, g view onto oroficium (preputial field) a, b P. weigeli sp. nov. c P. damascenum d, e P. salmoni f, g P. chevrolatii." figureDoi="10.3897/zookeys.1011.59449.figure3" httpUri="https://binary.pensoft.net/fig/497916" pageId="0" pageNumber="41">3f, g</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="34A5C05C9DC800FC5A862BD1ED6850C1" pageId="0" pageNumber="41">
|
||
The other microphthalmic species of the genus (
|
||
<taxonomicName id="725440CB864D3666EBD25CA6DA3598A5" lsidName="P. angustioculum" pageId="0" pageNumber="41" rank="species" species="angustioculum">
|
||
<emphasis id="0002D71A85BCD7B83130038139851D23" italics="true" pageId="0" pageNumber="41">P. angustioculum</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="CABACC6A8F12FB15BBD24838F0E6E478" originalValue="Hůrka">Hurka</normalizedToken>
|
||
, 1982,
|
||
<taxonomicName id="2CFD815B0FD39BB2A8A98AEFB5DDDBBA" lsidName="P. baeticum" pageId="0" pageNumber="41" rank="species" species="baeticum">
|
||
<emphasis id="E6C4E213C8CB908A1AFC5FFD22300111" italics="true" pageId="0" pageNumber="41">P. baeticum</emphasis>
|
||
</taxonomicName>
|
||
(K. & J. Daniel, 1898),
|
||
<taxonomicName id="7A7CD98A70E895B3CC70284CA3A40CAD" lsidName="P. punicum" pageId="0" pageNumber="41" rank="species" species="punicum">
|
||
<emphasis id="054C256E0CE7366493389950641178FA" italics="true" pageId="0" pageNumber="41">P. punicum</emphasis>
|
||
</taxonomicName>
|
||
(K. & J. Daniel, 1898),
|
||
<taxonomicName id="7CEA1C88DE25D6D18CDE4CD0F06C2876" lsidName="P. ramirezi" pageId="0" pageNumber="41" rank="species" species="ramirezi">
|
||
<emphasis id="4C9F28957E77504DA60FE598D639E154" italics="true" pageId="0" pageNumber="41">P. ramirezi</emphasis>
|
||
</taxonomicName>
|
||
J. & E. Vives, 1976) differ from
|
||
<taxonomicName id="9A2401D645C64FEB3C5EDFEF1F038B81" lsidName="P. weigeli" pageId="0" pageNumber="41" rank="species" species="weigeli">
|
||
<emphasis id="527030CB5F372758072B04D37AEB6F57" italics="true" pageId="0" pageNumber="41">P. weigeli</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by their (1) slender legs, (2) longer body lengths, (3) deviating proportions of antennomeres, (4) shape of median lobe of aedeagus and number and shape of sclerites (cf.
|
||
<bibRefCitation id="9E360A690356E6B7E55E1C44F45C002D" author="Hurka, K" journalOrPublisher="Acta Entomologica Bohemoslovaca" pageId="0" pageNumber="41" pagination="281 - 288" refId="B6" refString="Hurka, K, 1982. Taxonomic notes on Parazuphium, with descriptions of three new taxa (Coleoptera, Carabidae). Acta Entomologica Bohemoslovaca 79: 281 - 288" title="Taxonomic notes on Parazuphium, with descriptions of three new taxa (Coleoptera, Carabidae)." volume="79" year="1982">
|
||
<normalizedToken id="0A3EED8FD730EDF0170BEAB713C36C45" originalValue="Hůrka">Hurka</normalizedToken>
|
||
1982
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="A00C14C3160880A828F6BE217A1AB524" author="Hurka, K" journalOrPublisher="Acta Entomologica Bohemoslovaca" pageId="0" pageNumber="41" pagination="469 - 473" refId="B7" refString="Hurka, K, 1987. Separation of Parazuphium (P.) chevrolati and P. (P.) maroccanum, and description of a new subspecies (Col., Carabidae, Zuphiini). Acta Entomologica Bohemoslovaca 84: 469 - 473" title="Separation of Parazuphium (P.) chevrolati and P. (P.) maroccanum, and description of a new subspecies (Col., Carabidae, Zuphiini)." volume="84" year="1987">1987</bibRefCitation>
|
||
). These species occur exclusively in the south-western part of the Palaearctic (Morocco, Spain, and Sicily) (
|
||
<bibRefCitation id="1906061A3219CE6A0842ED264CDA8676" author="Huber, C" editor="Loebl, I" journalOrPublisher="Brill, Leiden, Boston" pageId="0" pageNumber="41" pagination="835 - 838" refId="B5" refString="Huber, C, Marggi, W, 2017. Tribe Zuphiini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Archostemata - Myxophaga - Adephaga)). Brill, Leiden, Boston: 835 - 838" title="Tribe Zuphiini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 1 (revised and updated edition, Archostemata - Myxophaga - Adephaga ))." year="2017">Huber and Marggi 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F1605BFBDD4D633EB848C005FF72FC1D" pageId="0" pageNumber="41" type="habitat">
|
||
<paragraph id="432A9B24638D2CC7DB50EAD1DE676921" pageId="0" pageNumber="41">Habitat.</paragraph>
|
||
<paragraph id="D5D88CF5388258CE383257842B41AC08" pageId="0" pageNumber="41">
|
||
Habitat is a subalpine grassland in 2500-2770 m a.s.l. The specimens were found under a large stone, deeply embedded in the soil, close to a snow field (Fig.
|
||
<figureCitation id="7FE819CA730C6D917414FCF6B5EC638F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitat of Parazuphium weigeli sp. nov. A landscape B microhabitat with the stone under which the two individuals were found. Photographs by Andreas Weigel." figureDoi="10.3897/zookeys.1011.59449.figure5" httpUri="https://binary.pensoft.net/fig/497918" pageId="0" pageNumber="41">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="475BBD4EC43C8D2C7D23D6698D49C8D7" doi="10.3897/zookeys.1011.59449.figure5" httpUri="https://binary.pensoft.net/fig/497918" pageId="0" pageNumber="41" start="Figure 5" startId="F5">
|
||
<paragraph id="F028E3F1E82DD3369EC85236D941ED64" pageId="0" pageNumber="41">
|
||
<emphasis id="7A588FE4C27D2A8EC00C477D56E383A4" bold="true" pageId="0" pageNumber="41">Figure 5.</emphasis>
|
||
Habitat of
|
||
<taxonomicName id="8CAE5B40DBB0BCABF90689E4B76D5A24" authorityName="Wrase & Assmann" authorityYear="2021" class="Insecta" family="Carabidae" genus="Parazuphium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parazuphium weigeli" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="weigeli">
|
||
<emphasis id="ABDE57C74C7904E0C21C4C6E59D22B60" italics="true" pageId="0" pageNumber="41">Parazuphium weigeli</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="77D4CF1C3A782DEDD4C77ACFD2EAF7FA" bold="true" pageId="0" pageNumber="41">A</emphasis>
|
||
landscape
|
||
<emphasis id="4B6997A51448A3FE08CAAB5EC5080E66" bold="true" pageId="0" pageNumber="41">B</emphasis>
|
||
microhabitat with the stone under which the two individuals were found. Photographs by Andreas Weigel.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="490108D7904F5929916A1EDF5FD543EA" pageId="0" pageNumber="41" type="dispersal power">
|
||
<paragraph id="D18BD91951BE14BD8434CDF860536A7A" pageId="0" pageNumber="41">Dispersal power.</paragraph>
|
||
<paragraph id="3BFE2EF28A66031DA10A8D3712A5C422" pageId="0" pageNumber="41">Brachypterous.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A90613D952D9002C51780AB18BD128FE" pageId="0" pageNumber="41" type="distribution">
|
||
<paragraph id="E605613C049476EF9C90E130006FF3FB" pageId="0" pageNumber="41">Distribution.</paragraph>
|
||
<paragraph id="3E881B0F7544309794E1665CAEFB5F94" pageId="0" pageNumber="41">Known so far only from the type locality. As the species is incapable of flight, its dispersal power is strongly limited, and a wider distribution range outside the Zagros Mountains is unlikely. Further studies may show if the occurrence of the species is restricted to high altitudes.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="326A915AFB7FDD331188637F31C22FFF" pageId="0" pageNumber="41" type="etymology">
|
||
<paragraph id="C63190901695589EF6757AC9D1A80477" pageId="0" pageNumber="41">Etymology.</paragraph>
|
||
<paragraph id="8A795EA16C46AE85EB529533A0376D9A" pageId="0" pageNumber="41">
|
||
Latinised patronym based on the surname of our colleague and friend, Andreas Weigel (
|
||
<normalizedToken id="F334A1B1B6F1631F29AE82734B2F38C3" originalValue="Pößneck">Poessneck</normalizedToken>
|
||
, Germany), specialist of
|
||
<taxonomicName id="86674127907FA88C23C0575FB4C3F30D" class="Insecta" family="Cerambycidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="family">Cerambycidae</taxonomicName>
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. He collected the first specimens of this new species and contributed to carabidology with numerous records of ground beetles, including new taxa.
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</paragraph>
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||
</subSubSection>
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</treatment>
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</document> |