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<document ID-CLB-Dataset="9370" ID-DOI="http://dx.doi.org/10.3897/phytokeys.169.57532" ID-GBIF-Dataset="353d0a27-32f0-4015-86f7-b60eed07bceb" ID-PMC="PMC7738470" ID-Pensoft-Pub="1314-2003-169-99" ID-Pensoft-UUID="1963D0A39F2B591E8BFDC11579CC4323" ID-PubMed="33354141" ModsDocID="1314-2003-169-99" checkinTime="1607471176246" checkinUser="pensoft" docAuthor="Lavandero, Nicolas, Rosende, Benito &amp; Perez, Maria Fernanda" docDate="2020" docId="C2CD131A1BBA53028CC7F0723D7790DF" docLanguage="en" docName="PhytoKeys 169: 99-117" docOrigin="PhytoKeys 169" docPubDate="2020-12-08" docSource="http://dx.doi.org/10.3897/phytokeys.169.57532" docTitle="Leucheria cantillanensis Lavandero 2020, sp. nov." docType="treatment" docVersion="6" id="1963D0A39F2B591E8BFDC11579CC4323" lastPageNumber="99" masterDocId="1963D0A39F2B591E8BFDC11579CC4323" masterDocTitle="Leucheria cantillanensis (Nassauvieae, Asteraceae), a new species endemic to Central Chile" masterLastPageNumber="117" masterPageNumber="99" pageNumber="99" updateTime="1732856593099" updateUser="ExternalLinkService">
<mods:mods id="4311FF0A60781838F1BB20CDD4F810E2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3D9EECDB3C1A2143907DDEDD8FB7F61C">
<mods:title id="5C5E0336CE46EAB3D4E14BF6E7C1B7E5">Leucheria cantillanensis (Nassauvieae, Asteraceae), a new species endemic to Central Chile</mods:title>
</mods:titleInfo>
<mods:name id="CFD65227034F08FE451AF5AE2E66817B" type="personal">
<mods:role id="3F5C3CCF67B8CA4C14BD98E08FBFF773">
<mods:roleTerm id="AC51771B4BEF99722B941E54B3070574">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2A2D1B5366C84DB90253696AF3C15DB8">Lavandero, Nicolas</mods:namePart>
<mods:nameIdentifier id="CD8E92C5314D1CBF687CDEBD543B1B5D" type="ORCID">https://orcid.org/0000-0002-2390-5078</mods:nameIdentifier>
<mods:affiliation id="88BE6C11449375139F149BEEFC5A5DA1">Departamento de Ecologia, Facultad de Ciencias Biologicas, Pontificia Universidad Catolica de Chile, Avenida Libertador B. O'Higgins 340, Santiago, Chile &amp; Instituto de Ecologia y Biodiversidad (IEB), Universidad de Chile, Las Palmeras 3425, Santiago, Chile</mods:affiliation>
<mods:nameIdentifier id="486140E298A2816853E994F5D1CC7FAC" type="email">nglavand@uc.cl</mods:nameIdentifier>
</mods:name>
<mods:name id="873E7C27A1AC7570F1872EAC7904C578" type="personal">
<mods:role id="BC522909EF120EBF68247975D8A1F3F7">
<mods:roleTerm id="7AA3F2311E08E5550C2AE2300B234F91">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D6812021A7C8409A670D0AE08510E98C">Rosende, Benito</mods:namePart>
<mods:affiliation id="A82AB4B1519B26FADFA67FAF17EAFA0B">Departamento de Ecologia, Facultad de Ciencias Biologicas, Pontificia Universidad Catolica de Chile, Avenida Libertador B. O'Higgins 340, Santiago, Chile</mods:affiliation>
</mods:name>
<mods:name id="9006B8F7CB474766EF4B283A6499419F" type="personal">
<mods:role id="34E17221568C2D64313CA84F422A12A9">
<mods:roleTerm id="10E2791328CFE986FA405E899804B006">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="462F9FC59615F9524E78526C2CFB23B8">Perez, Maria Fernanda</mods:namePart>
<mods:affiliation id="606626BEFAF0C310FBBF2EF7F6DC8D90">Departamento de Ecologia, Facultad de Ciencias Biologicas, Pontificia Universidad Catolica de Chile, Avenida Libertador B. O'Higgins 340, Santiago, Chile &amp; Instituto de Ecologia y Biodiversidad (IEB), Universidad de Chile, Las Palmeras 3425, Santiago, Chile</mods:affiliation>
</mods:name>
<mods:typeOfResource id="9C690427C9066AFC572C59CC88552F0D">text</mods:typeOfResource>
<mods:relatedItem id="4C29E540104120D09C85525E555E26EB" type="host">
<mods:titleInfo id="306122505CEA45A836AEC7CAA884E41C">
<mods:title id="2E791448C930F3AD487A3F4C4DB9D523">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="9CFE1AACAED48216A555B8712984BE16">
<mods:date id="7D5B27917C08995A09B177792DBF6CAC">2020</mods:date>
<mods:detail id="5DDAD6A9CCC64CB8CB3E018D5F3E3729" type="pubDate">
<mods:number id="4B03EBBB1009775A693261B4E9F8D00B">2020-12-08</mods:number>
</mods:detail>
<mods:detail id="2BE19404AECC4CC75D5700B393AE7DCB" type="volume">
<mods:number id="20D6ED51E0B1A5604E81B5423713B30D">169</mods:number>
</mods:detail>
<mods:extent id="00F124D418AB11254DAB353A53EB591C" unit="page">
<mods:start id="DDF3F70B3A1F23F916EEBBB02AC0DB8F">99</mods:start>
<mods:end id="8B86793A953FC01967F18B234D0C295F">117</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="823CDC3EFFA75C915DA7EE1D04F16CE4">
<mods:url id="1B6547D33C7866D673607C0379FD317C">http://dx.doi.org/10.3897/phytokeys.169.57532</mods:url>
</mods:location>
<mods:classification id="0EDE04824D7AE7CB2D91B2F202750AC3">journal article</mods:classification>
<mods:identifier id="5048E9D534FC85E4B2CC6AD2A33096A8" type="DOI">http://dx.doi.org/10.3897/phytokeys.169.57532</mods:identifier>
<mods:identifier id="E5696D8E267FD72A95DF41145A02E0D2" type="Pensoft-Pub">1314-2003-169-99</mods:identifier>
<mods:identifier id="1396BFE9226AB74CDBFDDFB67134C30E" type="Pensoft-UUID">1963D0A39F2B591E8BFDC11579CC4323</mods:identifier>
</mods:mods>
<treatment id="C2CD131A1BBA53028CC7F0723D7790DF" ID-GBIF-Taxon="170719407" LSID="urn:lsid:plazi:treatment:C2CD131A1BBA53028CC7F0723D7790DF" httpUri="http://treatment.plazi.org/id/C2CD131A1BBA53028CC7F0723D7790DF" lastPageNumber="99" pageId="0" pageNumber="99" scope_family="Asteraceae">
<subSubSection id="43D3778251783BAE615BB47F0EB6BD87" pageId="0" pageNumber="99" type="nomenclature">
<paragraph id="8888C763FF2E5DA668BB09166F1D003F" pageId="0" pageNumber="99">
<taxonomicName id="923216EC72C3B5D5516E5ADB1DB3F02A" ID-CoL="849PF" LSID="C2CD131A-1BBA-5302-8CC7-F0723D7790DF" authority="Lavandero" authorityName="Lavandero" authorityYear="2020" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria cantillanensis" order="Asterales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="cantillanensis" status="sp. nov.">Leucheria cantillanensis Lavandero</taxonomicName>
<taxonomicNameLabel id="F1684A3CF87D1CD6079E90DBD1E0C63A" pageId="0" pageNumber="99">sp. nov.</taxonomicNameLabel>
<figureCitation id="1B6F60F9422BBD33247E0274AEE0F8B8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leucheria cantillanensis Lavandero, sp. nov. A capitulum, sideways view B capitulum, detail of Middle involucral bracts C leaves D capitulum, upper view E plants growing in natural habitat F cypsela G detail of trichomes in cypsela. All photographs by Nicolas Lavandero." figureDoi="10.3897/phytokeys.169.57532.figure3" httpUri="https://binary.pensoft.net/fig/484182" pageId="0" pageNumber="99">Figures 3</figureCitation>
<figureCitation id="D9E760300414F7653EF765952F64249B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Leucheria cantillanensis Lavandero, sp. nov. A habit B capitulum C detail of flower D fruit E basal leaf F leaf trichome G stem trichome H inner Middle involucral bract I middle Middle involucral bract J outer Middle involucral bract K ciliate margin of inner Middle involucral bract (detail). Illustrations by Benito Rosende." figureDoi="10.3897/phytokeys.169.57532.figure4" httpUri="https://binary.pensoft.net/fig/484183" pageId="0" pageNumber="99">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="91C286D5271D3A14A0D7404C24273CEF" pageId="0" pageNumber="99" type="diagnosis">
<paragraph id="3A57095D91E49F1026FB654C55B20FC5" pageId="0" pageNumber="99">Diagnosis.</paragraph>
<paragraph id="851F7AA247FF34998D41CC51279F640A" pageId="0" pageNumber="99">
<taxonomicName id="D8B4DB64B78D838F9200FC44B9A6815D" authorityName="Lavandero" authorityYear="2020" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria cantillanensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="cantillanensis">
<emphasis id="E87B508195584AE604CB7D30255A518B" italics="true" pageId="0" pageNumber="99">Leucheria cantillanensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="F2A8B9EBA7F27C908BC3EA18AC1F5894" lsidName="L. salina" pageId="0" pageNumber="99" rank="species" species="salina">
<emphasis id="FCD67A38F409FF5715E5D86E247223AD" italics="true" pageId="0" pageNumber="99">L. salina</emphasis>
</taxonomicName>
but differs in its flat lamina and chartaceous texture (vs. foliar segments perpendicular or oblique to the lamina axis and leathery texture) (Figs
<figureCitation id="8DD40CDCCA9A24A0D304AB64EAAB9460" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Leaf morphology and habit of acaulescent / subacaulescent Leucheria from Central Chile A Leucheria scrobiculata (NL 433, CONC), inset: Habit B Leucheria candidissima (Lavandero, Santilli &amp; Ossa 36, CONC), inset: Habit C Leucheria salina (Lavandero &amp; Abello 40, CONC), inset: Habit D Leucheria cantillanensis (NL 700, CONC), inset: Habit. All photographs by Lavandero." figureDoi="10.3897/phytokeys.169.57532.figure5" httpUri="https://binary.pensoft.net/fig/484184" pageId="0" pageNumber="99">5C, D</figureCitation>
,
<figureCitation id="7189BEFDA7D86ECF7C27F7F6C317E14A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Leucheria salina (A, D, G, J) (Valle Nevado, Region Metropolitana, Chile), Lecheria runcinata (B, E, H, K) (Reserva Natural Altos de Cantillana, Region Metropolitana, Chile), and Leucheria cantillanensis (C, F, I, L) (Reserva Natural Altos de Cantillana, Region Metropolitana, Chile) A-C habit D-F capitula G-I abaxial side of leaves J-L belowground structures. All Photographs by Lavandero." figureDoi="10.3897/phytokeys.169.57532.figure6" httpUri="https://binary.pensoft.net/fig/484185" pageId="0" pageNumber="99">6G, I</figureCitation>
), conspicuously prominent venation, with the secondary and tertiary veins forming a raised pattern above and below the lamina (vs. non-prominent venation, with no raised pattern of veins in either side of the lamina) (Fig.
<figureCitation id="D180AAC781E42C2A9F6375100C0ED59F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Leucheria salina (A, D, G, J) (Valle Nevado, Region Metropolitana, Chile), Lecheria runcinata (B, E, H, K) (Reserva Natural Altos de Cantillana, Region Metropolitana, Chile), and Leucheria cantillanensis (C, F, I, L) (Reserva Natural Altos de Cantillana, Region Metropolitana, Chile) A-C habit D-F capitula G-I abaxial side of leaves J-L belowground structures. All Photographs by Lavandero." figureDoi="10.3897/phytokeys.169.57532.figure6" httpUri="https://binary.pensoft.net/fig/484185" pageId="0" pageNumber="99">6G, I</figureCitation>
), 27-30 flowers per capitula (vs. 40-60), purple anther apical appendages (vs. greenish-yellow) (Fig.
<figureCitation id="35EEA16C51AF89C4885778A55BD085A4" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Leucheria salina (A, D, G, J) (Valle Nevado, Region Metropolitana, Chile), Lecheria runcinata (B, E, H, K) (Reserva Natural Altos de Cantillana, Region Metropolitana, Chile), and Leucheria cantillanensis (C, F, I, L) (Reserva Natural Altos de Cantillana, Region Metropolitana, Chile) A-C habit D-F capitula G-I abaxial side of leaves J-L belowground structures. All Photographs by Lavandero." figureDoi="10.3897/phytokeys.169.57532.figure6" httpUri="https://binary.pensoft.net/fig/484185" pageId="0" pageNumber="99">6D, F</figureCitation>
).
<taxonomicName id="C2519D716ED8BFDD81759630B6368B9D" authorityName="Lavandero" authorityYear="2020" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria cantillanensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="cantillanensis">
<emphasis id="F8A958A336D99E0F156E7C564333A14C" italics="true" pageId="0" pageNumber="99">Leucheria cantillanensis</emphasis>
</taxonomicName>
is also similar to coastal forms of
<taxonomicName id="A59039B53DE3AD1328DEEE3631BF6993" lsidName="L. runcinata" pageId="0" pageNumber="99" rank="species" species="runcinata">
<emphasis id="B325808FFB053308034B9D71F69F7DAB" italics="true" pageId="0" pageNumber="99">L. runcinata</emphasis>
</taxonomicName>
D.Don with white corollas, but it differs by its underground creeping rhizome with nodes and leaf scars and remnants of dry leaf petioles along the rhizome (vs. a lignified taproot with leaf scars concentrated at the base of the stem at the surface of the ground) (Fig.
<figureCitation id="C6F541879FDD3CAABD0D0B82BC173093" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Leucheria salina (A, D, G, J) (Valle Nevado, Region Metropolitana, Chile), Lecheria runcinata (B, E, H, K) (Reserva Natural Altos de Cantillana, Region Metropolitana, Chile), and Leucheria cantillanensis (C, F, I, L) (Reserva Natural Altos de Cantillana, Region Metropolitana, Chile) A-C habit D-F capitula G-I abaxial side of leaves J-L belowground structures. All Photographs by Lavandero." figureDoi="10.3897/phytokeys.169.57532.figure6" httpUri="https://binary.pensoft.net/fig/484185" pageId="0" pageNumber="99">6K, L</figureCitation>
), entirely glandular indumentum in the leaves (vs. glandular on the adaxial side and glandular and lanate on the abaxial side) (Fig.
<figureCitation id="10E97A2C719B4479A61A1E94FBFF8CAC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Leucheria salina (A, D, G, J) (Valle Nevado, Region Metropolitana, Chile), Lecheria runcinata (B, E, H, K) (Reserva Natural Altos de Cantillana, Region Metropolitana, Chile), and Leucheria cantillanensis (C, F, I, L) (Reserva Natural Altos de Cantillana, Region Metropolitana, Chile) A-C habit D-F capitula G-I abaxial side of leaves J-L belowground structures. All Photographs by Lavandero." figureDoi="10.3897/phytokeys.169.57532.figure6" httpUri="https://binary.pensoft.net/fig/484185" pageId="0" pageNumber="99">6H, I</figureCitation>
), conspicuous prominent venation on both sides of the lamina (vs. inconspicuous and only midvein prominent on the abaxial side) and purple anther apical appendages (vs. greyish-blue) (Fig.
<figureCitation id="0EED54116E5D04E6542539789AF66CBB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Leucheria salina (A, D, G, J) (Valle Nevado, Region Metropolitana, Chile), Lecheria runcinata (B, E, H, K) (Reserva Natural Altos de Cantillana, Region Metropolitana, Chile), and Leucheria cantillanensis (C, F, I, L) (Reserva Natural Altos de Cantillana, Region Metropolitana, Chile) A-C habit D-F capitula G-I abaxial side of leaves J-L belowground structures. All Photographs by Lavandero." figureDoi="10.3897/phytokeys.169.57532.figure6" httpUri="https://binary.pensoft.net/fig/484185" pageId="0" pageNumber="99">6E, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FD65752DE92496850D85F327FEF27FBB" pageId="0" pageNumber="99" type="materials_examined">
<paragraph id="15E44E7207B4707C63CD31B4AB8C2041" pageId="0" pageNumber="99">Type.</paragraph>
<paragraph id="F9984C86A5A11AB08409FBE6BBBDF067" pageId="0" pageNumber="99">
Chile.
<normalizedToken id="C6C19F4CE65A27FA67E967CB1214D9D3" originalValue="Región">Region</normalizedToken>
Metropolitana: Provincia de Melipilla, entre el
<normalizedToken id="D7FC49C2F0BB12475D4F577EEE945878" originalValue="límite">limite</normalizedToken>
de
<normalizedToken id="6CC7CCD211A9A2E62F059B3004AFE7AE" originalValue="Alhué">Alhue</normalizedToken>
y Melipilla, Reserva Natural Altos de Cantillana,
<geoCoordinate id="AA38535B6954BE272008E820614835CC" degrees="33" direction="south" minutes="54" orientation="latitude" precision="1" seconds="54.24" value="-33.91507">33°54'54.24&quot;S</geoCoordinate>
,
<geoCoordinate id="FE3A3FC59789A4A7622632BEBC966CCB" degrees="70" direction="west" minutes="58" orientation="longitude" precision="1" seconds="43.57" value="-70.97877">70°58'43.57&quot;W</geoCoordinate>
, 2007 m., 27 December 2019, fl. And fr.,
<emphasis id="A22BEAC523951892F41475730AE4ED6D" italics="true" pageId="0" pageNumber="99">Lavandero</emphasis>
700 (holotype: CONC!; Isotype SGO!).
</paragraph>
</subSubSection>
<subSubSection id="E71EC14FF2844818F8B01248B0E90A62" pageId="0" pageNumber="99" type="description">
<paragraph id="87F2601BE8E93DDE4E59D6D5DFB171D0" pageId="0" pageNumber="99">Description.</paragraph>
<paragraph id="EFD9726BDB51C56B184A0DDA8D21938E" pageId="0" pageNumber="99">
<emphasis id="1E3F091BCC7FB85189BEB8F26E3AEE4F" bold="true" italics="true" pageId="0" pageNumber="99">Perennial</emphasis>
caulescent herb 15-30(-40) cm tall, decumbent, forming clumps of 5-6 aerial stems arising from the apex and nodes of the distal end of the rhizome.
<emphasis id="33E715438309E595B425C71326F15407" bold="true" italics="true" pageId="0" pageNumber="99">Rhizome</emphasis>
dark brown, round, 10-15 mm wide, oblique to creeping, leafless below, but with remnants of dry leaf petioles, roots arising from the internodes.
<emphasis id="E7DF0A5D4A1BFE1315232586F0B6554F" bold="true" italics="true" pageId="0" pageNumber="99">Roots</emphasis>
dark brown, ca. 2 mm wide, round in cross-section.
<emphasis id="4A8521B4CA8E5FC0A396FB4F4A68D1C9" bold="true" italics="true" pageId="0" pageNumber="99">Stems</emphasis>
purplish at the base, green at the top, 1.0-5.5 mm wide, simple or branching, round in cross-section, internodes up to 4 cm long, densely covered by glandular, capitate, (87-)115-180
<normalizedToken id="C2D34B4B58135AC4F0F650647B863DA2" originalValue="µm">µm</normalizedToken>
long, multicellular (6-12-celled) trichomes with clear resin, fragrant, with pungent citric scent (same indumentum up to the corolla tube).
<emphasis id="1C889C1E8B1E5DD64F210BEE3D05A2A0" bold="true" italics="true" pageId="0" pageNumber="99">Leaves</emphasis>
dark green, alternate; basal leaves petiolate, semi-densely arranged at the base; petiole compressed, winged, vaginate, 2-2.5(-3.5) cm long; upper leaves sessile, amplexicaul, loosely arranged, gradually reduced in size towards the capitulescences.
<emphasis id="5C7969D88C4EBA8BDE898E3747423C00" bold="true" italics="true" pageId="0" pageNumber="99">Lamina</emphasis>
obovate, (10-)50-100(-140)
<normalizedToken id="60E57D177A39AA4E8E1BF555DF768C58" originalValue="×">x</normalizedToken>
(5-)20-25(-35) mm; base attenuate, amplexicaul, apex mucronate; margin serrate, texture chartaceous, densely glandulous on both surfaces; pinnatisect to entire towards the tip; segments at the base entire, rarely 1(-2)-dentate, apex mucronate; segments in the middle (3-)4-6(-7)-dentate; apical segments fused, doubly dentate; venation conspicuously prominent, with the secondary and tertiary veins forming a raised pattern on both sides of the lamina, pinnate, semicraspedodromous, with primary vein ending in apical mucro, secondary veins either ending in second-order teeth or joining other distal secondary veins.
<emphasis id="93F9C336E6912FB063FAAD42ABB2D8C1" bold="true" italics="true" pageId="0" pageNumber="99">Capitulescences</emphasis>
a corymbiform cyme.
<emphasis id="15918F2883B7AB4F882D606CCA642545" bold="true" italics="true" pageId="0" pageNumber="99">Capitula</emphasis>
1-6 per stem, pedunculate, (0.5-)1.2-2.8(-3.6) cm long, homogamous, discoid.
<emphasis id="9C30B4974C9E25362D74876B6F169E8A" bold="true" italics="true" pageId="0" pageNumber="99">Involucres</emphasis>
hemispheric 7.0-7.2
<normalizedToken id="718F6EC350CA38FB5C3D55119AB9F302" originalValue="×">x</normalizedToken>
8.2-8.3 mm, two-seriate, alternate. A third series of Middle involucral bracts with intermediate characters between outer and inner Middle involucral bracts rarely present.
<emphasis id="FD40B389642532792BA7777C3CDA1A70" bold="true" italics="true" pageId="0" pageNumber="99">Receptacle</emphasis>
slightly convex, epaleate, glabrous.
<emphasis id="DE72232B4866BBE825FB669EA04E2E35" bold="true" italics="true" pageId="0" pageNumber="99">Outer Middle involucral bracts</emphasis>
5-6, green, lanceolate, concave on the inner face, 6.6-7.6
<normalizedToken id="14E41430E5607AC646974C290B5365A0" originalValue="×">x</normalizedToken>
1.10-1.21(-1.36) mm, with 3 dark-green longitudinal stripes (including the midrib), apex ciliate, margin entire, texture leaf-like, abaxial lamina and margins densely covered by glandular trichomes, adaxial lamina glabrous.
<emphasis id="A01DF1A1F3327C77BBB561E2F14B6821" bold="true" italics="true" pageId="0" pageNumber="99">Middle involucral bracts</emphasis>
rarely present, 1-2, green, lanceolate, concave to flat, with 6.5-7.0
<normalizedToken id="6D838A8D274FDE2B15C0A9A92884163A" originalValue="×">x</normalizedToken>
1.08-1.19 mm, with 3 dark-green longitudinal stripes (including the midrib), apex ciliate, texture leaf-like to hyaline towards the margins, margin ciliate, rarely glandular, central portion of the abaxial lamina covered by glandular trichomes, hyaline lamina glabrous.
<emphasis id="ED4A82AF52D02C9B94C6BBD5AD269169" bold="true" italics="true" pageId="0" pageNumber="99">Inner Middle involucral bracts</emphasis>
5-6(-8), green, lanceolate, concave to flat 6.2-7.1
<normalizedToken id="B3648EECEA3BC8A405726D54FE39CA44" originalValue="×">x</normalizedToken>
1.06-1.25(-1.51) mm, with 3 dark-green longitudinal stripes (including the midrib), apex acute, texture leaf-like to hyaline-membranaceous towards both lateral margins, margin ciliate, cilia (0.11-)0.17-0.21(-0.24) mm long, central portion of the abaxial lamina densely covered by glandular trichomes, hyaline lamina glabrous, adaxial lamina glabrous.
<emphasis id="A2660A35B4067F9F0B4C1FF8F2696D46" bold="true" italics="true" pageId="0" pageNumber="99">Flowers</emphasis>
isomorphic, bisexual, 27-30 per capitulum.
<emphasis id="FB8CBF85F65B57E472F4B84BA713AA40" bold="true" italics="true" pageId="0" pageNumber="99">Corollas</emphasis>
bilabiate, white, before anthesis pinkish-white, tube 3.7-4.0 mm long, 0.5-1.1 wide; corolla tube sparsely covered by glandular trichomes.
<emphasis id="3B83C682A3ACD71D3F3F11F6F264FAAA" bold="true" italics="true" pageId="0" pageNumber="99">Outer lip</emphasis>
oblanceolate, 3.6-3.9
<normalizedToken id="A536401529E2AAC48B18095FE1148DEC" originalValue="×">x</normalizedToken>
2.0-2.2 mm at its widest, apex 3-toothed, teeth equal, 4-veined, glabrous.
<emphasis id="3C53EB9A43F32F7E89B1ADF5F3208956" bold="true" italics="true" pageId="0" pageNumber="99">Inner lip</emphasis>
bifid, lacinae linear, 2.7-2.8
<normalizedToken id="4745BB6AAF12B05BEA441B252223E667" originalValue="×">x</normalizedToken>
0.24-0.37 mm at its widest, connivent, glabrous.
<emphasis id="9A878F9586645ADAB8DD6AECDB5F3F7C" bold="true" italics="true" pageId="0" pageNumber="99">Stamens</emphasis>
5, 3.8-4.0 mm long, glabrous.
<emphasis id="6C0B603418BF1CB9495934FBDAE090D2" bold="true" italics="true" pageId="0" pageNumber="99">Anthers</emphasis>
sagittate, 3.25-3.29 mm long; apical appendages purple, lanceolate, 1.19-1.23 mm long, apex acute; tails long, lanceolate, ca. 0.6 mm long, apex acute, smooth to ciliate.
<emphasis id="B3B5BAA745F3F465F996A8B980DDC2DC" bold="true" italics="true" pageId="0" pageNumber="99">Styles</emphasis>
white, 4.2-5.0 mm long, cleft into two truncate branches, branches 0.54-0.73 mm long, with stigmatic papillae on internal surface.
<emphasis id="026CD4D4B4F7E3E1D99996337E64E46D" bold="true" italics="true" pageId="0" pageNumber="99">Cypselae</emphasis>
dark-brown, 1.0-1.2
<normalizedToken id="FD3885F319B7F457F671438A354EFB78" originalValue="×">x</normalizedToken>
2.4-2.5 mm, obovoid, strigose; trichomes transparent, cylindric, terete, (150-)167-170(-180)
<normalizedToken id="B21DFBFD047BAB412BAF6512B2142FC9" originalValue="µm">µm</normalizedToken>
, ascending, unicellular, subtended by two globose exocarpic cells.
<taxonomicName id="8D7EEB0F23B384A8E28EFA9BA727A67F" class="Insecta" family="Miridae" genus="Pappus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pappus" order="Hemiptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="F6FB0A77E0F630E0FF9B9109F236EA60" bold="true" italics="true" pageId="0" pageNumber="99">Pappus</emphasis>
</taxonomicName>
uniseriate, fused at their bases into a ring, deciduous; bristles 19-20, white, capillary, sub-plumose, 4.5-5 mm long; pectines long, filiform, 0.21-0.35(-0.46) mm long, laterally inserted.
</paragraph>
<caption id="3A1FE41411144C124B1C4A533C58858E" doi="10.3897/phytokeys.169.57532.figure3" httpUri="https://binary.pensoft.net/fig/484182" pageId="0" pageNumber="99" start="Figure 3" startId="F3">
<paragraph id="6F7BE5A56B350FEA9B80334BEE078DA3" pageId="0" pageNumber="99">
<emphasis id="B59CAF64F3BC97918EEBDFD2FC068206" bold="true" pageId="0" pageNumber="99">Figure 3.</emphasis>
<taxonomicName id="CC0AC2089EF38F853EE06FADF6A716CC" authorityName="Lavandero" authorityYear="2020" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria cantillanensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="cantillanensis">
<emphasis id="3003B2FE0CA730BC7F3F62ED123D880E" italics="true" pageId="0" pageNumber="99">Leucheria cantillanensis</emphasis>
</taxonomicName>
Lavandero, sp. nov.
<emphasis id="901E3F5E87901AED25A3DB982CB59E7D" bold="true" pageId="0" pageNumber="99">A</emphasis>
capitulum, sideways view
<emphasis id="D0B4450C7F417505C3FD83168797B019" bold="true" pageId="0" pageNumber="99">B</emphasis>
capitulum, detail of Middle involucral bracts
<emphasis id="DBD3393AFE1CB268A255CBF649395688" bold="true" pageId="0" pageNumber="99">C</emphasis>
leaves
<emphasis id="B911431911530D68A5626834F71033FA" bold="true" pageId="0" pageNumber="99">D</emphasis>
capitulum, upper view
<emphasis id="EED97C772E6CB07400DC74FB7DB731CC" bold="true" pageId="0" pageNumber="99">E</emphasis>
plants growing in natural habitat
<emphasis id="B139990A8B50D2F6AE631821EBF4FE69" bold="true" pageId="0" pageNumber="99">F</emphasis>
cypsela
<emphasis id="501F3EE9F5FB08775E6437DDB0114F9A" bold="true" pageId="0" pageNumber="99">G</emphasis>
detail of trichomes in cypsela. All photographs by
<normalizedToken id="F08C32D36E606A7F1A1792D6522E46DF" originalValue="Nicolás">Nicolas</normalizedToken>
Lavandero.
</paragraph>
</caption>
<caption id="A47FFBA8F51E2E4F6ED1358E935BCDFF" doi="10.3897/phytokeys.169.57532.figure4" httpUri="https://binary.pensoft.net/fig/484183" pageId="0" pageNumber="99" start="Figure 4" startId="F4">
<paragraph id="CACC2FC1FC57BC2A22FA8C115F501C07" pageId="0" pageNumber="99">
<emphasis id="D49477A302C4CC0964875D1535E8E8C7" bold="true" pageId="0" pageNumber="99">Figure 4.</emphasis>
<taxonomicName id="8D5FB9CA2C3C9A8BFE3EFE1686BD6FE0" authorityName="Lavandero" authorityYear="2020" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria cantillanensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="cantillanensis">
<emphasis id="F2A59B3D79D565ABDD17190AA705B8FE" italics="true" pageId="0" pageNumber="99">Leucheria cantillanensis</emphasis>
</taxonomicName>
Lavandero, sp. nov.
<emphasis id="2F7ED7AEDF86E7DAB01F64510E01D1CB" bold="true" pageId="0" pageNumber="99">A</emphasis>
habit
<emphasis id="83E01019A99ACD56D9C72534B53C2362" bold="true" pageId="0" pageNumber="99">B</emphasis>
capitulum
<emphasis id="2BC6431282CDB15B9CB0D3032A90D234" bold="true" pageId="0" pageNumber="99">C</emphasis>
detail of flower
<emphasis id="F3E7B8AC058FA7856553E90D5A3680E8" bold="true" pageId="0" pageNumber="99">D</emphasis>
fruit
<emphasis id="A4C8450ABC3B75911B2A28F0486636C4" bold="true" pageId="0" pageNumber="99">E</emphasis>
basal leaf
<emphasis id="40C2EA9AFAA883A7948D20D63CBF404F" bold="true" pageId="0" pageNumber="99">F</emphasis>
leaf trichome
<emphasis id="BAD24193560AFCF577B276ED3E92272A" bold="true" pageId="0" pageNumber="99">G</emphasis>
stem trichome
<emphasis id="5B859DED2748503C90342ACABD30016F" bold="true" pageId="0" pageNumber="99">H</emphasis>
inner Middle involucral bract
<emphasis id="AF6736193407CB446ACB9AC2B3BF581D" bold="true" pageId="0" pageNumber="99">I</emphasis>
middle Middle involucral bract
<emphasis id="08D8BE1F130FBF4DDB0D206154348B55" bold="true" pageId="0" pageNumber="99">J</emphasis>
outer Middle involucral bract
<emphasis id="552503F220C17803E9648485497AEA83" bold="true" pageId="0" pageNumber="99">K</emphasis>
ciliate margin of inner Middle involucral bract (detail). Illustrations by Benito Rosende.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="434C1B21E0CF5184898E60D3ECE5ADA2" pageId="0" pageNumber="99" type="distribution">
<paragraph id="80CC8F2D7A7265726458C1815F739C48" pageId="0" pageNumber="99">Distribution and habitat.</paragraph>
<paragraph id="1F59E53394B99A39F167FFAC537B761D" pageId="0" pageNumber="99">
<taxonomicName id="E1F5109487D0E17691F14476E1752F78" authorityName="Lavandero" authorityYear="2020" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria cantillanensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="cantillanensis">
<emphasis id="0C6B6EB338BB5AEF1843D9B877CE1FEB" italics="true" pageId="0" pageNumber="99">Leucheria cantillanensis</emphasis>
</taxonomicName>
seems to be endemic to the Cantillana Mountain Range, which is part of the coastal mountain range of central Chile. It grows in shaded crevices of rocky outcrops near 2000 m a.s.l. with SW orientation (Fig.
<figureCitation id="A05F46AF57F5B3CCB2433C85AD5B31DD" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitat of Leucheria cantillanensis in Central Chile A southwest facing rock outcrops, ca. 2000 m elevation B rock outcrops in Cantillana, general view C general view of rock outcrops among Nothofagus macrocarpa (A. DC.) F. M. Vazquez &amp; R. A. Rodr. forests in the Cantillana plateau D Leucheria cantillanensis, detail of the plant growing between rock crevices. Photographs A, D by Lavandero, B, C by Fabiola Gamboa." figureDoi="10.3897/phytokeys.169.57532.figure7" httpUri="https://binary.pensoft.net/fig/484186" pageId="0" pageNumber="99">7</figureCitation>
). It is known thus far only from the type locality (Fig.
<figureCitation id="D37D04F63155D8CA93A3FF87DBBB314F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of Leucheria cantillanensis (star) in Chile, Region Metropolitana, based on the type locality." figureDoi="10.3897/phytokeys.169.57532.figure1" httpUri="https://binary.pensoft.net/fig/484180" pageId="0" pageNumber="99">1</figureCitation>
).
<taxonomicName id="A36BBD0A7EBF19C7E02DB8EE1C4A19A0" lsidName="L. cantillanensis" pageId="0" pageNumber="99" rank="species" species="cantillanensis">
<emphasis id="A5E09312E906A18814AD0E4A43F081CD" italics="true" pageId="0" pageNumber="99">L. cantillanensis</emphasis>
</taxonomicName>
occurs associated with other rupicolous taxa such as
<taxonomicName id="7967753E47A2B1A77042EF15A3BECEDB" class="Eudicots" family="Calceolariaceae" genus="Calceolaria" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Calceolaria andina" order="Lamiales" pageId="0" pageNumber="99" phylum="Magnoliophyta" rank="species" species="andina">
<emphasis id="C4F190BBF8B34E33FEE99618180C6E59" italics="true" pageId="0" pageNumber="99">Calceolaria andina</emphasis>
</taxonomicName>
Benth.
</paragraph>
</subSubSection>
<subSubSection id="2C100E4AFE2B343288BA77CD998C14AF" pageId="0" pageNumber="99" type="phenology">
<paragraph id="A5A6949BF37CC52162C9D94FADBB1BEB" pageId="0" pageNumber="99">Phenology.</paragraph>
<paragraph id="08B72C9DA35E16F4EB9534C5F97241AE" pageId="0" pageNumber="99">Collected flowering and fruiting in December.</paragraph>
</subSubSection>
<subSubSection id="53EE19D49A9058DB8D68C73A5E9E94DB" pageId="0" pageNumber="99" type="etymology">
<paragraph id="34BCD0DEB5D57FC4CC88B1D1F78A5B4A" pageId="0" pageNumber="99">Etymology.</paragraph>
<paragraph id="F268D6226AD40EFD95184BD3FCFA88E6" pageId="0" pageNumber="99">The specific epithet refers to the coastal mountain range where the species was found, Altos de Cantillana.</paragraph>
<caption id="E70A48ED950EA168F89D0042456A9047" doi="10.3897/phytokeys.169.57532.figure5" httpUri="https://binary.pensoft.net/fig/484184" pageId="0" pageNumber="99" start="Figure 5" startId="F5">
<paragraph id="7FABA9331FC7C42DC7559EC1FD1A1A95" pageId="0" pageNumber="99">
<emphasis id="CC8985C2F3FDBB3209CEBAC2B62520A0" bold="true" pageId="0" pageNumber="99">Figure 5.</emphasis>
Leaf morphology and habit of acaulescent/subacaulescent
<taxonomicName id="1CB56ECA7AB98725765B06CFBFDF047C" authorityName="Lagasca" authorityYear="1811" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="CD5DE8AE6E6243A4E5F41B22D1CF4814" italics="true" pageId="0" pageNumber="99">Leucheria</emphasis>
</taxonomicName>
from Central Chile
<emphasis id="49E3A5037C5EC3F6C3F5B3B12D3618E9" bold="true" pageId="0" pageNumber="99">A</emphasis>
<taxonomicName id="8046C95DC2D51EF0BC70D89B6A4A3EB9" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria scrobiculata" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="scrobiculata">
<emphasis id="32A24ACB0EDFC1838EB69B88CD1B42FC" italics="true" pageId="0" pageNumber="99">Leucheria scrobiculata</emphasis>
</taxonomicName>
(NL433, CONC), inset: Habit
<emphasis id="40CA1B9275D4BE78B56B6593343D746D" bold="true" pageId="0" pageNumber="99">B</emphasis>
<taxonomicName id="854723CEB883E0C9EE8190C743D2448A" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria candidissima" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="candidissima">
<emphasis id="3BAC6AF1DD6A13F8C681D2FE38873773" italics="true" pageId="0" pageNumber="99">Leucheria candidissima</emphasis>
</taxonomicName>
(Lavandero, Santilli &amp; Ossa 36, CONC), inset: Habit
<emphasis id="6C4FD2AEC88605A4F80586AF8B96C9F0" bold="true" pageId="0" pageNumber="99">C</emphasis>
<taxonomicName id="1815F59E8B227DA8419DB06C3C15E1AE" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria salina" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="salina">
<emphasis id="6F863B29453C8DACB99621CDEECC67BF" italics="true" pageId="0" pageNumber="99">Leucheria salina</emphasis>
</taxonomicName>
(Lavandero &amp; Abello 40, CONC), inset: Habit
<emphasis id="0D8AEA2ADB2D50551A2BF5F799641F4C" bold="true" pageId="0" pageNumber="99">D</emphasis>
<taxonomicName id="3717B302BA96CFDDB1291CEB78BEEE12" authorityName="Lavandero" authorityYear="2020" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria cantillanensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="cantillanensis">
<emphasis id="CB49D93CC7EFBF75114AFAB4183E8DD9" italics="true" pageId="0" pageNumber="99">Leucheria cantillanensis</emphasis>
</taxonomicName>
(NL700, CONC), inset: Habit. All photographs by Lavandero.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C5569A070878F4D3F906D2D9E4431923" pageId="0" pageNumber="99" type="conservation status">
<paragraph id="FFAA263AF814462B3CF74CDBA10546B7" pageId="0" pageNumber="99">Conservation status.</paragraph>
<paragraph id="D1DC6CD09484E44622C05E5919671964" pageId="0" pageNumber="99">
<taxonomicName id="3DA39F47143162E914738F568AEC43EC" authorityName="Lavandero" authorityYear="2020" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria cantillanensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="cantillanensis">
<emphasis id="734373071EEC9293308BF58CFA8CFEEA" italics="true" pageId="0" pageNumber="99">Leucheria cantillanensis</emphasis>
</taxonomicName>
is assessed here as Critically Endangered (CR) under the IUCN categories and criteria B2ab(i,ii,iii). Criterion B2 was selected because its Area of Occupancy is &lt;10 km2 (4 km2). Criterion
<normalizedToken id="82DFD16DD74A8DD3FA3A68D9161886A5" originalValue="“a”">&quot;a&quot;</normalizedToken>
was selected because it is known to exist at only a single location, with only one subpopulation. Criterion b(i,ii,iii) was selected because we expect a continuing decline of suitable area for the species to exist in since it is only found at the highest elevations within the mountain coastal range with very specific soil types and exposition. The quality of its habitat has also been deteriorating over time. The overall precipitation and snow cover in the Cantillana plateau has decreased dramatically over the past 20 years, affecting not only the Andean relict flora but all the vegetation in the area. Fog events which compensate for the drought over the summer, are not as common as before.
<taxonomicName id="A1BA993FE2FF823A57529CAC5D97A0DA" authorityName="Lavandero" authorityYear="2020" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria cantillanensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="cantillanensis">
<emphasis id="7F4E91D52060C594FF847F8E15AEA496" italics="true" pageId="0" pageNumber="99">Leucheria cantillanensis</emphasis>
</taxonomicName>
is present in the private reserve &quot;Reserva Natural Altos de Cantillana&quot;.
</paragraph>
<caption id="B2110BC3FCCDDBCB7468D79C9AEBDAAC" doi="10.3897/phytokeys.169.57532.figure6" httpUri="https://binary.pensoft.net/fig/484185" pageId="0" pageNumber="99" start="Figure 6" startId="F6">
<paragraph id="FBD08FE34FFB4C03869E6ED52D75C9E6" pageId="0" pageNumber="99">
<emphasis id="7DB808B8C19C186AA902A643405DA869" bold="true" pageId="0" pageNumber="99">Figure 6.</emphasis>
<taxonomicName id="D4E32A3DC58DE350919CD63CE0AEE473" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria salina" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="salina">
<emphasis id="264FAF8120F26179C005EB3A780A87BD" italics="true" pageId="0" pageNumber="99">Leucheria salina</emphasis>
</taxonomicName>
(
<emphasis id="D6BA346FC9D8415CE32F9D220833E61E" bold="true" pageId="0" pageNumber="99">A, D, G, J</emphasis>
) (Valle Nevado,
<normalizedToken id="7DBBEC4110927B1932483DC25C2F5F6D" originalValue="Región">Region</normalizedToken>
Metropolitana, Chile),
<taxonomicName id="635D9E4F15966E555BA09AFA61C2E604" genus="Lecheria" lsidName="Lecheria runcinata" pageId="0" pageNumber="99" rank="species" species="runcinata">
<emphasis id="72885999CB12E13EB8791CCE288D4553" italics="true" pageId="0" pageNumber="99">Lecheria runcinata</emphasis>
</taxonomicName>
(
<emphasis id="02206AB01703B4D27F0501BEA6217EA4" bold="true" pageId="0" pageNumber="99">B, E, H, K</emphasis>
) (Reserva Natural Altos de Cantillana,
<normalizedToken id="E5A0AED44865C34E2FEF66C704EE586C" originalValue="Región">Region</normalizedToken>
Metropolitana, Chile), and
<taxonomicName id="1DE766383FD5F0DF4AE05C5F3C5A7322" authorityName="Lavandero" authorityYear="2020" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria cantillanensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="cantillanensis">
<emphasis id="A4936D50FE9FE869C247403D55426F7F" italics="true" pageId="0" pageNumber="99">Leucheria cantillanensis</emphasis>
</taxonomicName>
(
<emphasis id="4B0EFB0A9CE6644638F2043ADBC60C57" bold="true" pageId="0" pageNumber="99">C, F, I, L</emphasis>
) (Reserva Natural Altos de Cantillana,
<normalizedToken id="10EEF8B86F7618495202A91449669BE5" originalValue="Región">Region</normalizedToken>
Metropolitana, Chile)
<emphasis id="F9D9F55307CBD98F264A8E6776F44443" bold="true" pageId="0" pageNumber="99">A-C</emphasis>
habit
<emphasis id="7CED8BBCAD4CC335D20B991897F2E0F6" bold="true" pageId="0" pageNumber="99">D-F</emphasis>
capitula
<emphasis id="1FC28E254F74C7B52274972BC595C468" bold="true" pageId="0" pageNumber="99">G-I</emphasis>
abaxial side of leaves
<emphasis id="A0272B3B14FC69FE9FAF4D5F1DD11D80" bold="true" pageId="0" pageNumber="99">J-L</emphasis>
belowground structures. All Photographs by Lavandero.
</paragraph>
</caption>
<caption id="3CA0EB181A13B3EEDBCE59F7C0CEAAB2" doi="10.3897/phytokeys.169.57532.figure7" httpUri="https://binary.pensoft.net/fig/484186" pageId="0" pageNumber="99" start="Figure 7" startId="F7">
<paragraph id="FC2FA21588A67FD31CAB43354DC5E8C3" pageId="0" pageNumber="99">
<emphasis id="37219F32C23E41845CE2E8A577D78213" bold="true" pageId="0" pageNumber="99">Figure 7.</emphasis>
Habitat of
<taxonomicName id="FF00633A884D59F09A54050DEEAA18A6" authorityName="Lavandero" authorityYear="2020" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria cantillanensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="cantillanensis">
<emphasis id="A9AB6DD5DDDF62218BD918BA2E566FBC" italics="true" pageId="0" pageNumber="99">Leucheria cantillanensis</emphasis>
</taxonomicName>
in Central Chile
<emphasis id="859D1C94F792FDEB5DFA572DD874BC68" bold="true" pageId="0" pageNumber="99">A</emphasis>
southwest facing rock outcrops, ca. 2000 m elevation
<emphasis id="67F0B8F519F876695724AE1FAC3C1EC5" bold="true" pageId="0" pageNumber="99">B</emphasis>
rock outcrops in Cantillana, general view
<emphasis id="77CCFB3EB77C15FC4FD82F3E6C8828CF" bold="true" pageId="0" pageNumber="99">C</emphasis>
general view of rock outcrops among
<taxonomicName id="CEF8AD95D2ABFC6AF0B06FAF2E1F5744" class="Rosopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Nothofagus macrocarpa" order="Fagales" pageId="0" pageNumber="99" phylum="Magnoliophyta" rank="species" species="macrocarpa">
<emphasis id="7BBEE6C7F688ED8B115F101D5A7CB195" italics="true" pageId="0" pageNumber="99">Nothofagus macrocarpa</emphasis>
</taxonomicName>
(A.DC.) F.M.
<normalizedToken id="DA19973241731F7C908629A0E364A61F" originalValue="Vázquez">Vazquez</normalizedToken>
&amp; R.A. Rodr. forests in the Cantillana plateau
<emphasis id="286716910CECE36FAB1C846BA265F95B" bold="true" pageId="0" pageNumber="99">D</emphasis>
<taxonomicName id="4794CE7342B02923975CD03592183275" authorityName="Lavandero" authorityYear="2020" class="Insecta" family="Asteraceae" genus="Leucheria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucheria cantillanensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="cantillanensis">
<emphasis id="8B713AF668E8CCAE608C667E57D6893C" italics="true" pageId="0" pageNumber="99">Leucheria cantillanensis</emphasis>
</taxonomicName>
, detail of the plant growing between rock crevices. Photographs
<emphasis id="2238DF6B06BD8CEB6CF6296225581835" bold="true" pageId="0" pageNumber="99">A, D</emphasis>
by Lavandero,
<emphasis id="636D11A15E194A53D38BB18906BC7A34" bold="true" pageId="0" pageNumber="99">B, C</emphasis>
by Fabiola Gamboa.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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