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<document id="1FCDB831F3149403C7AFC9BEECAE571C" ID-DOI="10.11646/zootaxa.5382.1.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="10279822" ID-ZooBank="E75EF166-6332-45F3-838B-0DF169BC0C46" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701937509306" checkinUser="plazi" docAuthor="Gil-Santana, Hélcio R." docDate="2023" docId="D72AF5687F2F173AFF6DFF2177C15D56" docLanguage="en" docName="zootaxa.5382.1.9.pdf" docOrigin="Zootaxa 5382 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5382.1.9/52425" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Brontostoma diringshofeni Gil-Santana &amp; Baena 2009" docType="treatment" docVersion="2" lastPageNumber="72" masterDocId="2B138D107F251730FFFAFFB6744D5E7F" masterDocTitle="A new species of Brontostoma Kirkaldy, description of the male of Brontostoma bahiensis Gil-Santana, Costa &amp; Marques and notes on Brontostoma doughertyae Gil-Santana et al. (Hemiptera: Heteroptera: Reduviidae: Ectrichodiinae)" masterLastPageNumber="86" masterPageNumber="62" pageNumber="72" updateTime="1701937952451" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0745F3B4C3E6E05947DDEF8CAFD94B5E">A new species of Brontostoma Kirkaldy, description of the male of Brontostoma bahiensis Gil-Santana, Costa &amp; Marques and notes on Brontostoma doughertyae Gil-Santana et al. (Hemiptera: Heteroptera: Reduviidae: Ectrichodiinae)</mods:title>
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<mods:date id="3FB1C55A87456D7085336668FD076C9A">2023</mods:date>
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<treatment id="D72AF5687F2F173AFF6DFF2177C15D56" ID-DOI="http://doi.org/10.5281/zenodo.10279836" ID-Zenodo-Dep="10279836" LSID="urn:lsid:plazi:treatment:D72AF5687F2F173AFF6DFF2177C15D56" httpUri="http://treatment.plazi.org/id/D72AF5687F2F173AFF6DFF2177C15D56" lastPageNumber="72" pageId="10" pageNumber="72">
<subSubSection id="179917F57F2F173AFF6DFF2177715ECE" box="[151,828,151,178]" pageId="10" pageNumber="72" type="nomenclature">
<paragraph id="5F3C447E7F2F173AFF6DFF2177715ECE" blockId="10.[151,828,151,214]" box="[151,828,151,178]" pageId="10" pageNumber="72">
<heading id="0474F3127F2F173AFF6DFF2177715ECE" bold="true" box="[151,828,151,178]" fontSize="11" level="1" pageId="10" pageNumber="72" reason="1">
<taxonomicName id="98833FFD7F2F173AFF6DFF2177715ECE" ID-CoL="5WR6M" authority="Gil-Santana &amp; Baena, 2009" authorityName="Gil-Santana &amp; Baena" authorityYear="2009" box="[151,828,151,178]" class="Insecta" family="Reduviidae" genus="Brontostoma" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="72" phylum="Arthropoda" rank="species" species="diringshofeni">
<emphasis id="6DF7986C7F2F173AFF6DFF2177715ECE" bold="true" box="[151,828,151,178]" pageId="10" pageNumber="72">
<emphasis id="6DF7986C7F2F173AFF6DFF2175935ECD" bold="true" box="[151,478,151,178]" italics="true" pageId="10" pageNumber="72">Brontostoma diringshofeni</emphasis>
<bibRefCitation id="3B12398F7F2F173AFE1FFF2177715ECE" author="Gil-Santana, H. R. &amp; Baena, M." box="[485,828,151,177]" pageId="10" pageNumber="72" pagination="41 - 52" refId="ref14524" refString="Gil-Santana, H. R. &amp; Baena, M. (2009) Two new species of Brontostoma Kirkaldy (Hemiptera: Heteroptera: Reduviidae: Ectrichodiinae) from Bolivia, with description of the male genitalia of two other species of the genus, and description of the female of B. doughertyae Gil-Santana, Lopes, Marques &amp; Jurberg. Zootaxa, 1979 (1), 41 - 52. https: // doi. org / 10.11646 / zootaxa. 1979.1. 4" type="journal article" year="2009">Gil-Santana &amp; Baena, 2009</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="179917F57F2F173AFF6DFF0D75635EA9" box="[151,302,187,214]" pageId="10" pageNumber="72" type="description">
<paragraph id="5F3C447E7F2F173AFF6DFF0D75635EA9" blockId="10.[151,828,151,214]" box="[151,302,187,214]" pageId="10" pageNumber="72">
(
<figureCitation id="C7B858FB7F2F173AFF65FF0D756A5EA9" box="[159,295,187,214]" captionStart="FIGURES 24–27" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-21@12.[165,1422,181,1919]" captionTargetPageId="12" captionText="FIGURES 24–27. 24–25, Brontostoma diringshofeni Gil-Santana &amp; Baena, 2009, male holotype deposited in MZUSP, 24, dorsal view, scale bar 5.0 mm; 25, labels; 26–27, Brontostoma doughertyae Gil-Santana, Lopes, Marques &amp; Jurberg, 2005, males, dorsal view, scale bars 5.0 mm, 26, holotype, previously deposited in MNRJ, 27, non-type specimen." figureDoi="http://doi.org/10.5281/zenodo.10279838" httpUri="https://zenodo.org/record/10279838/files/figure.png" pageId="10" pageNumber="72">Figs. 24–25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="179917F57F2F173AFF6DFEB6764C5F2A" pageId="10" pageNumber="72" type="reference_group">
<paragraph id="5F3C447E7F2F173AFF6DFEB6764C5F2A" blockId="10.[151,1437,255,341]" pageId="10" pageNumber="72">
<treatmentCitationGroup id="7F9363507F2F173AFF6DFEB6764C5F2A" pageId="10" pageNumber="72">
<treatmentCitation id="DE22626F7F2F173AFF6DFEB6776F5F68" ID-DOI="http://doi.org/10.5281/zenodo.6226714" ID-Zenodo-Dep="6226714" author="Gil-Santana, H. R. &amp; Baena, M." box="[151,802,255,280]" httpUri="http://treatment.plazi.org/id/0748AB34FFB0FFA9FF113CB3FC9BECEB" page=")" pageId="10" pageNumber="72" year="2009">
<taxonomicName id="98833FFD7F2F173AFF6DFEB6776F5F68" ID-CoL="5WR6M" authority="Gil-Santana &amp; Baena (2009: 46 - 48)" authorityName="Gil-Santana &amp; Baena" authorityPageNumber="46 - 48" authorityYear="2009" box="[151,802,255,280]" class="Insecta" family="Reduviidae" genus="Brontostoma" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="72" phylum="Arthropoda" rank="species" species="diringshofeni">
<emphasis id="6DF7986C7F2F173AFF6DFEB675EA5F67" box="[151,423,256,280]" italics="true" pageId="10" pageNumber="72">Brontostoma diringshofeni</emphasis>
<bibRefCitation id="3B12398F7F2F173AFE55FEB6776F5F68" author="Gil-Santana, H. R. &amp; Baena, M." box="[431,802,255,280]" pageId="10" pageNumber="72" pagination="41 - 52" refId="ref14524" refString="Gil-Santana, H. R. &amp; Baena, M. (2009) Two new species of Brontostoma Kirkaldy (Hemiptera: Heteroptera: Reduviidae: Ectrichodiinae) from Bolivia, with description of the male genitalia of two other species of the genus, and description of the female of B. doughertyae Gil-Santana, Lopes, Marques &amp; Jurberg. Zootaxa, 1979 (1), 41 - 52. https: // doi. org / 10.11646 / zootaxa. 1979.1. 4" type="journal article" year="2009">Gil-Santana &amp; Baena (2009: 46–48)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(description, figures 28–46), (50) (diagnostic characteristics);
<treatmentCitation id="DE22626F7F2F173AFF3DFEA8765A5F49" ID-DOI="http://doi.org/10.5281/zenodo.6160537" ID-Zenodo-Dep="6160537" author="Gil-Santana, H. R. &amp; Baena, M. &amp; Grillo, H." box="[199,535,286,310]" httpUri="http://treatment.plazi.org/id/DA46878AFFFA904BC3D1F8E6692EFEC5" page="62" pageId="10" pageNumber="72" year="2013">
<bibRefCitation id="3B12398F7F2F173AFF3DFEA8765A5F49" author="Gil-Santana, H. R. &amp; Baena, M. &amp; Grillo, H." box="[199,535,286,310]" pageId="10" pageNumber="72" pagination="60 - 74" refId="ref14612" refString="Gil-Santana, H. R., Baena, M. &amp; Grillo, H. (2013 a) Berengeria Gil-Santana &amp; Coletto-Silva, a junior synonym of Ectrichodiella Fracker &amp; Bruner, with new records and taxonomic notes on Ectrichodiinae from Brazil, and with keys to Ectrichodiinae and Reduviinae genera of the New World (Hemiptera: Heteroptera: Reduviidae). Zootaxa, 3652 (1), 60 - 74. https: // doi. org / 10.11646 / zootaxa. 3652.1. 2" type="journal article" year="2013">
Gil-Santana
<emphasis id="6DF7986C7F2F173AFEBCFEA9753C5F49" box="[326,369,286,310]" italics="true" pageId="10" pageNumber="72">et al</emphasis>
. (2013a: 62–63)
</bibRefCitation>
</treatmentCitation>
(record from
<collectingCountry id="279404EE7F2F173AFD5BFEA876AC5F49" box="[673,737,286,310]" name="Brazil" pageId="10" pageNumber="72">Brazil</collectingCountry>
, figure 3);
<treatmentCitation id="DE22626F7F2F173AFCB4FEA8701C5F49" author="Ferreira, V. S. &amp; Prado, L. P. &amp; Seripierri, D." box="[846,1105,286,310]" page="122" pageId="10" pageNumber="72" year="2016">
<bibRefCitation id="3B12398F7F2F173AFCB4FEA8701C5F49" author="Ferreira, V. S. &amp; Prado, L. P. &amp; Seripierri, D." box="[846,1105,286,310]" pageId="10" pageNumber="72" pagination="117 - 122" refId="ref13887" refString="Ferreira, V. S., Prado, L. P. &amp; Seripierri, D. (2016) The Entomological Collection of Ricardo von Diringshofen (1900 - 1986) and its incorporation to the Museu de Zoologia da Universidade de Sao Paulo. Revista Brasileira de Entomologia, 60, 117 - 122. https: // doi. org / 10.1016 / j. rbe. 2016.01.005" type="journal article" year="2016">
Ferreira
<emphasis id="6DF7986C7F2F173AFC5EFEA9779D5F49" box="[932,976,286,310]" italics="true" pageId="10" pageNumber="72">et al</emphasis>
. (2016: 122)
</bibRefCitation>
</treatmentCitation>
(citation);
<treatmentCitation id="DE22626F7F2F173AFB45FEA874905F2A" author="Carrenho, R. &amp; Rodrigues, H. D. D. &amp; Lima, A. C. &amp; Schwertner, C. F." page="4" pageId="10" pageNumber="72" year="2020">
<bibRefCitation id="3B12398F7F2F173AFB45FEA874905F2A" author="Carrenho, R. &amp; Rodrigues, H. D. D. &amp; Lima, A. C. &amp; Schwertner, C. F." pageId="10" pageNumber="72" pagination="20206017" refId="ref13735" refString="Carrenho, R., Rodrigues, H. D. D., Lima, A. C. &amp; Schwertner, C. F. (2020) Type specimens of true bugs (Hemiptera: Heteroptera) housed in the Museu de Zoologia da Universidade de Sao Paulo, Brazil. Papeis avulsos de Zoologia, 60, e 20206017. https: // doi. org / 10.11606 / 1807 - 0205 / 2020.60.17" type="journal article" year="2020">
Carrenho
<emphasis id="6DF7986C7F2F173AFAD9FEA971025F49" box="[1315,1359,286,310]" italics="true" pageId="10" pageNumber="72">et al</emphasis>
. (2020: 4)
</bibRefCitation>
</treatmentCitation>
(catalog of
<typeStatus id="8038FADC7F2F173AFEACFE8B75CF5F2A" box="[342,386,317,341]" pageId="10" pageNumber="72">type</typeStatus>
specimens).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="179917F57F2F173AFF6DFE3577C15D56" pageId="10" pageNumber="72" type="distribution">
<paragraph id="5F3C447E7F2F173AFF6DFE3577075FE2" blockId="10.[151,1437,387,810]" box="[151,842,387,414]" pageId="10" pageNumber="72">
<emphasis id="6DF7986C7F2F173AFF6DFE35757C5FE2" bold="true" box="[151,305,387,413]" pageId="10" pageNumber="72">Distribution.</emphasis>
<collectingCountry id="279404EE7F2F173AFECDFE3575C45FE2" box="[311,393,387,413]" name="Bolivia" pageId="10" pageNumber="72">Bolivia</collectingCountry>
and
<collectingCountry id="279404EE7F2F173AFE3AFE35764A5FE2" box="[448,519,387,413]" name="Brazil" pageId="10" pageNumber="72">Brazil</collectingCountry>
, in the state of
<collectingRegion id="9D478A9C7F2F173AFD4BFE32770B5FE2" box="[689,838,387,414]" country="Brazil" name="Mato Grosso" pageId="10" pageNumber="72">Mato Grosso</collectingRegion>
.
</paragraph>
<paragraph id="5F3C447E7F2F173AFF3DFE1177C15D56" blockId="10.[151,1437,387,810]" pageId="10" pageNumber="72">
The description of
<taxonomicName id="98833FFD7F2F173AFE61FE1E768B5FBE" authorityName="Gil-Santana &amp; Baena" authorityYear="2009" box="[411,710,423,449]" class="Insecta" family="Reduviidae" genus="Brontostoma" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="72" phylum="Arthropoda" rank="species" species="diringshofeni">
<emphasis id="6DF7986C7F2F173AFE61FE1E768B5FBE" box="[411,710,423,449]" italics="true" pageId="10" pageNumber="72">Brontostoma diringshofeni</emphasis>
</taxonomicName>
, based on a single male from
<collectingCountry id="279404EE7F2F173AFBEFFE11702A5FBE" box="[1045,1127,423,449]" name="Bolivia" pageId="10" pageNumber="72">Bolivia</collectingCountry>
(
<figureCitation id="C7B858FB7F2F173AFB8DFE11714D5FBD" box="[1143,1280,423,450]" captionStart="FIGURES 24–27" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-21@12.[165,1422,181,1919]" captionTargetPageId="12" captionText="FIGURES 24–27. 24–25, Brontostoma diringshofeni Gil-Santana &amp; Baena, 2009, male holotype deposited in MZUSP, 24, dorsal view, scale bar 5.0 mm; 25, labels; 26–27, Brontostoma doughertyae Gil-Santana, Lopes, Marques &amp; Jurberg, 2005, males, dorsal view, scale bars 5.0 mm, 26, holotype, previously deposited in MNRJ, 27, non-type specimen." figureDoi="http://doi.org/10.5281/zenodo.10279838" httpUri="https://zenodo.org/record/10279838/files/figure.png" pageId="10" pageNumber="72">Figs. 24–25</figureCitation>
), included the description and figures of the male genitalia (
<bibRefCitation id="3B12398F7F2F173AFD62FE7D779D5F9A" author="Gil-Santana, H. R. &amp; Baena, M." box="[664,976,459,486]" pageId="10" pageNumber="72" pagination="41 - 52" refId="ref14524" refString="Gil-Santana, H. R. &amp; Baena, M. (2009) Two new species of Brontostoma Kirkaldy (Hemiptera: Heteroptera: Reduviidae: Ectrichodiinae) from Bolivia, with description of the male genitalia of two other species of the genus, and description of the female of B. doughertyae Gil-Santana, Lopes, Marques &amp; Jurberg. Zootaxa, 1979 (1), 41 - 52. https: // doi. org / 10.11646 / zootaxa. 1979.1. 4" type="journal article" year="2009">Gil-Santana &amp; Baena 2009</bibRefCitation>
). This species was further recorded from
<collectingCountry id="279404EE7F2F173AFF6DFE5974965C76" box="[151,219,495,521]" name="Brazil" pageId="10" pageNumber="72">Brazil</collectingCountry>
by
<bibRefCitation id="3B12398F7F2F173AFEFCFE5976645C76" author="Gil-Santana, H. R. &amp; Baena, M. &amp; Grillo, H." box="[262,553,495,522]" pageId="10" pageNumber="72" pagination="60 - 74" refId="ref14612" refString="Gil-Santana, H. R., Baena, M. &amp; Grillo, H. (2013 a) Berengeria Gil-Santana &amp; Coletto-Silva, a junior synonym of Ectrichodiella Fracker &amp; Bruner, with new records and taxonomic notes on Ectrichodiinae from Brazil, and with keys to Ectrichodiinae and Reduviinae genera of the New World (Hemiptera: Heteroptera: Reduviidae). Zootaxa, 3652 (1), 60 - 74. https: // doi. org / 10.11646 / zootaxa. 3652.1. 2" type="journal article" year="2013">
Gil-Santana
<emphasis id="6DF7986C7F2F173AFE69FE4675895C76" box="[403,452,495,521]" italics="true" pageId="10" pageNumber="72">et al</emphasis>
. (2013a)
</bibRefCitation>
based on a single male too. Although
<bibRefCitation id="3B12398F7F2F173AFC2CFE5970B75C76" author="Gil-Santana, H. R. &amp; Baena, M. &amp; Grillo, H." box="[982,1274,495,522]" pageId="10" pageNumber="72" pagination="60 - 74" refId="ref14612" refString="Gil-Santana, H. R., Baena, M. &amp; Grillo, H. (2013 a) Berengeria Gil-Santana &amp; Coletto-Silva, a junior synonym of Ectrichodiella Fracker &amp; Bruner, with new records and taxonomic notes on Ectrichodiinae from Brazil, and with keys to Ectrichodiinae and Reduviinae genera of the New World (Hemiptera: Heteroptera: Reduviidae). Zootaxa, 3652 (1), 60 - 74. https: // doi. org / 10.11646 / zootaxa. 3652.1. 2" type="journal article" year="2013">
Gil-Santana
<emphasis id="6DF7986C7F2F173AFB99FE4670D95C76" box="[1123,1172,495,521]" italics="true" pageId="10" pageNumber="72">et al</emphasis>
. (2013a)
</bibRefCitation>
stated that the specimen examined by them was very similar to the
<typeStatus id="8038FADC7F2F173AFD0DFDA577115C52" box="[759,860,531,557]" pageId="10" pageNumber="72" type="holotype">holotype</typeStatus>
, they did not record that it showed a faint reddish tinge at basolateral portions of connexivum, somewhat more pronounced on the first segments (see their fig. 3), a characteristic not evident in the
<typeStatus id="8038FADC7F2F173AFE07FDED76125C0A" box="[509,607,603,629]" pageId="10" pageNumber="72" type="holotype">holotype</typeStatus>
(
<figureCitation id="C7B858FB7F2F173AFD95FDED768E5C0A" box="[623,707,603,629]" captionStart="FIGURES 24–27" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[165,1422,181,1919]" captionTargetId="figure-21@12.[165,1422,181,1919]" captionTargetPageId="12" captionText="FIGURES 24–27. 24–25, Brontostoma diringshofeni Gil-Santana &amp; Baena, 2009, male holotype deposited in MZUSP, 24, dorsal view, scale bar 5.0 mm; 25, labels; 26–27, Brontostoma doughertyae Gil-Santana, Lopes, Marques &amp; Jurberg, 2005, males, dorsal view, scale bars 5.0 mm, 26, holotype, previously deposited in MNRJ, 27, non-type specimen." figureDoi="http://doi.org/10.5281/zenodo.10279838" httpUri="https://zenodo.org/record/10279838/files/figure.png" pageId="10" pageNumber="72">Fig. 24</figureCitation>
). Nevertheless, this small difference is considered here as a color variation of the species, as commonly observed in ectrichodiines, including species of
<taxonomicName id="98833FFD7F2F173AFB98FD3670BF5CE6" authorityName="Kirkaldy" authorityYear="1904" box="[1122,1266,640,665]" class="Insecta" family="Reduviidae" genus="Brontostoma" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="6DF7986C7F2F173AFB98FD3670BF5CE6" box="[1122,1266,640,665]" italics="true" pageId="10" pageNumber="72">Brontostoma</emphasis>
</taxonomicName>
(
<bibRefCitation id="3B12398F7F2F173AFAFAFDC9749F5CC1" author="Wygodzinsky, P." pageId="10" pageNumber="72" pagination="35 - 52" refId="ref15689" refString="Wygodzinsky, P. (1951) Notas sobre Ectrichodiinae Neotropicales (Hemipt. Reduviidae). Revista de la Sociedad Entomologica Argentina, 15, 35 - 52." type="journal article" year="1951">Wygodzinsky 1951</bibRefCitation>
;
<bibRefCitation id="3B12398F7F2F173AFF18FD1575F35CC2" author="Gil-Santana, H. R." box="[226,446,675,701]" pageId="10" pageNumber="72" pagination="301 - 325" refId="ref14475" refString="Gil-Santana, H. R. (2020 b) New synonymies among species of Brontostoma (Hemiptera: Heteroptera: Reduviidae). Zootaxa, 4869 (3), 301 - 325. https: // doi. org / 10.11646 / zootaxa. 4869.3. 1" type="journal article" year="2020">Gil-Santana 2020b</bibRefCitation>
). Alternatively, taking into account that the specimen recorded by
<bibRefCitation id="3B12398F7F2F173AFB35FD1574A35C9E" author="Gil-Santana, H. R. &amp; Baena, M. &amp; Grillo, H." pageId="10" pageNumber="72" pagination="60 - 74" refId="ref14612" refString="Gil-Santana, H. R., Baena, M. &amp; Grillo, H. (2013 a) Berengeria Gil-Santana &amp; Coletto-Silva, a junior synonym of Ectrichodiella Fracker &amp; Bruner, with new records and taxonomic notes on Ectrichodiinae from Brazil, and with keys to Ectrichodiinae and Reduviinae genera of the New World (Hemiptera: Heteroptera: Reduviidae). Zootaxa, 3652 (1), 60 - 74. https: // doi. org / 10.11646 / zootaxa. 3652.1. 2" type="journal article" year="2013">
Gil-Santana
<emphasis id="6DF7986C7F2F173AFA9AFD1271D85CC2" box="[1376,1429,675,701]" italics="true" pageId="10" pageNumber="72">et al</emphasis>
. (2013a)
</bibRefCitation>
was collected more than 30 years after the
<typeStatus id="8038FADC7F2F173AFD29FD7177755C9E" box="[723,824,711,737]" pageId="10" pageNumber="72" type="holotype">holotype</typeStatus>
, it is possible that the
<typeStatus id="8038FADC7F2F173AFBD5FD7170DC5C9E" box="[1071,1169,711,737]" pageId="10" pageNumber="72" type="holotype">holotype</typeStatus>
was partially faded and had lost the reddish tinge of the connexivum. In order to clarify the actual significance of this variation, it would be necessary to examine more specimens, preferably freshly collected.
</paragraph>
</subSubSection>
</treatment>
</document>
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