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<document id="BE79FB4D78CEFA5401461F4F7B2B89B9" ID-CLB-Dataset="173358" ID-DOI="10.11646/zootaxa.5185.1.1" ID-GBIF-Dataset="500935fc-fd0d-4cd1-b994-390f35fddadb" ID-ISSN="1175-5326" ID-Zenodo-Dep="7073826" ID-ZooBank="D34A7F04-8EAD-4441-A859-CFD79F7740D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1663050621745" checkinUser="plazi" docAuthor="Pisanty, Gideon, Scheuchl, Erwin, Martin, Teresa, Cardinal, Sophie &amp; Wood, Thomas James" docDate="2022" docId="9C1E87C4C737FF82FF0BFA0CFEF4B3C0" docLanguage="en" docName="zootaxa.5185.1.1.pdf" docOrigin="Zootaxa 5185 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5185.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Andrena (incertae sedis) hulae Pisanty &amp; Scheuchl &amp; Martin &amp; Cardinal &amp; Wood 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="88" masterDocId="6027FFBCC761FFD5FF9CFFB2FF84B57C" masterDocTitle="Twenty-five new species of mining bees (Hymenoptera: Andrenidae: Andrena) from Israel and the Levant" masterLastPageNumber="109" masterPageNumber="1" pageNumber="87" updateTime="1699379981322" updateUser="ExternalLinkService">
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<mods:title id="94CAD9CBC9B309B91CD030791E5717AA">Twenty-five new species of mining bees (Hymenoptera: Andrenidae: Andrena) from Israel and the Levant</mods:title>
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<mods:namePart id="32EA986EB357B27C5B1F91A51ADB5EF3">Pisanty, Gideon</mods:namePart>
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<mods:namePart id="9CB7EE71C7B65BDF308DA5B3C8080060">Scheuchl, Erwin</mods:namePart>
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<mods:namePart id="CCA384064B348C41BD8A4E60C21BA2C4">Martin, Teresa</mods:namePart>
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<mods:namePart id="CF973CF0660D13112CC6484EE8230DA2">Wood, Thomas James</mods:namePart>
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<treatment id="9C1E87C4C737FF82FF0BFA0CFEF4B3C0" ID-DOI="http://doi.org/10.5281/zenodo.7255259" ID-GBIF-Taxon="203591847" ID-Zenodo-Dep="7255259" LSID="urn:lsid:plazi:treatment:9C1E87C4C737FF82FF0BFA0CFEF4B3C0" httpUri="http://treatment.plazi.org/id/9C1E87C4C737FF82FF0BFA0CFEF4B3C0" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87">
<subSubSection id="5CAD6559C737FF83FF0BFA0CFD51B0A5" box="[151,725,1470,1497]" pageId="86" pageNumber="87" type="nomenclature">
<paragraph id="140836D2C737FF83FF0BFA0CFD51B0A5" blockId="86.[151,725,1470,1533]" box="[151,725,1470,1497]" pageId="86" pageNumber="87">
<heading id="4F4081BEC737FF83FF0BFA0CFD51B0A5" bold="true" box="[151,725,1470,1497]" fontSize="11" level="2" pageId="86" pageNumber="87" reason="4">
<emphasis id="26C3EAC0C737FF83FF0BFA0CFD51B0A5" bold="true" box="[151,725,1470,1497]" pageId="86" pageNumber="87">
<taxonomicName id="D3B74D51C737FF83FF0BFA0CFDF4B0A4" ID-CoL="DRTC" authority="Pisanty" authorityName="Pisanty &amp; Scheuchl &amp; Martin &amp; Cardinal &amp; Wood" authorityYear="2022" box="[151,624,1470,1497]" class="Insecta" family="Andrenidae" kingdom="Animalia" order="Hymenoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hulae" status="sp. nov.">
<emphasis id="26C3EAC0C737FF83FF0BFA0CFE85B0A5" bold="true" box="[151,257,1470,1497]" italics="true" pageId="86" pageNumber="87">Andrena</emphasis>
(
<emphasis id="26C3EAC0C737FF83FE8EFA0DFE32B0A5" bold="true" box="[274,438,1470,1497]" italics="true" pageId="86" pageNumber="87">incertae sedis</emphasis>
)
<emphasis id="26C3EAC0C737FF83FE5BFA0CFD8FB0A5" bold="true" box="[455,523,1470,1497]" italics="true" pageId="86" pageNumber="87">hulae</emphasis>
Pisanty
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<taxonomicNameLabel id="3DF057BBC737FF83FDEBFA0DFD51B0A5" box="[631,725,1471,1497]" pageId="86" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="5CAD6559C737FF82FF0BFA51FE0CB6D8" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87" type="description">
<paragraph id="140836D2C737FF83FF0BFA51FECEB081" blockId="86.[151,725,1470,1533]" box="[151,330,1507,1533]" pageId="86" pageNumber="87">
(
<figureCitation id="8C8C2A57C737FF83FF03FA51FEC7B081" box="[159,323,1507,1533]" captionStart-0="FIGURES 204 212" captionStart-1="FIGURES 213221" captionStartId-0="85.[151,264,1943,1967]" captionStartId-1="88.[151,264,1943,1967]" captionTargetBox-0="[222,1367,181,1916]" captionTargetBox-1="[220,1369,181,1919]" captionTargetId-0="figure-18@85.[202,1386,181,1920]" captionTargetId-1="figure-28@88.[204,1383,181,1920]" captionTargetPageId-0="85" captionTargetPageId-1="88" captionText-0="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." captionText-1="FIGURES 213221. Andrena (incertae sedis) hulae sp. nov. 213. female metasoma. Andrena (incertae sedis) inusitata sp. nov. 214. male habitus, 215. male head, 216. male head and mesosoma, 217. male lateral part of propodeum, 218. male metasoma, 219. male genitalia, 220. male eighth sternum, ventral view, 221. male eighth sternum, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.7073884" figureDoi-1="http://doi.org/10.5281/zenodo.7073888" httpUri-0="https://zenodo.org/record/7073884/files/figure.png" httpUri-1="https://zenodo.org/record/7073888/files/figure.png" pageId="86" pageNumber="87">Figs. 210213</figureCitation>
)
</paragraph>
<paragraph id="140836D2C737FF83FF0BF999FEE9B339" blockId="86.[151,1437,1579,2037]" box="[151,365,1579,1605]" pageId="86" pageNumber="87">
<emphasis id="26C3EAC0C737FF83FF0BF999FF6AB339" bold="true" box="[151,238,1579,1605]" pageId="86" pageNumber="87">Female</emphasis>
(
<figureCitation id="8C8C2A57C737FF83FF62F999FEE5B339" box="[254,353,1579,1605]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="86" pageNumber="87">Fig. 210</figureCitation>
).
</paragraph>
<paragraph id="140836D2C737FF83FF5BF9FDFE41B315" blockId="86.[151,1437,1579,2037]" box="[199,453,1614,1641]" pageId="86" pageNumber="87">
<emphasis id="26C3EAC0C737FF83FF5BF9FDFEE4B315" bold="true" box="[199,352,1615,1641]" pageId="86" pageNumber="87">Body length:</emphasis>
<quantity id="D34F9B37C737FF83FEFBF9FCFE44B315" box="[359,448,1614,1641]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="86" pageNumber="87" unit="mm" value="8.5">8.5 mm</quantity>
.
</paragraph>
<paragraph id="140836D2C737FF83FF5BF9C0FB55B3A9" blockId="86.[151,1437,1579,2037]" pageId="86" pageNumber="87">
<emphasis id="26C3EAC0C737FF83FF5BF9C0FEA4B3F0" bold="true" box="[199,288,1650,1676]" pageId="86" pageNumber="87">Colour.</emphasis>
Head and mesosoma dark brown to black (
<figureCitation id="8C8C2A57C737FF83FC9CF9C1FC22B3F1" box="[768,934,1651,1677]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="86" pageNumber="87">Figs. 210212</figureCitation>
). Anterior side of flagellomeres 310 orange. Legs and metasoma brown (
<figureCitation id="8C8C2A57C737FF83FE4BF925FDC0B3CD" box="[471,580,1687,1713]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="86" pageNumber="87">Figs. 210</figureCitation>
,
<figureCitation id="8C8C2A57C737FF83FDCCF925FDFAB3CD" box="[592,638,1687,1713]" captionStart="FIGURES 213221" captionStartId="88.[151,264,1943,1967]" captionTargetBox="[220,1369,181,1919]" captionTargetId="figure-28@88.[204,1383,181,1920]" captionTargetPageId="88" captionText="FIGURES 213221. Andrena (incertae sedis) hulae sp. nov. 213. female metasoma. Andrena (incertae sedis) inusitata sp. nov. 214. male habitus, 215. male head, 216. male head and mesosoma, 217. male lateral part of propodeum, 218. male metasoma, 219. male genitalia, 220. male eighth sternum, ventral view, 221. male eighth sternum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7073888" httpUri="https://zenodo.org/record/7073888/files/figure.png" pageId="86" pageNumber="87">213</figureCitation>
). Last tarsal segment orange. Wings hyaline, veins brown, stigma dark orange to brown (
<figureCitation id="8C8C2A57C737FF83FEC2F909FE45B3A9" box="[350,449,1723,1749]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="86" pageNumber="87">Fig. 210</figureCitation>
). Tergal marginal zones reddish basally, yellowish apically (
<figureCitation id="8C8C2A57C737FF83FBFEF909FB41B3A9" box="[1122,1221,1723,1749]" captionStart="FIGURES 213221" captionStartId="88.[151,264,1943,1967]" captionTargetBox="[220,1369,181,1919]" captionTargetId="figure-28@88.[204,1383,181,1920]" captionTargetPageId="88" captionText="FIGURES 213221. Andrena (incertae sedis) hulae sp. nov. 213. female metasoma. Andrena (incertae sedis) inusitata sp. nov. 214. male habitus, 215. male head, 216. male head and mesosoma, 217. male lateral part of propodeum, 218. male metasoma, 219. male genitalia, 220. male eighth sternum, ventral view, 221. male eighth sternum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7073888" httpUri="https://zenodo.org/record/7073888/files/figure.png" pageId="86" pageNumber="87">Fig. 213</figureCitation>
).
</paragraph>
<paragraph id="140836D2C737FF83FF5BF96DFB96B289" blockId="86.[151,1437,1579,2037]" pageId="86" pageNumber="87">
<emphasis id="26C3EAC0C737FF83FF5BF96DFED3B385" bold="true" box="[199,343,1759,1785]" pageId="86" pageNumber="87">Pubescence.</emphasis>
Clypeus with short and thin whitish hair. Paraocular area with medium-lengthed white hair. Scape with short white hair. Frons with short white hair. Foveae with short whitish hair. Vertex with medium yellowishwhite hair (
<figureCitation id="8C8C2A57C737FF83FE89F895FE3DB23D" box="[277,441,1831,1857]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="86" pageNumber="87">Figs. 211212</figureCitation>
). Genal area with short to medium white hair. Mesonotum, scutellum and metanotum with short to medium yellowish-white hair, mostly on periphery (
<figureCitation id="8C8C2A57C737FF83FCDEF8F9FC6EB219" box="[834,1002,1867,1893]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="86" pageNumber="87">Figs. 210, 212</figureCitation>
). Mesepisternum with long white hair. Propodeal corbicula incomplete, posterodorsal fringe with long white plumose hairs, corbicular surface with medium to long white plumose hairs. Leg hair white to golden. Flocculus complete, white. Tibial scopal hairs simple, white (
<figureCitation id="8C8C2A57C737FF83FF78F805FEC3B2AD" box="[228,327,1975,2001]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="86" pageNumber="87">Fig. 210</figureCitation>
). Tergal discs almost hairless medially, laterally with short whitish hairs. Tergal marginal zones 24 with broadly interrupted bands of white hair. Terminal fringe golden (
<figureCitation id="8C8C2A57C737FF83FC3FF869FB81B289" box="[931,1029,2011,2037]" captionStart="FIGURES 213221" captionStartId="88.[151,264,1943,1967]" captionTargetBox="[220,1369,181,1919]" captionTargetId="figure-28@88.[204,1383,181,1920]" captionTargetPageId="88" captionText="FIGURES 213221. Andrena (incertae sedis) hulae sp. nov. 213. female metasoma. Andrena (incertae sedis) inusitata sp. nov. 214. male habitus, 215. male head, 216. male head and mesosoma, 217. male lateral part of propodeum, 218. male metasoma, 219. male genitalia, 220. male eighth sternum, ventral view, 221. male eighth sternum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7073888" httpUri="https://zenodo.org/record/7073888/files/figure.png" pageId="86" pageNumber="87">Fig. 213</figureCitation>
).
</paragraph>
<paragraph id="140836D2C736FF82FF5BFF24FCAEB4D0" blockId="87.[151,1437,150,1725]" pageId="87" pageNumber="88">
<emphasis id="26C3EAC0C736FF82FF5BFF24FE83B5CC" bold="true" box="[199,263,150,176]" pageId="87" pageNumber="88">Head</emphasis>
(
<figureCitation id="8C8C2A57C736FF82FE86FF24FE3BB5CC" box="[282,447,150,176]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="87" pageNumber="88">Figs. 211212</figureCitation>
). 1.2 times broader than long. Galea shagreened, matt. Labral process smooth, much broader than long, apical margin slightly arched. Clypeus moderately protuberant, strongly arched, basal half strongly and finely shagreened, apical half gradually smooth, punctation fine and dense, distance between punctures 11.5 puncture diameters (
<figureCitation id="8C8C2A57C736FF82FE22FEB0FD9AB460" box="[446,542,258,284]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="87" pageNumber="88">Fig. 211</figureCitation>
). Malar area length 0.2 times width of mandible base. Flagellomere 1 longer than 2+3. Frons longitudinally striated, interspersed with sparse, shallow fine punctures. Facial foveae broad, occupying slightly more than half distance between compound eye and lateral ocellus (
<figureCitation id="8C8C2A57C736FF82FB80FEF8FB47B418" box="[1052,1219,330,356]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="87" pageNumber="88">Figs. 211212</figureCitation>
). Distance of fovea from lateral ocellus 1.5 times width of lateral ocellus. Ocelloccipital distance 1.4 ocellus diameters. Vertex carinate. Genal area 1.2 times as broad as compound eye (
<figureCitation id="8C8C2A57C736FF82FD27FE20FC9AB4D0" box="[699,798,402,428]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="87" pageNumber="88">Fig. 212</figureCitation>
).
</paragraph>
<paragraph id="140836D2C736FF82FF5BFE04FCA8B668" blockId="87.[151,1437,150,1725]" pageId="87" pageNumber="88">
<emphasis id="26C3EAC0C736FF82FF5BFE04FEC1B4AC" bold="true" box="[199,325,438,464]" pageId="87" pageNumber="88">Mesosoma</emphasis>
(
<figureCitation id="8C8C2A57C736FF82FEC8FE04FE37B4AC" box="[340,435,438,464]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="87" pageNumber="88">Fig. 212</figureCitation>
). Dorsolateral angle of pronotum not elevated, pronotum not carinate. Mesonotum strongly and uniformly shagreened, weakly shiny, with dense, shallow but distinct, coarse crater-like punctures, distance between punctures 0.51 puncture diameters. Scutellum shagreened peripherally, shiny and weakly shagreened centrally, with regular (not crater-like) punctures, distance between punctures 1 puncture diameter (
<figureCitation id="8C8C2A57C736FF82FB7FFD90FAC1B740" box="[1251,1349,546,572]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="87" pageNumber="88">Fig. 212</figureCitation>
). Mesepisternum finely alveolate, densely and shallowly punctured, punctures merging onto underlying sculpture. Propodeal corbicula finely reticulate. Posterior part of propodeum clearly divided into basal, moderately sloping part, and apical vertical part. Posterolateral part of propodeum finely alveolate, shallowly, obliquely punctured. Propodeal triangle narrow, rugose basally, finely alveolate apically. Inner side of hind femur not carinate. Inner hind tibial spur almost straight, of uniform width. Hind pretarsal claw bidentate. Nervulus interstitial (
<figureCitation id="8C8C2A57C736FF82FB0BFD64FB7FB78C" box="[1175,1275,726,752]" captionStart="FIGURES 204 212" captionStartId="85.[151,264,1943,1967]" captionTargetBox="[222,1367,181,1916]" captionTargetId="figure-18@85.[202,1386,181,1920]" captionTargetPageId="85" captionText="FIGURES 204 212. Andrena (incertae sedis) guttata Warncke. 204. male habitus, 205. male head, 206. male head and mesosoma, 207. male terga 13, 208. male genitalia, 209. male eighth sternum. Andrena (incertae sedis) hulae sp. nov. 210. female habitus, 211. female head, 212. female head and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.7073884" httpUri="https://zenodo.org/record/7073884/files/figure.png" pageId="87" pageNumber="88">Fig. 210</figureCitation>
). Submarginal crossvein 1 meets marginal cell 4 vein widths from stigma.
</paragraph>
<paragraph id="140836D2C736FF82FF5BFCACFAAFB6FD" blockId="87.[151,1437,150,1725]" pageId="87" pageNumber="88">
<emphasis id="26C3EAC0C736FF82FF5BFCACFEC7B644" bold="true" box="[199,323,798,824]" pageId="87" pageNumber="88">Metasoma</emphasis>
(
<figureCitation id="8C8C2A57C736FF82FECBFCACFE3EB644" box="[343,442,798,824]" captionStart="FIGURES 213221" captionStartId="88.[151,264,1943,1967]" captionTargetBox="[220,1369,181,1919]" captionTargetId="figure-28@88.[204,1383,181,1920]" captionTargetPageId="88" captionText="FIGURES 213221. Andrena (incertae sedis) hulae sp. nov. 213. female metasoma. Andrena (incertae sedis) inusitata sp. nov. 214. male habitus, 215. male head, 216. male head and mesosoma, 217. male lateral part of propodeum, 218. male metasoma, 219. male genitalia, 220. male eighth sternum, ventral view, 221. male eighth sternum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7073888" httpUri="https://zenodo.org/record/7073888/files/figure.png" pageId="87" pageNumber="88">Fig. 213</figureCitation>
). Tergal discs impunctate, 12 finely shagreened, 34 weakly, finely shagreened and shiny. Tergal marginal zones similarly sculptured, smooth near apex, 24 slightly depressed, centrally occupying 1/31/2 of tergum length. Pygidial plate finely alveolate, rounded apically, without elevated central area.
</paragraph>
<paragraph id="140836D2C736FF82FF5BFC38FE0CB6D8" blockId="87.[151,1437,150,1725]" box="[199,392,906,932]" pageId="87" pageNumber="88">
<emphasis id="26C3EAC0C736FF82FF5BFC38FE8FB6D8" bold="true" box="[199,267,906,932]" pageId="87" pageNumber="88">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="5CAD6559C736FF82FF5BFC1CFEB3B0E0" pageId="87" pageNumber="88" type="diagnosis">
<paragraph id="140836D2C736FF82FF5BFC1CFEB3B0E0" blockId="87.[151,1437,150,1725]" pageId="87" pageNumber="88">
<emphasis id="26C3EAC0C736FF82FF5BFC1CFEC6B6B4" bold="true" box="[199,322,942,968]" pageId="87" pageNumber="88">Diagnosis.</emphasis>
<taxonomicName id="D3B74D51C736FF82FED5FC1CFE6AB6B4" authorityName="Pisanty &amp; Scheuchl &amp; Martin &amp; Cardinal &amp; Wood" authorityYear="2022" box="[329,494,942,968]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="hulae">
<emphasis id="26C3EAC0C736FF82FED5FC1CFE6AB6B4" box="[329,494,942,968]" italics="true" pageId="87" pageNumber="88">Andrena hulae</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="D3B74D51C736FF82FD5EFC1DFC06B6B4" authority="Wood" authorityName="Wood" box="[706,898,942,969]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="26C3EAC0C736FF82FD5EFC1DFCB2B6B4" box="[706,822,943,969]" italics="true" pageId="87" pageNumber="88">A. ramosa</emphasis>
Wood
</taxonomicName>
, recently described from southern
<collectingCountry id="6CA07642C736FF82FA95FC1CFAC8B6B4" box="[1289,1356,942,968]" name="Spain" pageId="87" pageNumber="88">Spain</collectingCountry>
, which is morphologically related to the subgenus
<taxonomicName id="D3B74D51C736FF82FD1CFC60FC7FB690" authority="(Wood et al. 2022)" baseAuthorityName="Wood" baseAuthorityYear="2022" box="[640,1019,978,1004]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="subGenus" subGenus="Planiandrena">
<emphasis id="26C3EAC0C736FF82FD1CFC60FC9DB690" box="[640,793,978,1004]" italics="true" pageId="87" pageNumber="88">Planiandrena</emphasis>
(
<bibRefCitation id="70264B23C736FF82FCB5FC60FC70B690" author="Wood, T. J. &amp; Molina, F. P. &amp; Bartomeus, I." box="[809,1012,978,1004]" pageId="87" pageNumber="88" pagination="1 - 13" refId="ref90021" refString="Wood, T. J., Molina, F. P. &amp; Bartomeus, I. (2022) A new Andrena species (Hymenoptera: Andrenidae) from the overlooked Donana Protected Areas of southern Spain. Belgian Journal of Entomology, 126, 1 - 13. https: // doi. org / 10.47446 / OSMIA 10.1" type="journal article" year="2022">
Wood
<emphasis id="26C3EAC0C736FF82FCE9FC61FC2BB690" box="[885,943,978,1004]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
2022
</bibRefCitation>
)
</taxonomicName>
. However, recent molecular findings show that the COI barcode of
<taxonomicName id="D3B74D51C736FF82FE78FC45FDD3B16C" box="[484,599,1015,1041]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="26C3EAC0C736FF82FE78FC45FDD3B16C" box="[484,599,1015,1041]" italics="true" pageId="87" pageNumber="88">A. ramosa</emphasis>
</taxonomicName>
falls closest to species of
<taxonomicName id="D3B74D51C736FF82FCEBFC44FC76B16C" box="[887,1010,1014,1040]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="38" phylum="Arthropoda" rank="subGenus" subGenus="Euandrena">
<emphasis id="26C3EAC0C736FF82FCEBFC44FC76B16C" box="[887,1010,1014,1040]" italics="true" pageId="87" pageNumber="88">Euandrena</emphasis>
</taxonomicName>
, questioning its subgeneric placement (TJW, unpublished results). We therefore place both
<taxonomicName id="D3B74D51C736FF82FD73FBA9FCCAB148" authorityName="Pisanty &amp; Scheuchl &amp; Martin &amp; Cardinal &amp; Wood" authorityYear="2022" box="[751,846,1050,1077]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="hulae">
<emphasis id="26C3EAC0C736FF82FD73FBA9FCCAB148" box="[751,846,1050,1077]" italics="true" pageId="87" pageNumber="88">A. hulae</emphasis>
</taxonomicName>
and
<taxonomicName id="D3B74D51C736FF82FC1BFBA9FC78B148" box="[903,1020,1051,1077]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="26C3EAC0C736FF82FC1BFBA9FC78B148" box="[903,1020,1051,1077]" italics="true" pageId="87" pageNumber="88">A. ramosa</emphasis>
</taxonomicName>
as
<emphasis id="26C3EAC0C736FF82FBB8FBA9FB44B148" box="[1060,1216,1050,1076]" italics="true" pageId="87" pageNumber="88">incertae sedis</emphasis>
for the time being.
<taxonomicName id="D3B74D51C736FF82FF0BFB8CFEB8B124" authorityName="Pisanty &amp; Scheuchl &amp; Martin &amp; Cardinal &amp; Wood" authorityYear="2022" box="[151,316,1086,1112]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="hulae">
<emphasis id="26C3EAC0C736FF82FF0BFB8CFEB8B124" box="[151,316,1086,1112]" italics="true" pageId="87" pageNumber="88">Andrena hulae</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="D3B74D51C736FF82FE4DFB8CFDEEB124" box="[465,618,1086,1112]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="subGenus" subGenus="Planiandrena">
<emphasis id="26C3EAC0C736FF82FE4DFB8CFDEEB124" box="[465,618,1086,1112]" italics="true" pageId="87" pageNumber="88">Planiandrena</emphasis>
</taxonomicName>
species in its short and broad labral process and its relatively broad propodeal triangle with short longitudinal rugae basally, but its fovea is substantially broader, the hind tibial spur is not broadened submedially, and the pygidial plate lacks a centrally elevated area. Compared to
<taxonomicName id="D3B74D51C736FF82FB0EFB35FA80B1DC" box="[1170,1284,1159,1185]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="26C3EAC0C736FF82FB0EFB35FA80B1DC" box="[1170,1284,1159,1185]" italics="true" pageId="87" pageNumber="88">A. ramosa</emphasis>
</taxonomicName>
,
<taxonomicName id="D3B74D51C736FF82FA8EFB35FAEBB1DC" authorityName="Pisanty &amp; Scheuchl &amp; Martin &amp; Cardinal &amp; Wood" authorityYear="2022" box="[1298,1391,1158,1185]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="hulae">
<emphasis id="26C3EAC0C736FF82FA8EFB35FAEBB1DC" box="[1298,1391,1158,1185]" italics="true" pageId="87" pageNumber="88">A. hulae</emphasis>
</taxonomicName>
can be separated by its smaller body size of
<quantity id="D34F9B37C736FF82FDC2FB18FD31B1B9" box="[606,693,1194,1221]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="87" pageNumber="88" unit="mm" value="8.5">8.5 mm</quantity>
(
<quantity id="D34F9B37C736FF82FD58FB18FC91B1B9" box="[708,789,1194,1221]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="87" pageNumber="88" unit="mm" value="12.0">12 mm</quantity>
in
<taxonomicName id="D3B74D51C736FF82FCA6FB19FC2AB1B8" box="[826,942,1195,1221]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="26C3EAC0C736FF82FCA6FB19FC2AB1B8" box="[826,942,1195,1221]" italics="true" pageId="87" pageNumber="88">A. ramosa</emphasis>
</taxonomicName>
), its broad foveae that occupy over half the space between the compound eye and the lateral ocellus (occupying 1/3 of this space, separated from lateral ocellus by over two times diameter of lateral ocellus in
<taxonomicName id="D3B74D51C736FF82FD29FB41FCACB070" box="[693,808,1267,1293]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="26C3EAC0C736FF82FD29FB41FCACB070" box="[693,808,1267,1293]" italics="true" pageId="87" pageNumber="88">A. ramosa</emphasis>
</taxonomicName>
), propodeal triangle with short basal longitudinal rugae (without rugae basally in
<taxonomicName id="D3B74D51C736FF82FE2AFAA5FDADB04C" box="[438,553,1303,1329]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="26C3EAC0C736FF82FE2AFAA5FDADB04C" box="[438,553,1303,1329]" italics="true" pageId="87" pageNumber="88">A. ramosa</emphasis>
</taxonomicName>
), its flat and centrally alveolate pygidial plate (pygidial plate with smooth surface, with weakly raised longitudinal ridge in
<taxonomicName id="D3B74D51C736FF82FD03FA89FC97B028" box="[671,787,1339,1365]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="26C3EAC0C736FF82FD03FA89FC97B028" box="[671,787,1339,1365]" italics="true" pageId="87" pageNumber="88">A. ramosa</emphasis>
</taxonomicName>
), and by its weakly plumose hairs on the mesepisternum, propodeal corbiculae, flocculus, and femoral scopae (these areas with extremely long and extremely plumose hairs in
<taxonomicName id="D3B74D51C736FF82FF28FA31FEA3B0E0" box="[180,295,1411,1437]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="ramosa">
<emphasis id="26C3EAC0C736FF82FF28FA31FEA3B0E0" box="[180,295,1411,1437]" italics="true" pageId="87" pageNumber="88">A. ramosa</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="5CAD6559C736FF82FF5BFA14FE54B375" pageId="87" pageNumber="88" type="distribution">
<paragraph id="140836D2C736FF82FF5BFA14FD98B0BC" blockId="87.[151,1437,150,1725]" box="[199,540,1446,1472]" pageId="87" pageNumber="88">
<emphasis id="26C3EAC0C736FF82FF5BFA14FEE7B0BC" bold="true" box="[199,355,1446,1472]" pageId="87" pageNumber="88">Distribution:</emphasis>
Northern
<collectingCountry id="6CA07642C736FF82FE4AFA14FD9CB0BC" box="[470,536,1446,1472]" name="Israel" pageId="87" pageNumber="88">Israel</collectingCountry>
.
</paragraph>
<paragraph id="140836D2C736FF82FF5BFA78FE41B098" blockId="87.[151,1437,150,1725]" box="[199,453,1482,1508]" pageId="87" pageNumber="88">
<emphasis id="26C3EAC0C736FF82FF5BFA78FEE9B098" bold="true" box="[199,365,1482,1508]" pageId="87" pageNumber="88">Flight period:</emphasis>
March.
</paragraph>
<paragraph id="140836D2C736FF82FF5BFA5CFE54B375" blockId="87.[151,1437,150,1725]" box="[199,464,1518,1545]" pageId="87" pageNumber="88">
<emphasis id="26C3EAC0C736FF82FF5BFA5CFE01B374" bold="true" box="[199,389,1518,1544]" pageId="87" pageNumber="88">Flower records:</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="5CAD6559C736FF82FF5BF9A0FCEBB308" pageId="87" pageNumber="88" type="materials_examined">
<paragraph id="140836D2C736FF82FF5BF9A0FE11B32C" blockId="87.[151,1437,150,1725]" pageId="87" pageNumber="88">
<materialsCitation id="A4DF3C8FC736FF82FF5BF9A0FE11B32C" ID-GBIF-Occurrence="3947000690" collectingDate="1995-03-19" collectionCode="SMNHTAU" collectorName="N Rosh-Pina &amp; R. Kasher" country="Israel" location="Sde Eliezer" pageId="87" pageNumber="88" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="26C3EAC0C736FF82FF5BF9A0FE4FB350" bold="true" box="[199,459,1554,1580]" pageId="87" pageNumber="88">
<typeStatus id="CB0C8870C736FF82FF5BF9A0FEBCB350" box="[199,312,1554,1580]" pageId="87" pageNumber="88" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="6CA07642C736FF82FEC6F9A0FE43B350" box="[346,455,1554,1580]" name="Israel" pageId="87" pageNumber="88">ISRAEL</collectingCountry>
:
</emphasis>
<location id="11686009C736FF82FE74F9A0FDF9B350" LSID="urn:lsid:plazi:treatment:9C1E87C4C737FF82FF0BFA0CFEF4B3C0:11686009C736FF82FE74F9A0FDF9B350" box="[488,637,1554,1580]" country="Israel" name="Sde Eliezer" pageId="87" pageNumber="88">Sde Eliezer</location>
[Sede Eliezer],
<collectorName id="B9425304C736FF82FCE7F9A1FBA7B350" box="[891,1059,1554,1581]" pageId="87" pageNumber="88">N Rosh-Pina</collectorName>
,
<date id="60091012C736FF82FBDFF9A0FB3BB351" box="[1091,1215,1554,1581]" pageId="87" pageNumber="88" value="1995-03-19">
<collectingDate id="704DE9FAC736FF82FBDFF9A0FB3BB351" box="[1091,1215,1554,1581]" pageId="87" pageNumber="88" value="1995-03-19">19.iii.1995</collectingDate>
</date>
,
<collectorName id="B9425304C736FF82FB7CF9A1FAE3B350" box="[1248,1383,1554,1581]" pageId="87" pageNumber="88">R. Kasher</collectorName>
,
<specimenCount id="02B1FD5BC736FF82FA1BF9A1FA18B351" box="[1415,1436,1555,1581]" pageId="87" pageNumber="88" type="female"></specimenCount>
(
<collectionCode id="72A6AE17C736FF82FF03F984FEB5B32C" box="[159,305,1590,1616]" pageId="87" pageNumber="88">SMNHTAU</collectionCode>
:367741).
</materialsCitation>
</paragraph>
<paragraph id="140836D2C736FF82FF5BF9E8FCEBB308" blockId="87.[151,1437,150,1725]" box="[199,879,1626,1653]" pageId="87" pageNumber="88">
<materialsCitation id="A4DF3C8FC736FF82FF5BF9E8FCEFB308" ID-GBIF-Occurrence="3947000833" box="[199,875,1626,1653]" collectionCode="SMNHTAU" country="Israel" location="ISRAEL" pageId="87" pageNumber="88" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="26C3EAC0C736FF82FF5BF9E8FE45B308" bold="true" box="[199,449,1626,1652]" pageId="87" pageNumber="88">
<typeStatus id="CB0C8870C736FF82FF5BF9E8FEC7B308" box="[199,323,1626,1652]" pageId="87" pageNumber="88" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="6CA07642C736FF82FED3F9E8FE38B308" box="[335,444,1626,1652]" name="Israel" pageId="87" pageNumber="88">ISRAEL</collectingCountry>
:
</emphasis>
same as
<typeStatus id="CB0C8870C736FF82FDBBF9E8FD0DB308" box="[551,649,1626,1652]" pageId="87" pageNumber="88" type="holotype">holotype</typeStatus>
(
<specimenCount id="02B1FD5BC736FF82FD07F9E8FD3EB308" box="[667,698,1626,1652]" pageId="87" pageNumber="88" type="female">1♀</specimenCount>
) (
<collectionCode id="72A6AE17C736FF82FD48F9E8FCE7B308" box="[724,867,1626,1652]" pageId="87" pageNumber="88">SMNHTAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5CAD6559C736FF82FF5BF9CCFEF4B3C0" pageId="87" pageNumber="88" type="etymology">
<paragraph id="140836D2C736FF82FF5BF9CCFEF4B3C0" blockId="87.[151,1437,150,1725]" pageId="87" pageNumber="88">
<emphasis id="26C3EAC0C736FF82FF5BF9CCFECAB3E4" bold="true" box="[199,334,1662,1688]" pageId="87" pageNumber="88">Etymology.</emphasis>
Named after the Hula Valley in northern
<collectingCountry id="6CA07642C736FF82FCBFF9CCFCE0B3E4" box="[803,868,1662,1688]" name="Israel" pageId="87" pageNumber="88">Israel</collectingCountry>
, where the species was collected. The species epithet is an adjective.
</paragraph>
</subSubSection>
</treatment>
</document>