treatments-xml/data/AD/5B/CC/AD5BCC59AE2AFFC7FF0DFDF85455FE7D.xml
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<document ID-DOI="10.11646/zootaxa.5278.3.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7906748" ID-ZooBank="8397A532-CFAB-4BCA-AF98-D76BE5079B04" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683531519212" checkinUser="plazi" docAuthor="Bitar, Murilo Victor Silva, Mendonça, Maria Thayane Da Silva &amp; Fernandes, José Antônio Marin" docDate="2023" docId="AD5BCC59AE2AFFC7FF0DFDF85455FE7D" docLanguage="en" docName="zootaxa.5278.3.9.pdf" docOrigin="Zootaxa 5278 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.3.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Edessa fuscopunctata Bitar &amp; Mendonça &amp; Fernandes 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="590" masterDocId="5162B421AE20FFCBFF9AFFBE5775FF88" masterDocTitle="Description of seven new species of Edessa Fabricius, 1803 from Central America (Heteroptera, Pentatomidae, Edessinae)" masterLastPageNumber="600" masterPageNumber="578" pageNumber="588" updateTime="1683532061405" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Description of seven new species of Edessa Fabricius, 1803 from Central America (Heteroptera, Pentatomidae, Edessinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bitar, Murilo Victor Silva</mods:namePart>
<mods:affiliation>murilobioufpa @ gmail. com; https: // orcid. org / 0000 - 0002 - 1109 - 4386</mods:affiliation>
<mods:nameIdentifier type="email">murilobioufpa@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mendonça, Maria Thayane Da Silva</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-6504-247X</mods:nameIdentifier>
<mods:affiliation>thaymendonca 4 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6504 - 247 X</mods:affiliation>
<mods:nameIdentifier type="email">thaymendonca4@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fernandes, José Antônio Marin</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-7450-5296</mods:nameIdentifier>
<mods:affiliation>joseamf @ ufpa. br; https: // orcid. org / 0000 - 0001 - 7450 - 5296</mods:affiliation>
<mods:nameIdentifier type="email">joseamf@ufpa.br</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5278</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>578</mods:start>
<mods:end>600</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.3.9</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5278.3.9</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7906748</mods:identifier>
<mods:identifier type="ZooBank">8397A532-CFAB-4BCA-AF98-D76BE5079B04</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7906774" ID-Zenodo-Dep="7906774" LSID="urn:lsid:plazi:treatment:AD5BCC59AE2AFFC7FF0DFDF85455FE7D" httpUri="http://treatment.plazi.org/id/AD5BCC59AE2AFFC7FF0DFDF85455FE7D" lastPageId="12" lastPageNumber="590" pageId="10" pageNumber="588">
<subSubSection box="[151,488,582,609]" pageId="10" pageNumber="588" type="nomenclature">
<paragraph blockId="10.[151,488,582,645]" box="[151,488,582,609]" pageId="10" pageNumber="588">
<heading bold="true" box="[151,488,582,609]" fontSize="11" level="1" pageId="10" pageNumber="588" reason="1">
<emphasis bold="true" box="[151,488,582,609]" pageId="10" pageNumber="588">
<taxonomicName authority="Bitar &amp; Mendonça &amp; Fernandes, 2023" authorityName="Bitar &amp; Mendonça &amp; Fernandes" authorityYear="2023" box="[151,414,582,609]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="588" phylum="Arthropoda" rank="species" species="fuscopunctata" status="sp. n.">
<emphasis bold="true" box="[151,414,582,609]" italics="true" pageId="10" pageNumber="588">Edessa fuscopunctata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[422,488,583,609]" pageId="10" pageNumber="588" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,403,618,645]" pageId="10" pageNumber="588" type="description">
<paragraph blockId="10.[151,488,582,645]" box="[151,403,618,645]" pageId="10" pageNumber="588">
(
<figureCitation box="[159,239,618,645]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Figs. 5</figureCitation>
;
<figureCitation box="[252,328,619,645]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">9 A, D</figureCitation>
;
<figureCitation box="[340,396,619,645]" captionStart="FIGURE 10" captionStartId="19.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-25@19.[203,1385,181,1919]" captionTargetPageId="19" captionText="FIGURE 10. AC. Distribution map of the species: A, Edessa bituberculata sp. n., Edessa fuscolimbata sp. n., Edessa fuscopunctatasp.n.;B,Edessabivenulatasp.n.,Edessaenargocelidasp.n.,Edessaholochloratasp.n.;C,Edessahelvoalatasp.n." figureDoi="http://doi.org/10.5281/zenodo.7906784" httpUri="https://zenodo.org/record/7906784/files/figure.png" pageId="10" pageNumber="588">10 A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[151,1373,691,717]" pageId="10" pageNumber="588" type="etymology">
<paragraph blockId="10.[151,1437,691,2013]" box="[151,1373,691,717]" pageId="10" pageNumber="588">
<emphasis bold="true" box="[151,286,691,717]" pageId="10" pageNumber="588">Etymology.</emphasis>
The name refers to the black color of the pronotal punctuation (L.
<emphasis box="[1035,1103,691,716]" italics="true" pageId="10" pageNumber="588">Fusco</emphasis>
- black;
<emphasis box="[1195,1291,691,716]" italics="true" pageId="10" pageNumber="588">punctum</emphasis>
- hole).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="588" type="materials_examined">
<paragraph blockId="10.[151,1437,691,2013]" pageId="10" pageNumber="588">
<emphasis bold="true" box="[199,430,726,752]" pageId="10" pageNumber="588">Material examined.</emphasis>
<materialsCitation collectingDate="1979-08-05" collectingDateMax="1979-08-11" collectingDateMin="1979-08-05" collectionCode="INBio" collectorName="D. H. Janzen" country="Costa Rica" location="Osa Peninsula" municipality="Corcovado National Park" pageId="10" pageNumber="588" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus box="[434,538,727,753]" pageId="10" pageNumber="588">Holotype</typeStatus>
male.
<emphasis bold="true" box="[607,776,726,752]" pageId="10" pageNumber="588">
<collectingCountry box="[607,776,726,752]" name="Costa Rica" pageId="10" pageNumber="588">COSTA RICA</collectingCountry>
</emphasis>
.
<emphasis bold="true" box="[787,933,727,753]" pageId="10" pageNumber="588">
<collectingRegion box="[787,928,727,753]" country="Costa Rica" name="Puntarenas" pageId="10" pageNumber="588">Puntarenas</collectingRegion>
:
</emphasis>
1♁,
<collectingMunicipality box="[983,1264,727,753]" pageId="10" pageNumber="588">Corcovado National Park</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:AD5BCC59AE2AFFC7FF0DFDF85455FE7D:202D2B94AE2AFFC1FB63FD6952EDFD79" box="[1273,1432,727,753]" country="Costa Rica" municipality="Corcovado National Park" name="Osa Peninsula" pageId="10" pageNumber="588" stateProvince="Puntarenas">Osa Peninsula</location>
,
<date box="[151,329,762,789]" pageId="10" pageNumber="588" value="1979-08-05" valueMax="1979-08-11" valueMin="1979-08-05">
<collectingDate box="[151,329,762,789]" pageId="10" pageNumber="588" value="1979-08-05" valueMax="1979-08-11" valueMin="1979-08-05">511/VIII/1979</collectingDate>
</date>
,
<collectorName box="[340,487,763,789]" pageId="10" pageNumber="588">D. H. Janzen</collectorName>
, INBIO CRI001 702800 (
<collectionCode box="[778,854,763,789]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="10" pageNumber="588">INBio</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="10.[151,1437,691,2013]" pageId="10" pageNumber="588">
<materialsCitation collectingDate="1980-03-13" collectingDateMax="1980-03-22" collectingDateMin="1980-03-13" collectionCode="INBio" collectorName="Corcovado Nat. Pk. &amp; D. H. Janzen &amp; W. Hallwachs" country="Costa Rica" location="Osa Peninsula" municipality="Sirena" pageId="10" pageNumber="588" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
<typeStatus box="[199,312,799,825]" pageId="10" pageNumber="588">Paratypes</typeStatus>
.
<emphasis bold="true" box="[321,492,798,824]" pageId="10" pageNumber="588">
<collectingCountry box="[321,492,798,824]" name="Costa Rica" pageId="10" pageNumber="588">COSTA RICA</collectingCountry>
</emphasis>
.
<emphasis bold="true" box="[505,651,799,825]" pageId="10" pageNumber="588">
<collectingRegion box="[505,646,799,825]" country="Costa Rica" name="Puntarenas" pageId="10" pageNumber="588">Puntarenas</collectingRegion>
:
</emphasis>
1♁,
<collectingMunicipality box="[705,779,799,825]" pageId="10" pageNumber="588">Sirena</collectingMunicipality>
,
<collectorName box="[789,1011,799,825]" pageId="10" pageNumber="588">Corcovado Nat. Pk.</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:AD5BCC59AE2AFFC7FF0DFDF85455FE7D:202D2B94AE2AFFC1FC67FCA153EAFCB1" box="[1021,1183,799,825]" country="Costa Rica" municipality="Sirena" name="Osa Peninsula" pageId="10" pageNumber="588" stateProvince="Puntarenas">Osa Peninsula</location>
,
<date box="[1193,1366,799,825]" pageId="10" pageNumber="588" value="1980-03-13" valueMax="1980-03-22" valueMin="1980-03-13">
<collectingDate box="[1193,1366,799,825]" pageId="10" pageNumber="588" value="1980-03-13" valueMax="1980-03-22" valueMin="1980-03-13">1322/III/1980</collectingDate>
</date>
,
<collectorName pageId="10" pageNumber="588">D. H. Janzen</collectorName>
&amp;
<collectorName box="[261,420,835,861]" pageId="10" pageNumber="588">W. Hallwachs</collectorName>
, INBIO CRI001 715378 (
<collectionCode box="[708,784,835,861]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="10" pageNumber="588">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1979-03-15" collectingDateMax="1979-03-22" collectingDateMin="1979-03-15" collectionCode="INBio" collectorName="Corcovado Nat. Pk. Osa Penin &amp; D. H. Janzen" country="Costa Rica" pageId="10" pageNumber="588" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
1♁,
<collectorName box="[851,1194,835,861]" pageId="10" pageNumber="588">Corcovado Nat. Pk. Osa Penin</collectorName>
,
<date box="[1204,1376,834,861]" pageId="10" pageNumber="588" value="1979-03-15" valueMax="1979-03-22" valueMin="1979-03-15">
<collectingDate box="[1204,1376,834,861]" pageId="10" pageNumber="588" value="1979-03-15" valueMax="1979-03-22" valueMin="1979-03-15">1522/III/1979</collectingDate>
</date>
, coll.
<collectorName box="[151,298,871,897]" pageId="10" pageNumber="588">D. H. Janzen</collectorName>
, INBIO CRI001 702253 (
<collectionCode box="[589,665,871,897]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="10" pageNumber="588">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1995-04-05" collectingDateMax="1995-04-24" collectingDateMin="1995-04-05" collectionCode="INBio" collectorName="B. Gamboa" country="Costa Rica" elevation="51" pageId="10" pageNumber="588" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount box="[684,719,871,897]" pageId="10" pageNumber="588" type="female">1♀</specimenCount>
, Est. Sirena, ACOSA.
<quantity box="[975,1070,871,897]" metricMagnitude="1" metricUnit="m" metricValue="5.05" metricValueMax="10.0" metricValueMin="0.1" pageId="10" pageNumber="588" unit="m" value="50.5" valueMax="100.0" valueMin="1.0">
<elevation box="[975,1070,871,897]" metricMagnitude="1" metricUnit="m" metricValue="5.05" metricValueMax="10.0" metricValueMin="0.1" pageId="10" pageNumber="588" unit="m" value="50.5" valueMax="100.0" valueMin="1.0">1100m</elevation>
</quantity>
,
<date box="[1081,1240,870,897]" pageId="10" pageNumber="588" value="1995-04-05" valueMax="1995-04-24" valueMin="1995-04-05">
<collectingDate box="[1081,1240,870,897]" pageId="10" pageNumber="588" value="1995-04-05" valueMax="1995-04-24" valueMin="1995-04-05">524/IV/1995</collectingDate>
</date>
,
<collectorName box="[1252,1382,871,897]" pageId="10" pageNumber="588">B. Gamboa</collectorName>
, L N 270500 507900#4738, INBIO CRI002 188039 (
<collectionCode box="[690,766,907,933]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="10" pageNumber="588">INBio</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="588" type="description">
<paragraph blockId="10.[151,1437,691,2013]" pageId="10" pageNumber="588">
<emphasis bold="true" box="[199,374,943,969]" pageId="10" pageNumber="588">Measurements</emphasis>
(n= 4): antennomeres length: 1st:
<quantity box="[768,927,942,969]" metricMagnitude="-4" metricUnit="m" metricValue="5.9" metricValueMax="6.7" metricValueMin="5.1" pageId="10" pageNumber="588" unit="mm" value="0.59" valueMax="0.67" valueMin="0.51">0.510.67mm</quantity>
; 2nd:
<quantity box="[997,1157,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" metricValueMax="1.14" metricValueMin="1.02" pageId="10" pageNumber="588" unit="mm" value="1.08" valueMax="1.14" valueMin="1.02">1.021.14mm</quantity>
; 3rd:
<quantity box="[1221,1381,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="2.395" metricValueMax="2.89" metricValueMin="1.9" pageId="10" pageNumber="588" unit="mm" value="2.395" valueMax="2.89" valueMin="1.9">1.902.89mm</quantity>
; 4th:
<quantity box="[151,311,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.525" metricValueMax="3.57" metricValueMin="3.48" pageId="10" pageNumber="588" unit="mm" value="3.525" valueMax="3.57" valueMin="3.48">3.483.57mm</quantity>
; 5th:
<quantity box="[369,465,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.37" pageId="10" pageNumber="588" unit="mm" value="3.37">3.37mm</quantity>
; head length:
<quantity box="[617,777,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="1.415" metricValueMax="1.49" metricValueMin="1.34" pageId="10" pageNumber="588" unit="mm" value="1.415" valueMax="1.49" valueMin="1.34">1.341.49mm</quantity>
; head width:
<quantity box="[922,1081,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="2.84" metricValueMax="2.85" metricValueMin="2.83" pageId="10" pageNumber="588" unit="mm" value="2.84" valueMax="2.85" valueMin="2.83">2.832.85mm</quantity>
; pronotal length:
<quantity box="[1273,1432,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="2.89" metricValueMax="2.99" metricValueMin="2.79" pageId="10" pageNumber="588" unit="mm" value="2.89" valueMax="2.99" valueMin="2.79">2.792.99mm</quantity>
; pronotal width:
<quantity box="[331,490,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="7.074999999999999" metricValueMax="7.27" metricValueMin="6.88" pageId="10" pageNumber="588" unit="mm" value="7.074999999999999" valueMax="7.27" valueMin="6.88">6.887.27mm</quantity>
; scutellum length:
<quantity box="[703,862,1014,1041]" metricMagnitude="-3" metricUnit="m" metricValue="5.755" metricValueMax="5.96" metricValueMin="5.55" pageId="10" pageNumber="588" unit="mm" value="5.755" valueMax="5.96" valueMin="5.55">5.555.96mm</quantity>
; scutellum width:
<quantity box="[1068,1228,1014,1041]" metricMagnitude="-3" metricUnit="m" metricValue="4.325" metricValueMax="4.5" metricValueMin="4.15" pageId="10" pageNumber="588" unit="mm" value="4.325" valueMax="4.5" valueMin="4.15">4.154.50mm</quantity>
; abdominal width:
<quantity box="[151,311,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="7.23" metricValueMin="6.87" pageId="10" pageNumber="588" unit="mm" value="7.05" valueMax="7.23" valueMin="6.87">6.877.23mm</quantity>
; total length:
<quantity box="[463,650,1051,1077]" metricMagnitude="-2" metricUnit="m" metricValue="1.3335" metricValueMax="1.3679999999999999" metricValueMin="1.299" pageId="10" pageNumber="588" unit="mm" value="13.335" valueMax="13.68" valueMin="12.99">12.9913.68mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="588" type="diagnosis">
<paragraph blockId="10.[151,1437,691,2013]" pageId="10" pageNumber="588">
<emphasis bold="true" box="[199,324,1086,1112]" pageId="10" pageNumber="588">Diagnosis:</emphasis>
body dorsally green except variegated brown corium (
<figureCitation box="[960,1061,1087,1113]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Antennae brown, basal portion of 5th antennomeres whitish (
<figureCitation box="[503,608,1123,1149]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Anterolateral margin of pronotum with narrow yellowish stripe (
<figureCitation captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Pronotum with punctures black, coarse, covering most of pronotum except on cicatrices, anterior margin and anterolateral margins (
<figureCitation box="[459,561,1195,1221]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Black minute longitudinal callosity adjacent to humeral angles (
<figureCitation box="[1322,1424,1195,1221]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Scutellum with transverse diffuse brown stripe on midposterior region (
<figureCitation box="[968,1067,1231,1257]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Corium with diffuse yellowish and reddish shade, black longitudinal stripe on basal portion of lateral margin (
<figureCitation box="[1069,1170,1267,1293]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Connexival segments with yellowish spots (
<figureCitation box="[399,497,1303,1329]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Last connexival segment mostly black (
<figureCitation box="[967,1064,1303,1329]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Ventral surface: yellowish with thoracic dark lines restricted to propleura and abdominal dark lines narrow and incomplete (
<figureCitation box="[1218,1321,1339,1365]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 D</figureCitation>
). Anterior bifurcation of the metasternal process with arms narrow, apices acuminated (
<figureCitation box="[1019,1119,1375,1401]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 D</figureCitation>
). Male genitalia: pygophore subrectangular (
<figureCitation box="[333,464,1410,1437]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Fig. 5 A, B</figureCitation>
). Dorsal rim freely projected over slightly developed posterolateral angles (
<figureCitation box="[1325,1424,1410,1437]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Fig. 5 A</figureCitation>
). Superior process of the genital cup arrowhead-like, slightly bent, medially excavated (
<figureCitation box="[1114,1244,1446,1473]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Fig. 5 C, D</figureCitation>
). Parameres with anterior lobe triangular and developed (
<figureCitation box="[599,731,1482,1509]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Fig. 5 C, D</figureCitation>
); dorsal margin deeply excavated (
<figureCitation box="[1127,1258,1482,1509]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Fig. 5 C, D</figureCitation>
); posterior lobe short, digitiform and straight (
<figureCitation box="[497,634,1518,1545]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Fig. 5 C, D</figureCitation>
). Proctiger slightly excavated laterally (
<figureCitation box="[1091,1227,1518,1545]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Fig. 5 C, D</figureCitation>
). Ventral rim with median notch subrectangular; expansions developed and triangular (
<figureCitation box="[911,1004,1554,1581]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Fig. 5 B</figureCitation>
). Female genitalia (
<figureCitation box="[1225,1319,1554,1581]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Fig. 5 E</figureCitation>
): valvifers VIII punctured medially; medially largely excavated.
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="590" pageId="10" pageNumber="588" type="description">
<paragraph blockId="10.[151,1437,691,2013]" lastBlockId="12.[151,1437,150,501]" lastPageId="12" lastPageNumber="590" pageId="10" pageNumber="588">
<emphasis bold="true" box="[199,422,1626,1653]" pageId="10" pageNumber="588">Description: head:</emphasis>
clypeus and jugae, slightly ridged. Bucculae subtriangular, harboring first labial segment almost completely.
<emphasis bold="true" box="[370,468,1663,1689]" pageId="10" pageNumber="588">Thorax:</emphasis>
dorsal surface: anterior margin of pronotum with subtle intermittent row of punctures (
<figureCitation box="[159,259,1699,1725]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Humeral angle with small black stripe at the apex, short, about as long as the width of an eye (
<figureCitation captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Scutellum very acute, punctures coarse, grouped much tighter on brown stripe and adjacent areas; distal part unpunctured (
<figureCitation box="[305,398,1771,1797]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Membrane transparent, slightly dark (
<figureCitation box="[832,925,1771,1797]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
). Ventral surface: prothorax with black, dense, irregularly distributed punctures (
<figureCitation box="[529,628,1807,1833]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 D</figureCitation>
). Mesothorax with few minute, shallow punctures (
<figureCitation box="[1208,1307,1807,1833]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 D</figureCitation>
). Peritreme ruga-like, reaching 2/3 of the distance between ostiole of the scent gland and lateral margin of metapleuron (
<figureCitation box="[1330,1424,1843,1869]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 D</figureCitation>
). Metasternal process with anterior arms slightly divergent, anterior bifurcation receiving only fourth rostral segment (
<figureCitation box="[159,255,1915,1941]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 D</figureCitation>
).
<emphasis bold="true" box="[274,400,1914,1940]" pageId="10" pageNumber="588">Abdomen:</emphasis>
dorsal surface: posterolateral angles of connexival segments with distal black spot (
<figureCitation box="[1330,1424,1915,1941]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="10" pageNumber="588">Fig. 9 A</figureCitation>
).
<emphasis bold="true" box="[151,330,1950,1977]" pageId="10" pageNumber="588">Male genitalia:</emphasis>
dorsal rim light-brown, slightly sinuous, broadly excavated (
<figureCitation box="[1004,1097,1950,1977]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Fig. 5 A</figureCitation>
). Posterolateral angles slightly developed, gently excavated on inner face (
<figureCitation box="[622,712,1986,2013]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Fig. 5A</figureCitation>
). Ventral rim with expansions short and slightly swollen (
<figureCitation box="[1333,1422,1986,2013]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="10" pageNumber="588">Fig. 5 B</figureCitation>
).
<emphasis bold="true" box="[151,347,150,177]" pageId="12" pageNumber="590">Female genitalia</emphasis>
(
<figureCitation box="[361,450,150,177]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="12" pageNumber="590">Fig. 5 E</figureCitation>
): valvifers VIII slightly convex; distal margin slightly arched; median excavation widely exposing valvulae IX. Laterotergites VIII sharp distally, outer lateral margin almost straight; free distal spinose part about 1/5 of its length. Valvifers IX smooth, trapezoidal. Laterotergites IX acute distally, barely surpassing sclerite uniting laterotergites VIII.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906764" ID-Zenodo-Dep="7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="11" pageNumber="589" startId="11.[152,255,1943,1967]" targetBox="[203,1385,179,1917]" targetPageId="11" targetType="figure">
<paragraph blockId="11.[151,1436,1943,2004]" pageId="11" pageNumber="589">
<emphasis bold="true" box="[152,286,1943,1968]" pageId="11" pageNumber="589">FIGURE 5.</emphasis>
AE.
<taxonomicName authority="Bitar &amp; Mendonça &amp; Fernandes, 2023" authorityName="Bitar &amp; Mendonça &amp; Fernandes" authorityYear="2023" box="[362,588,1944,1968]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="589" phylum="Arthropoda" rank="species" species="fuscopunctata" status="sp. n.">
<emphasis box="[362,588,1944,1968]" italics="true" pageId="11" pageNumber="589">Edessa fuscopunctata</emphasis>
</taxonomicName>
<emphasis bold="true" box="[600,663,1944,1968]" pageId="11" pageNumber="589">
<taxonomicNameLabel box="[600,663,1944,1968]" pageId="11" pageNumber="589" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm.
</paragraph>
</caption>
<subSubSection pageId="12" pageNumber="590" type="diagnosis">
<paragraph blockId="12.[151,1437,150,501]" pageId="12" pageNumber="590">
<emphasis bold="true" box="[199,465,294,320]" pageId="12" pageNumber="590">Differential diagnosis:</emphasis>
<taxonomicName authority="Bitar &amp; Mendonça &amp; Fernandes, 2023" authorityName="Bitar &amp; Mendonça &amp; Fernandes" authorityYear="2023" box="[473,717,295,321]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="590" phylum="Arthropoda" rank="species" species="fuscopunctata" status="sp. n.">
<emphasis box="[473,717,295,321]" italics="true" pageId="12" pageNumber="590">Edessa fuscopunctata</emphasis>
</taxonomicName>
<emphasis bold="true" box="[725,789,295,321]" pageId="12" pageNumber="590">
<taxonomicNameLabel box="[725,789,295,321]" pageId="12" pageNumber="590" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is more similar to
<taxonomicName authority="Bitar &amp; Mendonça &amp; Fernandes, 2023" authorityName="Bitar &amp; Mendonça &amp; Fernandes" authorityYear="2023" box="[1003,1180,295,321]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="590" phylum="Arthropoda" rank="species" species="fuscolimbata" status="sp. n.">
<emphasis box="[1003,1180,295,321]" italics="true" pageId="12" pageNumber="590">E. fuscolimbata</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1188,1252,295,321]" pageId="12" pageNumber="590">
<taxonomicNameLabel box="[1188,1252,295,321]" pageId="12" pageNumber="590" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(see differential diagnosis of
<taxonomicName authority="Bitar &amp; Mendonça &amp; Fernandes, 2023" authorityName="Bitar &amp; Mendonça &amp; Fernandes" authorityYear="2023" box="[298,477,331,357]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="590" phylum="Arthropoda" rank="species" species="fuscolimbata" status="sp. n.">
<emphasis box="[298,477,331,357]" italics="true" pageId="12" pageNumber="590">E. fuscolimbata</emphasis>
</taxonomicName>
<emphasis bold="true" box="[485,551,331,357]" pageId="12" pageNumber="590">
<taxonomicNameLabel box="[485,551,331,357]" pageId="12" pageNumber="590" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
). But
<taxonomicName authority="Bitar &amp; Mendonça &amp; Fernandes, 2023" authorityName="Bitar &amp; Mendonça &amp; Fernandes" authorityYear="2023" box="[625,871,331,357]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="590" phylum="Arthropoda" rank="species" species="fuscopunctata" status="sp. n.">
<emphasis box="[625,871,331,357]" italics="true" pageId="12" pageNumber="590">Edessa fuscopunctata</emphasis>
</taxonomicName>
<emphasis bold="true" box="[880,945,331,357]" pageId="12" pageNumber="590">
<taxonomicNameLabel box="[880,945,331,357]" pageId="12" pageNumber="590" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be separated from
<taxonomicName authority="Bitar &amp; Mendonça &amp; Fernandes, 2023" authorityName="Bitar &amp; Mendonça &amp; Fernandes" authorityYear="2023" box="[1216,1394,331,357]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="590" phylum="Arthropoda" rank="species" species="fuscolimbata" status="sp. n.">
<emphasis box="[1216,1394,331,357]" italics="true" pageId="12" pageNumber="590">E. fuscolimbata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="12" pageNumber="590">
<taxonomicNameLabel pageId="12" pageNumber="590" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the brown midposterior stripe on scutellum (
<figureCitation box="[736,847,367,393]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="12" pageNumber="590">Figs. 9 A</figureCitation>
;
<figureCitation box="[862,910,367,393]" captionStart="FIGURE 8" captionStartId="17.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-25@17.[203,1385,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. AH. Dorsal and ventral surface: AD, Edessa bituberculata sp. n.; BE, Edessa bivenulata sp. n.; CF, Edessa enargocelida sp. n.; GH, Edessa fuscolimbata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906776" httpUri="https://zenodo.org/record/7906776/files/figure.png" pageId="12" pageNumber="590">8 G</figureCitation>
); humeral angles very short (
<figureCitation box="[1249,1360,367,393]" captionStart="FIGURE 9" captionStartId="18.[152,255,1757,1781]" captionTargetBox="[151,1436,181,1733]" captionTargetId="figure-20@18.[151,1436,181,1733]" captionTargetPageId="18" captionText="FIGURE 9. AF. Dorsal and ventral surface: AD, Edessa fuscopunctata sp. n.; BE, Edessa helvoalata sp. n.; CF, Edessa holochlorata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906780" httpUri="https://zenodo.org/record/7906780/files/figure.png" pageId="12" pageNumber="590">Figs. 9 A</figureCitation>
;
<figureCitation box="[1375,1423,367,393]" captionStart="FIGURE 8" captionStartId="17.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-25@17.[203,1385,181,1919]" captionTargetPageId="17" captionText="FIGURE 8. AH. Dorsal and ventral surface: AD, Edessa bituberculata sp. n.; BE, Edessa bivenulata sp. n.; CF, Edessa enargocelida sp. n.; GH, Edessa fuscolimbata sp. n. Scale 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906776" httpUri="https://zenodo.org/record/7906776/files/figure.png" pageId="12" pageNumber="590">8 G</figureCitation>
); pygophore with genital cup process arrowhead-like (
<figureCitation box="[756,903,402,429]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="12" pageNumber="590">Figs. 5 C, D</figureCitation>
;
<figureCitation box="[917,999,403,429]" captionStart="FIGURE 4" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[186,1401,179,1917]" captionTargetId="figure-25@9.[186,1401,179,1917]" captionTargetPageId="9" captionText="FIGURE 4.AD.Edessa fuscolimbata sp. n.:AD, pygophore in dorsal, ventral,posterior and posterolateral views, respectively. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906760" httpUri="https://zenodo.org/record/7906760/files/figure.png" pageId="12" pageNumber="590">4 C, D</figureCitation>
), and dorsal rim projected freely over humeral angles (
<figureCitation box="[337,442,438,465]" captionStart="FIGURE 5" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[203,1385,179,1917]" captionTargetId="figure-25@11.[203,1385,179,1917]" captionTargetPageId="11" captionText="FIGURE 5. AE. Edessa fuscopunctata sp. n.: AD, pygophore in dorsal, ventral, posterior and posterolateral views, respectively; E, genital plates. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906764" httpUri="https://zenodo.org/record/7906764/files/figure.png" pageId="12" pageNumber="590">Figs. 5 A</figureCitation>
;
<figureCitation box="[454,498,439,465]" captionStart="FIGURE 4" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[186,1401,179,1917]" captionTargetId="figure-25@9.[186,1401,179,1917]" captionTargetPageId="9" captionText="FIGURE 4.AD.Edessa fuscolimbata sp. n.:AD, pygophore in dorsal, ventral,posterior and posterolateral views, respectively. Scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906760" httpUri="https://zenodo.org/record/7906760/files/figure.png" pageId="12" pageNumber="590">4 A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[199,800,474,501]" pageId="12" pageNumber="590" type="distribution">
<paragraph blockId="12.[151,1437,150,501]" box="[199,800,474,501]" pageId="12" pageNumber="590">
<emphasis bold="true" box="[199,345,474,500]" pageId="12" pageNumber="590">Distribution</emphasis>
(
<figureCitation box="[361,469,475,501]" captionStart="FIGURE 10" captionStartId="19.[152,255,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-25@19.[203,1385,181,1919]" captionTargetPageId="19" captionText="FIGURE 10. AC. Distribution map of the species: A, Edessa bituberculata sp. n., Edessa fuscolimbata sp. n., Edessa fuscopunctatasp.n.;B,Edessabivenulatasp.n.,Edessaenargocelidasp.n.,Edessaholochloratasp.n.;C,Edessahelvoalatasp.n." figureDoi="http://doi.org/10.5281/zenodo.7906784" httpUri="https://zenodo.org/record/7906784/files/figure.png" pageId="12" pageNumber="590">Fig. 10 A</figureCitation>
):
<collectingCountry box="[492,658,475,501]" name="Costa Rica" pageId="12" pageNumber="590">COSTA RICA</collectingCountry>
:
<collectingRegion box="[669,796,475,501]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="590">Puntarenas</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>