treatments-xml/data/B3/64/87/B36487D8FFD8D525D108FA5BFEC9F9F8.xml
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<document id="1590DA3A3BA14D6D75E104B6303061B3" ID-DOI="10.5281/zenodo.5393630" ID-ISSN="1638-9387" ID-Zenodo-Dep="5393630" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630319175263" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; Spiegel, Didier Van Den" docDate="2007" docId="B36487D8FFD8D525D108FA5BFEC9F9F8" docLanguage="en" docName="Zoosystema.2007.29.3.417-456.pdf" docOrigin="Zoosystema 29 (3)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Glyphiulus paracostulifer Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel 2007, n. sp." docType="treatment" docVersion="3" lastPageNumber="446" masterDocId="4F5DFFA0FFC1D538D20EFFF0FFA5FC3D" masterDocTitle="Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group" masterLastPageNumber="456" masterPageNumber="417" pageNumber="442" updateTime="1699225420998" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="34E6742D35E02CD8C0CBA738DEC5AF18">Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group</mods:title>
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<mods:namePart id="16EAB268F2DBF97EB60189E59DA6759C">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="4C4D5096D0EC9D49CC10257548355F98">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
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<mods:namePart id="55A4C5956C697CC1EA8D2C07B391532C">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="DBF75E6E4FF40D015EC6867A66FCFD5E">Muséum national dHistoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
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<mods:namePart id="7024181D080979F77AF90C1CA66D3F8F">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="5DC532E6B2C32B2EF788B13F791905F5">Muséum national dHistoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
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<mods:namePart id="41030FF44CA1F6CE1E179878124FD84B">Spiegel, Didier Van Den</mods:namePart>
<mods:affiliation id="DF9B9542B01468FECAFDDF6E06F8D461">Musée royal de lAfrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be</mods:affiliation>
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<mods:title id="CCD97E0AD1048B17BD52F81C75FC5D01">Zoosystema</mods:title>
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<mods:date id="868A2C013051420AD341E22C29479A94">2007</mods:date>
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<heading id="603A81A2FFD8D521D108FA5BFBFDF9FB" box="[774,1112,1451,1478]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="442" reason="2">
<taxonomicName id="FCCD4D4DFFD8D521D108FA5BFBB2F9FB" ID-CoL="3GGKH" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[774,1047,1451,1478]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="442" phylum="Arthropoda" rank="species" species="paracostulifer" status="sp. nov.">
<emphasis id="09B9EADCFFD8D521D108FA5BFBB2F9FB" bold="true" box="[774,1047,1451,1478]" italics="true" pageId="25" pageNumber="442">Glyphiulus paracostulifer</emphasis>
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<taxonomicNameLabel id="128A57A7FFD8D521D613FA5CFBFDF9FB" box="[1053,1112,1452,1478]" pageId="25" pageNumber="442" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="3B7236CEFFD8D521D163FA3BFC54F9D8" blockId="25.[774,1112,1451,1509]" box="[877,1009,1483,1509]" pageId="25" pageNumber="442">
(
<figureCitation id="A3F62A4BFFD8D521D17AFA3BFC4CF9D8" box="[884,1001,1483,1509]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="28.[113,124,1657,1674]" captionStartId-1="29.[113,124,1119,1136]" captionStartId-2="30.[113,124,1286,1303]" captionTargetBox-0="[116,1205,215,1587]" captionTargetBox-1="[117,1205,214,1085]" captionTargetBox-2="[115,1207,204,1257]" captionTargetId-0="figure-146@28.[628,876,845,1029]" captionTargetId-1="figure-124@29.[614,868,643,820]" captionTargetId-2="figure-23@30.[115,1207,204,1257]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionTargetPageId-2="30" captionText-0="FIG. 22.— Glyphiulus paracostulifer n.sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: 0.5 mm." captionText-1="FIG. 23. — Glyphiulus paracostulifer n. sp., paratype ♀: A, midbody segment section; B, midbody leg; C, claw; D, antennal sensilla. Scale bars: A, 0.5 mm; B, 0.2 mm; C, 0.1 mm; D, 0.05 mm." captionText-2="FIG. 24. — Glyphiulus paracostulifer n. sp., paratype ♂: A, antenna; B, leg 1, frontal view; C, leg 2, caudal view; D, leg 3; E, leg 11; F, anterior gonopods, caudal view; G, posterior gonopods, frontal view. Scale bar: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5393680" figureDoi-1="http://doi.org/10.5281/zenodo.5393682" figureDoi-2="http://doi.org/10.5281/zenodo.5393684" httpUri-0="https://zenodo.org/record/5393680/files/figure.png" httpUri-1="https://zenodo.org/record/5393682/files/figure.png" httpUri-2="https://zenodo.org/record/5393684/files/figure.png" pageId="25" pageNumber="442">Figs 22-24</figureCitation>
)
</paragraph>
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<subSubSection id="73D76545FFD8D521D0ABF9D2FCD9FA9E" pageId="25" pageNumber="442" type="materials_examined">
<paragraph id="3B7236CEFFD8D521D0ABF9D2FCD9FA9E" blockId="25.[677,1211,1569,1699]" pageId="25" pageNumber="442">
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<collectionCode id="5DDCAE0BFFD8D521D0ABF9D2FCE4FA05" box="[677,833,1570,1593]" pageId="25" pageNumber="442">TYPE MATERIAL</collectionCode>
. —
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<collectingCountry id="43DA765EFFD8D521D17AF9D3FC11FA06" box="[884,948,1571,1595]" name="China" pageId="25" pageNumber="442">China</collectingCountry>
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.
<collectingRegion id="F909F82CFFD8D521D1CAF9D1FBFBFA04" box="[964,1118,1569,1593]" country="China" name="Guizhou" pageId="25" pageNumber="442">Guizhou Prov.</collectingRegion>
,
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,
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,
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,
<date id="4F73100EFFD8D521D614F9CCFB2BFA69" box="[1050,1166,1596,1620]" pageId="25" pageNumber="442" value="2003-11-21">
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, leg.
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&amp;
<collectorName id="96385318FFD8D521D128F9A6FC0AFA53" box="[806,943,1622,1646]" pageId="25" pageNumber="442">S. Meggiorini</collectorName>
,
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(
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;
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(
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;
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(
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;
<materialsCitation id="8BA53C93FFD8D521D0ABF97BFCDDFA9E" box="[677,888,1675,1699]" collectionCode="MCSNV, SEM" country="China" pageId="25" pageNumber="442" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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(
<collectionCode id="5DDCAE0BFFD8D521D0D8F97BFC96FA9E" box="[726,819,1675,1699]" pageId="25" pageNumber="442">MCSNV</collectionCode>
/
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)
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.
</paragraph>
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<caption id="6FB26646FFDBD522D27FFA29FF6CFA25" ID-DOI="http://doi.org/10.5281/zenodo.5393676" ID-Zenodo-Dep="5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="26" pageNumber="443" startId="26.[113,124,1497,1514]" targetBox="[118,1205,215,1453]" targetPageId="26">
<paragraph id="3B7236CEFFDBD522D27FFA29FF6CFA25" blockId="26.[113,1211,1497,1560]" pageId="26" pageNumber="443">
FIG. 20. —
<taxonomicName id="FCCD4D4DFFDBD522D2C5FA29FE28F9D7" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[203,397,1497,1514]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="26" pageNumber="443" phylum="Arthropoda" rank="species" species="percostulifer" status="sp. nov.">
<emphasis id="09B9EADCFFDBD522D2C5FA29FE28F9D7" box="[203,397,1497,1514]" italics="true" pageId="26" pageNumber="443">Glyphiulus percostulifer</emphasis>
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, paratype ♀:
<emphasis id="09B9EADCFFDBD522D021FA29FD99F9D7" bold="true" box="[559,572,1497,1514]" pageId="26" pageNumber="443">A</emphasis>
, anterior part of body, lateral view;
<emphasis id="09B9EADCFFDBD522D151FA29FCC9F9D7" bold="true" box="[863,876,1497,1514]" pageId="26" pageNumber="443">B</emphasis>
, midbody segment section;
<emphasis id="09B9EADCFFDBD522D656FA29FBC3F9D7" bold="true" box="[1112,1126,1497,1514]" pageId="26" pageNumber="443">C</emphasis>
, midbody segments, lateral view;
<emphasis id="09B9EADCFFDBD522D333FA00FEEEFA3C" bold="true" box="[317,331,1520,1537]" pageId="26" pageNumber="443">D</emphasis>
, same, dorsal view;
<emphasis id="09B9EADCFFDBD522D3F0FA00FDAFFA3C" bold="true" box="[510,522,1520,1537]" pageId="26" pageNumber="443">E</emphasis>
, caudal body end, lateral view;
<emphasis id="09B9EADCFFDBD522D111FA00FC8FFA3C" bold="true" box="[799,810,1520,1537]" pageId="26" pageNumber="443">F</emphasis>
, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm.
</paragraph>
</caption>
<subSubSection id="73D76545FFDBD522D27FF9CBFEBBFA51" pageId="26" pageNumber="443" type="etymology">
<paragraph id="3B7236CEFFDBD522D27FF9CBFEBBFA51" blockId="26.[113,646,1594,1644]" pageId="26" pageNumber="443">
ETYMOLOGY. — To emphasize the close resemblance to
<taxonomicName id="FCCD4D4DFFDBD522D27FF9A5FF46FA51" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[113,227,1620,1644]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="26" pageNumber="443" phylum="Arthropoda" rank="species" species="costulifer" status="sp. nov.">
<emphasis id="09B9EADCFFDBD522D27FF9A5FF46FA51" box="[113,227,1620,1644]" italics="true" pageId="26" pageNumber="443">G. costulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFDBD522D2E7F9A5FEBBFA51" box="[233,286,1621,1644]" pageId="26" pageNumber="443" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="73D76545FFDBD522D27FF97AFD4BFA9C" pageId="26" pageNumber="443" type="diagnosis">
<paragraph id="3B7236CEFFDBD522D27FF97AFD4BFA9C" blockId="26.[113,646,1673,1698]" lastBlockId="26.[677,1209,1594,1697]" pageId="26" pageNumber="443">DIAGNOSIS. — Differs from congeners in the relatively large size, the completely unpigmented tegument and ocelli, the undivided tergal crests, coupled with certain details of leg and gonopod structure (see also key below).</paragraph>
</subSubSection>
<caption id="6FB26646FFDAD523D27FFB5CFB36F8EE" ID-DOI="http://doi.org/10.5281/zenodo.5393678" ID-Zenodo-Dep="5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="27" pageNumber="444" startId="27.[113,124,1196,1213]" targetBox="[128,1201,226,1152]" targetPageId="27">
<paragraph id="3B7236CEFFDAD523D27FFB5CFB36F8EE" blockId="27.[113,1211,1196,1235]" pageId="27" pageNumber="444">
FIG. 21. —
<taxonomicName id="FCCD4D4DFFDAD523D2C0FB5CFE34F880" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[206,401,1196,1213]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="27" pageNumber="444" phylum="Arthropoda" rank="species" species="percostulifer" status="sp. nov.">
<emphasis id="09B9EADCFFDAD523D2C0FB5CFE34F880" box="[206,401,1196,1213]" italics="true" pageId="27" pageNumber="444">Glyphiulus percostulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFDAD523D399FB5CFE6FF880" box="[407,458,1196,1213]" pageId="27" pageNumber="444" rank="species">n. sp.</taxonomicNameLabel>
, paratype ♂:
<emphasis id="09B9EADCFFDAD523D034FB5CFDE2F880" bold="true" box="[570,583,1196,1213]" pageId="27" pageNumber="444">A</emphasis>
, antenna;
<emphasis id="09B9EADCFFDAD523D0AEFB5CFD08F880" bold="true" box="[672,685,1196,1213]" pageId="27" pageNumber="444">B</emphasis>
, gnathochilarium, ventral view;
<emphasis id="09B9EADCFFDAD523D1B9FB5CFC60F880" bold="true" box="[951,965,1196,1213]" pageId="27" pageNumber="444">C</emphasis>
, leg 1, caudal view;
<emphasis id="09B9EADCFFDAD523D67CFB5CFB25F880" bold="true" box="[1138,1152,1196,1213]" pageId="27" pageNumber="444">D</emphasis>
, leg 2, caudal view;
<emphasis id="09B9EADCFFDAD523D2D0FB32FF4FF8EE" bold="true" box="[222,234,1218,1235]" pageId="27" pageNumber="444">E</emphasis>
, leg 3;
<emphasis id="09B9EADCFFDAD523D326FB32FE96F8EE" bold="true" box="[296,307,1218,1235]" pageId="27" pageNumber="444">F</emphasis>
, leg 13;
<emphasis id="09B9EADCFFDAD523D374FB32FE2DF8EE" bold="true" box="[378,392,1218,1235]" pageId="27" pageNumber="444">G</emphasis>
, anterior gonopods, caudal view;
<emphasis id="09B9EADCFFDAD523D0AAFB32FD17F8EE" bold="true" box="[676,690,1218,1235]" pageId="27" pageNumber="444">H</emphasis>
, right posterior gonopod, frontal view. Scale bar: 0.2 mm.
</paragraph>
</caption>
<subSubSection id="73D76545FFDAD525D27FFAFDFD92F93A" lastPageId="29" lastPageNumber="446" pageId="27" pageNumber="444" type="description">
<paragraph id="3B7236CEFFDAD523D27FFAFDFEA7F918" blockId="27.[113,647,1293,1700]" box="[113,258,1293,1319]" pageId="27" pageNumber="444">DESCRIPTION</paragraph>
<paragraph id="3B7236CEFFDAD523D27FFADDFE9CF99B" blockId="27.[113,647,1293,1700]" pageId="27" pageNumber="444">
<typeStatus id="E476886CFFDAD523D27FFADDFF72F97A" box="[113,215,1325,1351]" pageId="27" pageNumber="444" type="holotype">Holotype</typeStatus>
with 65p+1a+T, about
<quantity id="FC359B2BFFDAD523D3DAFADDFD87F97A" box="[468,546,1325,1351]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="27" pageNumber="444" unit="mm" value="43.0">43 mm</quantity>
long and 2.0 mm wide; adult
<typeStatus id="E476886CFFDAD523D350FABDFE6DF95A" box="[350,456,1357,1383]" pageId="27" pageNumber="444" type="paratype">paratypes</typeStatus>
with 52p+2a+T, 55p+3a+T or 57p+1a+T, all about
<quantity id="FC359B2BFFDAD523D3F5FA9CFDE8F9BA" box="[507,589,1388,1415]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="27" pageNumber="444" unit="mm" value="36.0">36 mm</quantity>
long and 2.0 mm wide.
</paragraph>
<paragraph id="3B7236CEFFDAD523D286FA5BFE93FA19" blockId="27.[113,647,1293,1700]" pageId="27" pageNumber="444">Coloration rather uniformly pale yellowish to brownish, only some ozoporiferous cones a little darker; ocelli unpigmented, a few barely discernible per eye patch.</paragraph>
<paragraph id="3B7236CEFFDAD525D286F9DAFD92F93A" blockId="27.[113,647,1293,1700]" lastBlockId="29.[113,646,1198,1288]" lastPageId="29" lastPageNumber="446" pageId="27" pageNumber="444">
Gnathochilarium (mentum divided, n = 1), postcollar constriction (
<figureCitation id="A3F62A4BFFDAD523D334F9BAFE3DFA59" box="[314,408,1610,1636]" captionStart="FIG" captionStartId="28.[113,124,1657,1674]" captionTargetBox="[116,1205,215,1587]" captionTargetId="figure-146@28.[628,876,845,1029]" captionTargetPageId="28" captionText="FIG. 22.— Glyphiulus paracostulifer n.sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393680" httpUri="https://zenodo.org/record/5393680/files/figure.png" pageId="27" pageNumber="444">Fig. 22B</figureCitation>
), segments texture and cross-section (
<figureCitation id="A3F62A4BFFDAD523D30FF999FEEFFAB9" box="[257,330,1641,1668]" captionStart="FIG" captionStartId="28.[113,124,1657,1674]" captionTargetBox="[116,1205,215,1587]" captionTargetId="figure-146@28.[628,876,845,1029]" captionTargetPageId="28" captionText="FIG. 22.— Glyphiulus paracostulifer n.sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393680" httpUri="https://zenodo.org/record/5393680/files/figure.png" pageId="27" pageNumber="444">Fig. 22</figureCitation>
A-E), most of telson (
<figureCitation id="A3F62A4BFFDAD523D02AF999FF27FA99" captionStart="FIG" captionStartId="28.[113,124,1657,1674]" captionTargetBox="[116,1205,215,1587]" captionTargetId="figure-146@28.[628,876,845,1029]" captionTargetPageId="28" captionText="FIG. 22.— Glyphiulus paracostulifer n.sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393680" httpUri="https://zenodo.org/record/5393680/files/figure.png" pageId="27" pageNumber="444">Fig. 22E, F</figureCitation>
), ventral flaps of male segment 7, male legs 2 and 3 (
<figureCitation id="A3F62A4BFFDAD523D0A2FAFDFC8CF91A" box="[684,809,1293,1319]" captionStart="FIG" captionStartId="30.[113,124,1286,1303]" captionTargetBox="[115,1207,204,1257]" captionTargetId="figure-23@30.[115,1207,204,1257]" captionTargetPageId="30" captionText="FIG. 24. — Glyphiulus paracostulifer n. sp., paratype ♂: A, antenna; B, leg 1, frontal view; C, leg 2, caudal view; D, leg 3; E, leg 11; F, anterior gonopods, caudal view; G, posterior gonopods, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393684" httpUri="https://zenodo.org/record/5393684/files/figure.png" pageId="27" pageNumber="444">Fig. 24C, D</figureCitation>
), posterior gonopods (
<figureCitation id="A3F62A4BFFDAD523D600FAFDFBCEF91A" box="[1038,1131,1293,1319]" captionStart="FIG" captionStartId="30.[113,124,1286,1303]" captionTargetBox="[115,1207,204,1257]" captionTargetId="figure-23@30.[115,1207,204,1257]" captionTargetPageId="30" captionText="FIG. 24. — Glyphiulus paracostulifer n. sp., paratype ♂: A, antenna; B, leg 1, frontal view; C, leg 2, caudal view; D, leg 3; E, leg 11; F, anterior gonopods, caudal view; G, posterior gonopods, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393684" httpUri="https://zenodo.org/record/5393684/files/figure.png" pageId="27" pageNumber="444">Fig. 24G</figureCitation>
) generally as in
<taxonomicName id="FCCD4D4DFFDAD523D10FFADDFC3FF97A" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[769,922,1325,1351]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="27" pageNumber="444" phylum="Arthropoda" rank="species" species="subcostulifer" status="sp. nov.">
<emphasis id="09B9EADCFFDAD523D10FFADDFC3FF97A" box="[769,922,1325,1351]" italics="true" pageId="27" pageNumber="444">G. subcostulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFDAD523D190FADDFC72F97A" box="[926,983,1325,1351]" pageId="27" pageNumber="444" rank="species">n. sp.</taxonomicNameLabel>
, but antennae slightly more elongate and less rich in sensilla (
<figureCitation id="A3F62A4BFFDAD523D613FABCFBDFF95B" box="[1053,1146,1356,1382]" captionStart="FIG" captionStartId="28.[113,124,1657,1674]" captionTargetBox="[116,1205,215,1587]" captionTargetId="figure-146@28.[628,876,845,1029]" captionTargetPageId="28" captionText="FIG. 22.— Glyphiulus paracostulifer n.sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393680" httpUri="https://zenodo.org/record/5393680/files/figure.png" pageId="27" pageNumber="444">Figs 22A</figureCitation>
;
<figureCitation id="A3F62A4BFFDAD523D68DFABCFB10F95B" box="[1155,1205,1356,1382]" captionStart="FIG" captionStartId="29.[113,124,1119,1136]" captionTargetBox="[117,1205,214,1085]" captionTargetId="figure-124@29.[614,868,643,820]" captionTargetPageId="29" captionText="FIG. 23. — Glyphiulus paracostulifer n. sp., paratype ♀: A, midbody segment section; B, midbody leg; C, claw; D, antennal sensilla. Scale bars: A, 0.5 mm; B, 0.2 mm; C, 0.1 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393682" httpUri="https://zenodo.org/record/5393682/files/figure.png" pageId="27" pageNumber="444">23D</figureCitation>
;
<figureCitation id="A3F62A4BFFDAD523D0ABFA9CFD71F9BB" box="[677,724,1388,1414]" captionStart="FIG" captionStartId="30.[113,124,1286,1303]" captionTargetBox="[115,1207,204,1257]" captionTargetId="figure-23@30.[115,1207,204,1257]" captionTargetPageId="30" captionText="FIG. 24. — Glyphiulus paracostulifer n. sp., paratype ♂: A, antenna; B, leg 1, frontal view; C, leg 2, caudal view; D, leg 3; E, leg 11; F, anterior gonopods, caudal view; G, posterior gonopods, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393684" httpUri="https://zenodo.org/record/5393684/files/figure.png" pageId="27" pageNumber="444">24A</figureCitation>
); collum (
<figureCitation id="A3F62A4BFFDAD523D137FA9CFC0BF9BA" box="[825,942,1388,1415]" captionStart="FIG" captionStartId="28.[113,124,1657,1674]" captionTargetBox="[116,1205,215,1587]" captionTargetId="figure-146@28.[628,876,845,1029]" captionTargetPageId="28" captionText="FIG. 22.— Glyphiulus paracostulifer n.sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393680" httpUri="https://zenodo.org/record/5393680/files/figure.png" pageId="27" pageNumber="444">Fig. 22A, B</figureCitation>
) with a nearly typical carinotaxy formula (I-III+4c+5a+pc+ma+pc+5a+4c+III- I), crests evident; all tergal crests (
<figureCitation id="A3F62A4BFFDAD523D60EFA5BFBE9F9F8" box="[1024,1100,1451,1477]" captionStart="FIG" captionStartId="28.[113,124,1657,1674]" captionTargetBox="[116,1205,215,1587]" captionTargetId="figure-146@28.[628,876,845,1029]" captionTargetPageId="28" captionText="FIG. 22.— Glyphiulus paracostulifer n.sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393680" httpUri="https://zenodo.org/record/5393680/files/figure.png" pageId="27" pageNumber="444">Figs 22</figureCitation>
A-E; 23A) on subsequent body segments higher, especially so until about segment 9 or 10, virtually not divided (2+I/i+3+I/i+2), ozoporiferous cones only barely lower than wide; epiproct with an evident dorsal rib (
<figureCitation id="A3F62A4BFFDAD523D0DFF9BAFC95FA59" box="[721,816,1610,1636]" captionStart="FIG" captionStartId="28.[113,124,1657,1674]" captionTargetBox="[116,1205,215,1587]" captionTargetId="figure-146@28.[628,876,845,1029]" captionTargetPageId="28" captionText="FIG. 22.— Glyphiulus paracostulifer n.sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393680" httpUri="https://zenodo.org/record/5393680/files/figure.png" pageId="27" pageNumber="444">Fig. 22E</figureCitation>
); legs about as long as body diameter (
<figureCitation id="A3F62A4BFFDAD523D0A2F999FC8CFAB9" box="[684,809,1641,1668]" captionStart="FIG" captionStartId="29.[113,124,1119,1136]" captionTargetBox="[117,1205,214,1085]" captionTargetId="figure-124@29.[614,868,643,820]" captionTargetPageId="29" captionText="FIG. 23. — Glyphiulus paracostulifer n. sp., paratype ♀: A, midbody segment section; B, midbody leg; C, claw; D, antennal sensilla. Scale bars: A, 0.5 mm; B, 0.2 mm; C, 0.1 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393682" httpUri="https://zenodo.org/record/5393682/files/figure.png" pageId="27" pageNumber="444">Fig. 23A, B</figureCitation>
), at most with only a vestigial spine at claw base (
<figureCitation id="A3F62A4BFFDAD523D11DF979FCD3FA9E" box="[787,886,1673,1699]" captionStart="FIG" captionStartId="29.[113,124,1119,1136]" captionTargetBox="[117,1205,214,1085]" captionTargetId="figure-124@29.[614,868,643,820]" captionTargetPageId="29" captionText="FIG. 23. — Glyphiulus paracostulifer n. sp., paratype ♀: A, midbody segment section; B, midbody leg; C, claw; D, antennal sensilla. Scale bars: A, 0.5 mm; B, 0.2 mm; C, 0.1 mm; D, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393682" httpUri="https://zenodo.org/record/5393682/files/figure.png" pageId="27" pageNumber="444">Figs 23C</figureCitation>
;
<figureCitation id="A3F62A4BFFDAD523D18EF979FC0AFA9E" box="[896,943,1673,1699]" captionStart="FIG" captionStartId="30.[113,124,1286,1303]" captionTargetBox="[115,1207,204,1257]" captionTargetId="figure-23@30.[115,1207,204,1257]" captionTargetPageId="30" captionText="FIG. 24. — Glyphiulus paracostulifer n. sp., paratype ♂: A, antenna; B, leg 1, frontal view; C, leg 2, caudal view; D, leg 3; E, leg 11; F, anterior gonopods, caudal view; G, posterior gonopods, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393684" httpUri="https://zenodo.org/record/5393684/files/figure.png" pageId="27" pageNumber="444">24E</figureCitation>
); male leg 1 with reduced 5-segmented telopodites and large paramedian coxal processes (
<figureCitation id="A3F62A4BFFDCD525D2EFFB3DFEE6F8D5" box="[225,323,1229,1256]" captionStart="FIG" captionStartId="30.[113,124,1286,1303]" captionTargetBox="[115,1207,204,1257]" captionTargetId="figure-23@30.[115,1207,204,1257]" captionTargetPageId="30" captionText="FIG. 24. — Glyphiulus paracostulifer n. sp., paratype ♂: A, antenna; B, leg 1, frontal view; C, leg 2, caudal view; D, leg 3; E, leg 11; F, anterior gonopods, caudal view; G, posterior gonopods, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393684" httpUri="https://zenodo.org/record/5393684/files/figure.png" pageId="29" pageNumber="446">Fig. 24B</figureCitation>
); each anterior gonopod plate with a high distomedial process (
<figureCitation id="A3F62A4BFFDCD525D3C2FB1DFD8EF93A" box="[460,555,1261,1287]" captionStart="FIG" captionStartId="30.[113,124,1286,1303]" captionTargetBox="[115,1207,204,1257]" captionTargetId="figure-23@30.[115,1207,204,1257]" captionTargetPageId="30" captionText="FIG. 24. — Glyphiulus paracostulifer n. sp., paratype ♂: A, antenna; B, leg 1, frontal view; C, leg 2, caudal view; D, leg 3; E, leg 11; F, anterior gonopods, caudal view; G, posterior gonopods, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393684" httpUri="https://zenodo.org/record/5393684/files/figure.png" pageId="29" pageNumber="446">Fig. 24F</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="6FB26646FFDDD524D27FF989FBADFA9C" ID-DOI="http://doi.org/10.5281/zenodo.5393680" ID-Zenodo-Dep="5393680" httpUri="https://zenodo.org/record/5393680/files/figure.png" pageId="28" pageNumber="445" startId="28.[113,124,1657,1674]" targetBox="[116,1205,215,1587]" targetPageId="28">
<paragraph id="3B7236CEFFDDD524D27FF989FBADFA9C" blockId="28.[113,1211,1657,1697]" pageId="28" pageNumber="445">
FIG. 22.—
<taxonomicName id="FCCD4D4DFFDDD524D2C9F989FE2AFAB7" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[199,399,1657,1674]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="28" pageNumber="445" phylum="Arthropoda" rank="species" species="paracostulifer" status="sp. nov.">
<emphasis id="09B9EADCFFDDD524D2C9F989FE2AFAB7" box="[199,399,1657,1674]" italics="true" pageId="28" pageNumber="445">Glyphiulus paracostulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFDDD524D39DF989FE66FAB7" box="[403,451,1657,1674]" pageId="28" pageNumber="445" rank="species">n.sp.</taxonomicNameLabel>
, paratype ♀:
<emphasis id="09B9EADCFFDDD524D025F989FD9DFAB7" bold="true" box="[555,568,1657,1674]" pageId="28" pageNumber="445">A</emphasis>
, anterior part of body,lateral view;
<emphasis id="09B9EADCFFDDD524D15EF989FCF8FAB7" bold="true" box="[848,861,1657,1674]" pageId="28" pageNumber="445">B</emphasis>
, same,dorsal view;
<emphasis id="09B9EADCFFDDD524D60FF989FBAAFAB7" bold="true" box="[1025,1039,1657,1674]" pageId="28" pageNumber="445">C</emphasis>
, midbody segments, lateral view;
<emphasis id="09B9EADCFFDDD524D2D7F960FF42FA9C" bold="true" box="[217,231,1680,1697]" pageId="28" pageNumber="445">D</emphasis>
, same, dorsal view;
<emphasis id="09B9EADCFFDDD524D39CF960FE3BFA9C" bold="true" box="[402,414,1680,1697]" pageId="28" pageNumber="445">E</emphasis>
, caudal body end, lateral view;
<emphasis id="09B9EADCFFDDD524D0A8F960FD14FA9C" bold="true" box="[678,689,1680,1697]" pageId="28" pageNumber="445">F</emphasis>
, same, ventral view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="6FB26646FFDCD525D27FFBAFFDF8F8BA" ID-DOI="http://doi.org/10.5281/zenodo.5393682" ID-Zenodo-Dep="5393682" httpUri="https://zenodo.org/record/5393682/files/figure.png" pageId="29" pageNumber="446" startId="29.[113,124,1119,1136]" targetBox="[117,1205,214,1085]" targetPageId="29">
<paragraph id="3B7236CEFFDCD525D27FFBAFFDF8F8BA" blockId="29.[113,1211,1119,1159]" pageId="29" pageNumber="446">
FIG. 23. —
<taxonomicName id="FCCD4D4DFFDCD525D2C3FBAFFE3EF84D" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[205,411,1119,1136]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="29" pageNumber="446" phylum="Arthropoda" rank="species" species="paracostulifer" status="sp. nov.">
<emphasis id="09B9EADCFFDCD525D2C3FBAFFE3EF84D" box="[205,411,1119,1136]" italics="true" pageId="29" pageNumber="446">Glyphiulus paracostulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFDCD525D3AEFBAFFE76F84D" box="[416,467,1119,1136]" pageId="29" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
, paratype ♀:
<emphasis id="09B9EADCFFDCD525D04FFBAFFDEBF84D" bold="true" box="[577,590,1119,1136]" pageId="29" pageNumber="446">A</emphasis>
, midbody segment section;
<emphasis id="09B9EADCFFDCD525D133FBAFFCEFF84D" bold="true" box="[829,842,1119,1136]" pageId="29" pageNumber="446">B</emphasis>
, midbody leg;
<emphasis id="09B9EADCFFDCD525D1C9FBAFFC70F84D" bold="true" box="[967,981,1119,1136]" pageId="29" pageNumber="446">C</emphasis>
, claw;
<emphasis id="09B9EADCFFDCD525D61FFBAFFBBAF84D" bold="true" box="[1041,1055,1119,1136]" pageId="29" pageNumber="446">D</emphasis>
, antennal sensilla. Scale bars: A, 0.5 mm; B, 0.2 mm; C, 0.1 mm; D, 0.05 mm.
</paragraph>
</caption>
<subSubSection id="73D76545FFDCD525D27FFADDFEC9F9F8" pageId="29" pageNumber="446" type="discussion">
<paragraph id="3B7236CEFFDCD525D27FFADDFF70F978" blockId="29.[113,646,1325,1478]" box="[113,213,1325,1351]" pageId="29" pageNumber="446">REMARKS</paragraph>
<paragraph id="3B7236CEFFDCD525D27FFABCFEC9F9F8" blockId="29.[113,646,1325,1478]" pageId="29" pageNumber="446">Based on several troglomorphic traits, such as the unpigmented body and ocelli, as well as the somewhat elongated antennae and legs, this species seems to be troglobitic.</paragraph>
</subSubSection>
</treatment>
</document>