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<mods:title id="CE3C338BB863B34AB4D563A1A1C2B43C">A new iphiculid crab (Crustacea, Brachyura, Leucosioidea) from the Middle Miocene of Austria, with notes on palaeobiogeography of Iphiculus</mods:title>
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<treatment id="9605E23F9270FFCE96D8F90BFBF2FE51" ID-DOI="http://doi.org/10.5281/zenodo.6083809" ID-GBIF-Taxon="124560776" ID-Zenodo-Dep="6083809" LSID="urn:lsid:plazi:treatment:9605E23F9270FFCE96D8F90BFBF2FE51" httpUri="http://treatment.plazi.org/id/9605E23F9270FFCE96D8F90BFBF2FE51" lastPageId="3" lastPageNumber="266" pageId="2" pageNumber="265">
<subSubSection id="56B600A29270FFCF96D8F90BFDFDF93D" box="[151,551,1702,1729]" pageId="2" pageNumber="265" type="nomenclature">
<paragraph id="1E1353299270FFCF96D8F90BFDFDF93D" blockId="2.[151,551,1702,1729]" box="[151,551,1702,1729]" pageId="2" pageNumber="265">
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<taxonomicName id="D9AC28AA9270FFCF96D8F90BFDFDF93D" ID-CoL="7PLPF" authority="Adams &amp; White, 1849" authorityName="Adams &amp; White" authorityYear="1849" box="[151,551,1702,1729]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="2CD88F3B9270FFCF96D8F90BFDFDF93D" bold="true" box="[151,551,1702,1729]" pageId="2" pageNumber="265">
<emphasis id="2CD88F3B9270FFCF96D8F90BFED1F93C" bold="true" box="[151,267,1702,1728]" italics="true" pageId="2" pageNumber="265">Iphiculus</emphasis>
<bibRefCitation id="7A3D2ED89270FFCF975DF90AFDFDF93D" author="Adams" box="[274,551,1703,1729]" pageId="2" pageNumber="265" refString="Adams, A. &amp; White, A. (1849) Crustacea, Part II. In: Adams, A. (Ed.), Zoology of the Voyage of H. M. S. Samarang; under the command of Captain Sir Edward Belcher, C. B., F. R. A. S., F. G. S., during the years 1843 - 1846. Reeve, Benham &amp; Reeve, London, pp. 33 - 67, pls. 7 - 13." type="book chapter" year="1849">Adams &amp; White, 1849</bibRefCitation>
</emphasis>
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<paragraph id="1E1353299270FFCF96D8F943FCA4F8FF" blockId="2.[151,894,1773,1795]" box="[151,894,1773,1795]" pageId="2" pageNumber="265">
Type species.
<taxonomicName id="D9AC28AA9270FFCF976AF943FD32F8FF" authority="Adams &amp; White, 1849" authorityName="Adams &amp; White" authorityYear="1849" box="[293,744,1773,1795]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="265" phylum="Arthropoda" rank="species" species="spongiosus">
<emphasis id="2CD88F3B9270FFCF976AF943FE21F8FF" box="[293,507,1774,1795]" italics="true" pageId="2" pageNumber="265">Iphiculus spongiosus</emphasis>
<bibRefCitation id="7A3D2ED89270FFCF944FF940FD32F8FF" author="Adams" box="[512,744,1773,1795]" pageId="2" pageNumber="265" refString="Adams, A. &amp; White, A. (1849) Crustacea, Part II. In: Adams, A. (Ed.), Zoology of the Voyage of H. M. S. Samarang; under the command of Captain Sir Edward Belcher, C. B., F. R. A. S., F. G. S., during the years 1843 - 1846. Reeve, Benham &amp; Reeve, London, pp. 33 - 67, pls. 7 - 13." type="book chapter" year="1849">Adams &amp; White, 1849</bibRefCitation>
</taxonomicName>
, by monotypy.
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<paragraph id="1E1353299270FFCE96D8F89DFBF2FEE0" blockId="2.[151,1436,1840,2009]" lastBlockId="3.[151,1436,151,429]" lastPageId="3" lastPageNumber="266" pageId="2" pageNumber="265">
<emphasis id="2CD88F3B9270FFCF96D8F89DFED1F8B5" bold="true" box="[151,267,1840,1865]" pageId="2" pageNumber="265">Remarks.</emphasis>
A handful of species are currently included in this genus. Its identification in the fossil record (
<bibRefCitation id="7A3D2ED89270FFCF937DF89DFEF6F891" author="Morris" pageId="2" pageNumber="265" refString="Morris, S. F. &amp; Collins, J. S. H. (1991) Neogene crabs from Brunei, Sabah and Sarawak. Bulletin of the British Museum of Natural History (Geology), 47, 1 - 33." type="journal article" year="1991">Morris &amp; Collins 1991</bibRefCitation>
;
<bibRefCitation id="7A3D2ED89270FFCF9775F8F9FDCBF891" author="Collins" box="[314,529,1876,1901]" pageId="2" pageNumber="265" refString="Collins, J. S. H., Lee, C. &amp; Noad, J. (2003) Miocene and Pleistocene crabs (Crustacea, Decapoda) from Sabah and Sarawak. Journal of Systematic Palaeontology, 1, 187 - 226. http: // dx. doi. org / 10.1017 / S 1477201903001068" type="journal article" year="2003">
Collins
<emphasis id="2CD88F3B9270FFCF97DBF8FBFE14F891" box="[404,462,1876,1901]" italics="true" pageId="2" pageNumber="265">et al.</emphasis>
2003
</bibRefCitation>
) is largely based on the carapace outline, configuration of spines on the lateral margins and carapace surface ornamentation. In this respect it should be noted that some species of
<taxonomicName id="D9AC28AA9270FFCF9360F8D5FE93F849" authority="Stoliczka, 1871" authorityName="Stoliczka" authorityYear="1871" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="2CD88F3B9270FFCF9360F8D5FA41F86D" box="[1327,1435,1912,1937]" italics="true" pageId="2" pageNumber="265">Typilobus</emphasis>
<bibRefCitation id="7A3D2ED89270FFCF96D8F831FE93F849" author="Stoliczka" box="[151,329,1948,1973]" pageId="2" pageNumber="265" refString="Stoliczka, F. (1871) Observations of fossil crabs from Tertiary deposits in Sind and Kutch. Memoirs of the Geological Survey of India, Palaeontologica Indica, Series 7, XIV, Vol. 1, part 1, 1 - 16, 5 pls." type="book chapter" year="1871">Stoliczka, 1871</bibRefCitation>
</taxonomicName>
, a leucosiid genus known exclusively from the fossil record, resemble extant representatives of
<taxonomicName id="D9AC28AA9270FFCF96D8F86DFF25F825" box="[151,255,1984,2009]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="2CD88F3B9270FFCF96D8F86DFF25F825" box="[151,255,1984,2009]" italics="true" pageId="2" pageNumber="265">Iphiculus</emphasis>
</taxonomicName>
.
<bibRefCitation id="7A3D2ED89270FFCF9741F86DFDD6F825" author="Artal" box="[270,524,1984,2009]" pageId="2" pageNumber="265" refString="Artal, P. &amp; Hyzny, M. (2016) An appraisal of Typilobus Stoliczka, 1871 (Crustacea, Brachyura, Leucosioidea), with description of a new family and genus. Zootaxa, 4117 (3), 387 - 398. http: // dx. doi. org / 10.11646 / zootaxa. 4117.3.6" type="journal article" year="2016">Artal &amp; Hyžný (2016)</bibRefCitation>
presented an appraisal of the fossil leucosiid genus
<taxonomicName id="D9AC28AA9270FFCF9211F86DFB13F825" box="[1118,1225,1984,2009]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="2CD88F3B9270FFCF9211F86DFB13F825" box="[1118,1225,1984,2009]" italics="true" pageId="2" pageNumber="265">Typilobus</emphasis>
</taxonomicName>
. They pointed out the heterogeneity of the taxon, as previously emphasized by
<bibRefCitation id="7A3D2ED89271FFCE9573FF35FBDDFF4C" box="[828,1031,151,176]" pageId="3" pageNumber="266" refString="Via Boada, L. (1969) Crustaceos decapodos del Eoceno espanol. Pirineos, 91 - 94, 1 - 469." type="book chapter">Vía Boada (1969)</bibRefCitation>
,
<bibRefCitation id="7A3D2ED89271FFCE925CFF3AFB68FF4C" author="Muller" box="[1043,1202,151,176]" pageId="3" pageNumber="266" refString="Muller, P. (1993) Neogene decapod crustaceans from Catalonia. Scripta Musei Geologici Seminarii Barcinonensis, 225, 1 - 39." type="journal article" year="1993">Müller (1993)</bibRefCitation>
, and
<bibRefCitation id="7A3D2ED89271FFCE92A2FF3AFF3AFF28" pageId="3" pageNumber="266" refString="Feldmann, R. M., Schweitzer, C. E., Bennett, O., Frantescu, O., Resar, N. &amp; Trudeau, A. (2011) New Eocene Brachyura (Crustacea: Decapoda) from Egypt. Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen, 262, 323 - 353. http: // dx. doi. org / 10.1127 / 0077 - 7749 / 2011 / 0202" type="journal article">
Feldmann
<emphasis id="2CD88F3B9271FFCE932BFF34FA46FF4C" box="[1380,1436,151,176]" italics="true" pageId="3" pageNumber="266">et al.</emphasis>
(2011)
</bibRefCitation>
and reassigned
<taxonomicName id="D9AC28AA9271FFCE97EEFF11FCE8FF29" authority="Via Boada, 1959" authorityName="Via Boada" authorityYear="1959" box="[417,818,188,213]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="266" phylum="Arthropoda" rank="species" species="boscoi">
<emphasis id="2CD88F3B9271FFCE97EEFF11FDBBFF29" box="[417,609,188,213]" italics="true" pageId="3" pageNumber="266">Typilobus boscoi</emphasis>
<bibRefCitation id="7A3D2ED89271FFCE9423FF11FCE8FF29" author="Via" box="[620,818,188,213]" pageId="3" pageNumber="266" refString="Via Boada, L. (1959) Decapodos fosiles del Eoceno espanol. Boletin del Instituto Geologico y Minero de Espana, 70, 331 - 402." type="journal article" year="1959">Vía Boada, 1959</bibRefCitation>
</taxonomicName>
to a new genus and family. More re-assignments of species now classified within the broadly defined
<taxonomicName id="D9AC28AA9271FFCE948CFF72FCEAFF04" box="[707,816,223,248]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis id="2CD88F3B9271FFCE948CFF72FCEAFF04" box="[707,816,223,248]" italics="true" pageId="3" pageNumber="266">Typilobus</emphasis>
</taxonomicName>
are likely once the genus is reviewed. It is possible that some of them will fall within the range of characters which now define
<taxonomicName id="D9AC28AA9271FFCE95F7FEA9FBFAFEE1" box="[952,1056,260,285]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis id="2CD88F3B9271FFCE95F7FEA9FBFAFEE1" box="[952,1056,260,285]" italics="true" pageId="3" pageNumber="266">Iphiculus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1E1353299271FFCE9688FE85FBF2FE51" blockId="3.[151,1436,151,429]" pageId="3" pageNumber="266">
Current assignment of a newly described species to
<taxonomicName id="D9AC28AA9271FFCE9554FE8AFC5EFEBC" box="[795,900,295,320]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis id="2CD88F3B9271FFCE9554FE8AFC5EFEBC" box="[795,900,295,320]" italics="true" pageId="3" pageNumber="266">Iphiculus</emphasis>
</taxonomicName>
is based on striking morphological similarities with extant
<taxonomicName id="D9AC28AA9271FFCE9750FEE0FE45FE98" box="[287,415,333,356]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="266" phylum="Arthropoda" rank="species" species="convexus">
<emphasis id="2CD88F3B9271FFCE9750FEE0FE45FE98" box="[287,415,333,356]" italics="true" pageId="3" pageNumber="266">I. convexus</emphasis>
</taxonomicName>
(
<figureCitation id="86974FAC9271FFCE97FFFEE1FE2DFE98" box="[432,503,332,357]" captionStart="FIGURE 2" captionStartId="3.[151,249,1386,1408]" captionTargetBox="[151,1435,447,1361]" captionTargetId="figure@3.[151,1435,447,1361]" captionTargetPageId="3" captionText="FIGURE 2. Fossil and extant Iphiculus. A, Iphiculus eliasi sp. nov., holotype UMJGP 75.612 in dorsal view (dry and uncoated); B – D, Iphiculus convexus Ihle, 1918, ZRC 2009.0462 (male specimen from Vanuatu) in dorsal (B), posterior (C), and posterodorsal view (D). Photos in B – D by P. K. L. Ng." httpUri="https://zenodo.org/record/163904/files/figure.png" pageId="3" pageNumber="266">Fig. 2</figureCitation>
), especially on the general shape of carapace, short anterolateral spines and large rounded tubercles covering evenly the dorsal carapace surface. Since, the new fossil material consists only of isolated carapaces, comparison with sternum, pleon and chelipeds is not possible.
</paragraph>
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</treatment>
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