152 lines
17 KiB
XML
152 lines
17 KiB
XML
<document id="61BC82FE976228A7091A2BB09D7EBE6C" ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZooBank="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="D7A7B0F1E44D2398243DE6DE1D29D698" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus productus Regimbart 1906" docType="treatment" docVersion="5" lastPageNumber="18" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="17" updateTime="1701314941532" updateUser="plazi">
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<mods:title id="663CE5FD12B900D29321C1EB6BC9D022">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="BFC0CB8ACF635C5250C7AE1D3A6DBFC7">Bistroem, Olof</mods:namePart>
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<mods:namePart id="5D74E2C71138472CC6301441252C579C">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="199FF810002566C271B47CC0E886C7FE">Bergsten, Johannes</mods:namePart>
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<mods:typeOfResource id="80C3CBC315CE7786FA4BFD0D67669417">text</mods:typeOfResource>
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<mods:date id="CECA1C60FF624789D255DFA19CF62491">2015</mods:date>
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<mods:detail id="4EDA038E0C1BE3FA276E762438AA8984" type="volume">
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<mods:number id="E34B44EE2832CD8AB7CC7881AC430167">542</mods:number>
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<mods:url id="4A4C3FA53853D7919D99D858855B346F">http://dx.doi.org/10.3897/zookeys.542.5975</mods:url>
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<mods:classification id="6E15D875AE5EE6101116A1D5123AC47B">journal article</mods:classification>
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<mods:identifier id="DD317C020DEEA0CD2EEF93E5695BDA1F" type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
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<treatment id="D7A7B0F1E44D2398243DE6DE1D29D698" ID-GBIF-Taxon="127862713" LSID="urn:lsid:plazi:treatment:D7A7B0F1E44D2398243DE6DE1D29D698" httpUri="http://treatment.plazi.org/id/D7A7B0F1E44D2398243DE6DE1D29D698" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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<subSubSection id="2652172482B4DA2DEBCBF3B925AB2FEE" pageId="16" pageNumber="17" type="multiple">
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<paragraph id="1214953F7DA026AA0552E5777D538D42" pageId="16" pageNumber="17">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="10260E5D520916210332E8C28EC388D2" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="28E434610563E24C8F35DAC4F9282FEC" pageId="16" pageNumber="17">
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<taxonomicName id="088DE6ED98772BAEFE748BC37ED5925E" ID-CoL="6NRNQ" authority="Regimbart, 1906" authorityName="Regimbart" authorityYear="1906" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus productus" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="productus">
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Laccophilus productus
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<normalizedToken id="02E0897E0BA57AFEB67C1F4F34AECC31" originalValue="Régimbart">Regimbart</normalizedToken>
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, 1906
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</taxonomicName>
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Figs 17-18, 219-221, 382, 527
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</paragraph>
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</subSubSection>
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<subSubSection id="3B468A9CE4E3016F49B9E80659FEEC37" pageId="17" pageNumber="18" type="reference_group">
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<paragraph id="C5B54471267EBE39D17ACBCA76676224" pageId="17" pageNumber="18">
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<taxonomicName id="DE3FC4E7E662132102EAEED3FC758BFB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus productus" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="productus">
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<pageBreakToken id="27A378280BA939FE79CA5FCEAB9A839E" pageId="17" pageNumber="18" start="start">Laccophilus</pageBreakToken>
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productus
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</taxonomicName>
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<bibRefCitation id="323F3526D9BF328017020D71C14C3DF8" author="Regimbart, M" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="262" pageNumber="263" pagination="235 - 278" title="Dytiscidae, Gyrinidae, Hydrophilidae. Voyage de M. Ch. Alluaud dans l'Afrique orientale." volume="75" year="1906">
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<normalizedToken id="CCB34D927FA84477C95CF69230EDAE32" originalValue="Régimbart">Regimbart</normalizedToken>
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1906
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</bibRefCitation>
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: 249 (original description, faunistics);
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<bibRefCitation id="783879E47343B8E20D85DCE9FC78A1C7" author="Zimmermann, A" journalOrPublisher="Junk, Berlin" pageId="263" pageNumber="264" title="Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Vol. 4 pars 71. W." year="1920 a">Zimmermann 1920a</bibRefCitation>
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: 25 (catalogue);
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<bibRefCitation id="5CC29C89A7A96126DC2CA1536969BE49" author="Peschet, R" journalOrPublisher="Imprimerie Generale Lahure, Paris" pageId="261" pageNumber="262" title="Dytiscidae et Gyrinidae. Voyage de M. G. Babault dans l'Afrique Orientale Anglaise" year="1921">Peschet 1921</bibRefCitation>
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: 6 (discussion, description, faunistics);
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<bibRefCitation id="C1B5C771905BDCF5CC21FE0AD8173AD8" author="Zimmermann, A" journalOrPublisher="Entomologische Blaetter" pageId="263" pageNumber="264" pagination="18 - 31" title="Schwimmkaefer-Material aus Deutsch-Ost-Afrika, gesammelt von Geh. Regierungsrat Methner in den Jahren 1902 - 1916." volume="22" year="1926">Zimmermann 1926</bibRefCitation>
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: 23 (faunistics);
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<bibRefCitation id="6FF150A4AE47D6DE57553D3112E807E1" author="Guignot, F" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle Paris" pageId="256" pageNumber="257" pagination="215 - 322" title="Mission scientifique de l'Omo 6 (58). ColeopteraDytiscidae (2 e partie)." volume="19" year="1946 c">Guignot 1946c</bibRefCitation>
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: 284, 313 (description, faunistics);
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<bibRefCitation id="119C88DEFDBD0B4B23DF3CF39C54FFB2" author="Guignot, F" journalOrPublisher="La Naturaliste Malgache" pageId="258" pageNumber="259" pagination="71 - 74" title="Un haliplide et deux dytiscides nouveaux de Madagascar." volume="9" year="1957 b">Guignot 1957b</bibRefCitation>
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: 73 (discussion, faunistics);
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<bibRefCitation id="69E794410D2FD271DB7368F575F3A2CB" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
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: 585, 586 (redescription, faunistics);
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<bibRefCitation id="577521CA73B9402F0BE934FCEDCCB48E" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
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: 249 (catalogue, faunistics);
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<bibRefCitation id="6E4B166944843EE67DDFCDC7AF03BD78" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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: 216 (catalogue, faunistics).
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</paragraph>
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</subSubSection>
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<subSubSection id="365680DB5BFC4A42AE68F51C92D7B7BE" pageId="17" pageNumber="18" type="type locality">
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<paragraph id="5B8B6B2EA3D3AEF558D23E0FEEF56B68" pageId="17" pageNumber="18">Type locality.</paragraph>
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<paragraph id="5C945F02AAE7746A3D801E735E203EDA" pageId="17" pageNumber="18">Kenya: Samburu.</paragraph>
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</subSubSection>
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<subSubSection id="8517B1752DEB556A92D98A88402A80D5" pageId="17" pageNumber="18" type="type material">
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<paragraph id="4088EBCA6DEF6CB6CD490FD46E4370E8" pageId="17" pageNumber="18">Type material studied</paragraph>
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<paragraph id="129BE5C171B77A93AB57A43FE09C8B5A" pageId="17" pageNumber="18">
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(5 exs.). Lectotype (by present designation): male: "Afrique Orle Anglaise Samburu (Wa-Nyika) Dr. Alluaud IV. 1904 / Museum Paris coll. Ch. Alluaud / TYPE /
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<taxonomicName id="C23F66A28EEDE84CB2CC9E4E425FDD85" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus productus" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="productus">Laccophilus productus</taxonomicName>
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<normalizedToken id="DE9B94A09B529A182124E017BB88A9F1" originalValue="Rég">Reg</normalizedToken>
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. sp. n. typ" (MNHN; top specimen on pin with two additional paralectotypes). - Paralectotypes: Similar data and on same pin as lectotype (2 exs. MNHN); "Samburu Wa-Nyika / Afr. Orle Angl. Alluaud / Museum Paris coll. Maurice
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<bibRefCitation id="DBBA0A45E6C5F937DF2E8C9F38C4F40D" author="Regimbart, M" journalOrPublisher="Almquist & Wiksells Boktryckeri-A. B., Uppsala" pageId="262" pageNumber="263" title="Coleoptera. 1. Dytiscidae, Gyrinidae, Hydrophilidae. In Sjoestedt, Y Wissenschaftliche Ergebnisse der schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und der umgebenden Massaisteppen Deutsch-Ostafrikas 1905 - 1906" year="1908">
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<normalizedToken id="BB658B9B79B049A50E42A7237D33BEAF" originalValue="Régimbart">Regimbart</normalizedToken>
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1908
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</bibRefCitation>
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/
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<taxonomicName id="3D28865CE76763D7F4A5643A645405D8" lsidName="productus" pageId="17" pageNumber="18" rank="species" species="productus">productus</taxonomicName>
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<normalizedToken id="69F7A0887822C3A7AE3F831260C78C94" originalValue="Rég.”">Reg."</normalizedToken>
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(2 exs. MNHN; habitus in Fig. 382).
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</paragraph>
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</subSubSection>
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<subSubSection id="3C9B532004D40ABDDC7EC7D6F283323E" pageId="17" pageNumber="18" type="materials_examined">
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<paragraph id="4682EA1BE7EF4C1834E20709FA3C9A5F" pageId="17" pageNumber="18">Additional material studied</paragraph>
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<paragraph id="6CC70B4945CDF5A5920C6CF3C18DF3E3" pageId="17" pageNumber="18">
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(3 exs.): Tanzania: "Kwakiyembe D.O.Afr. April 1916 Methner /
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<taxonomicName id="5EDCA5F13590B8E6206CE3AECFDE5797" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus productus" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="productus">Laccophilus productus</taxonomicName>
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<normalizedToken id="36F5F9C795D8BA6FEBFB1A97A2100A51" originalValue="Rég">Reg</normalizedToken>
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. det. Brancucci 1982" (1 ex. ZMHB); "Narobi b. Tanga 5. 1915 Methner" (1 ex. ZMHB); "Nord-Rabeho D.O. Afr. leg. Methner" (1 ex. ZMHB).
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</paragraph>
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</subSubSection>
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<subSubSection id="42991B2AA0B1202F2F72C3844E983FB6" pageId="17" pageNumber="18" type="diagnosis">
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<paragraph id="63CED71E1621C8BA63E0C69900C49D33" pageId="17" pageNumber="18">Diagnosis.</paragraph>
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<paragraph id="3752B3661FE9E201F7548E9A5F40E3E4" pageId="17" pageNumber="18">
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<taxonomicName id="59B5D0346F153566095772A720926421" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus productus" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="productus">Laccophilus productus</taxonomicName>
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is characterized by quite large but slender body and by peculiar dorsal colour pattern and male genitalia (penis apical half slightly twisted; extreme apex bent leftwards). The species resembles most of
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<taxonomicName id="B6FE6B4E7750654F53B16DC76BA44C56" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus morondavensis" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="morondavensis">Laccophilus morondavensis</taxonomicName>
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which occurs in Madagascar; diagnostic features are given under diagnosis of
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<taxonomicName id="8FB9F8FE6FFC1327314F6472119E6343" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus morondavensis" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="morondavensis">Laccophilus morondavensis</taxonomicName>
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on p. 22.
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</paragraph>
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</subSubSection>
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<subSubSection id="5EB08A10E6B2CCF5F5A4B9A51F8BE8FF" pageId="17" pageNumber="18" type="description">
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<paragraph id="045EE07D0F465AD80018CA5FFC8C8360" pageId="17" pageNumber="18">Description.</paragraph>
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<paragraph id="917E692B9DBDBEE6ED2C5668BD276D11" pageId="17" pageNumber="18">Body length 4.1-4.9, width 1.9-2.5 mm. Elytra with distinct colour pattern (Fig. 382); only minor variation exhibited.</paragraph>
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<paragraph id="37A539367ABD4CB3231B4A73C318F7DF" pageId="17" pageNumber="18">Head: Pale ferrugineous. Submat, finely and densely microsculptured. Reticulation simple; only with small, distinct meshes. Impunctate, except at eyes with scattered, fine, punctures.</paragraph>
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<paragraph id="C258E013F4EDAFF8C1AF3FFFF74934A0" pageId="17" pageNumber="18">Pronotum: Pale ferrugineous, mediobasally with a vague ferrugineous to dark ferrugineous marking. Submat, finely and densely microsculptured. Reticulation simple; only with small, distinct meshes. Impunctate, except at margins; with fine, somewhat sparse and irregular punctures. Mediobasally punctures absent or indistinct.</paragraph>
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<paragraph id="D3CCFA00342913B325B0AF7F0451BBB1" pageId="17" pageNumber="18">Elytra: Dark ferrugineous, with subbasal, preapical and apical, pale ferrugineous area (Fig. 382). Colour pattern stable and exhibits only minor variation. Submat, finely and densely microsculptured. Reticulation simple; only with small, distinct meshes. Fine, sparse and somewhat irregular punctures form a discal row. Dorsolateral and lateral rows indicated by scattered, fine punctures. Laterally with a comparatively long, finely pubescent, pre-apical furrow.</paragraph>
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<paragraph id="1A477F2A1A7F9D47636441BD653DD3E6" pageId="17" pageNumber="18">Ventral aspect: Dark ferrugineous to ferrugineous; colour pattern vague, indistinct. Rather shiny, finely microsculptured. Abdomen with fine, curved striae. Almost impunctate, except for apical ventrite; with scattered irregular punctures and shape symmetric (Fig. 17). Metacoxal plates with 13-15 almost transverse, fine, shallow furrows which in part are rather indistinct. Prosternal process slightly enlarged; apex moderately extended, pointed.</paragraph>
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<paragraph id="CB036485B00F2082A526756405B67CF5" pageId="17" pageNumber="18">Legs: Pro- and mesotarsus slightly enlarged, provided with suckers.</paragraph>
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<paragraph id="EFB2635676B631A49B100DD070762252" pageId="17" pageNumber="18">Male genitalia: Apical half of penis in dorsal aspect slightly sinuate; extreme apex slightly bent to left (Figs 219-221).</paragraph>
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<paragraph id="5390FC77428BFA5268B80002C915FB79" pageId="17" pageNumber="18">Female: Apical ventrite apically extended (Fig. 18). Pro- and mesotarsus slender, somewhat extended.</paragraph>
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</subSubSection>
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<subSubSection id="1D0D0C48743B767FBE01D2CF0E285EB8" pageId="17" pageNumber="18" type="distribution">
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<paragraph id="6107845AB229FE4B0804A74660B18B60" pageId="17" pageNumber="18">Distribution.</paragraph>
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<paragraph id="3C9B1FC77D7341D3E7075E414DE6B077" pageId="17" pageNumber="18">Kenya, Tanzania (Fig. 527).</paragraph>
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</subSubSection>
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<subSubSection id="4287274B3FBF1BF6568B9033585034D0" pageId="17" pageNumber="18" type="collecting circumstances">
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<paragraph id="492599FCAEDDE10F34704C263D3785A5" pageId="17" pageNumber="18">Collecting circumstances.</paragraph>
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<paragraph id="E5F24FCEA3470EA11427D2ECB361F09A" pageId="17" pageNumber="18">Unknown.</paragraph>
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</subSubSection>
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</treatment>
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