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<document id="0A439405D1506AB12D1A623E7A6544EC" ID-CLB-Dataset="186676" ID-DOI="10.11646/zootaxa.5205.5.5" ID-GBIF-Dataset="8bf46ddf-12ef-4b59-b92e-674efe374a44" ID-ISSN="1175-5326" ID-Zenodo-Dep="7318559" ID-ZooBank="DD3A8499-DC11-464F-950A-89AB5F885245" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668421593984" checkinUser="plazi" docAuthor="Xue, Guoxi, Qu, Peng, Li, Meng, Chiba, Hideyuki &amp; Li, Wenwei" docDate="2022" docId="8D1487CFFFA9F925FF665958FCA5FEF1" docLanguage="en" docName="zootaxa.5205.5.5.pdf" docOrigin="Zootaxa 5205 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ancistroides armatus subsp. armatus" docType="treatment" docVersion="8" lastPageNumber="488" masterDocId="712DFFB7FFAFF922FFF15C51FFBAFFEC" masterDocTitle="Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae)" masterLastPageNumber="490" masterPageNumber="481" pageNumber="487" updateTime="1699411284781" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="75A91AFFC5B772D415489BF5750A0CC8">Molecular and morphological evidences confirm the statuses of Ancistroides othonias (Hewitson, 1878) and the subspecies of A. nigrita (Latreille, [1824]), sensu Evans 1949 (Lepidoptera, Hesperiidae)</mods:title>
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<mods:namePart id="1D0D163ED1B0A2BBB0DBFE46525C44EB">Qu, Peng</mods:namePart>
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<mods:affiliation id="0A7121519B162E1BE7BC664489943711">College of Food and Bioengineering, Zhengzhou University of Light Industry, No. 136 Ke Xue Avenue, Zhengzhou, Henan Province, 450000, P. R. China &amp; limengjeff @ 126. com; ORCID iD: https: // orcid. org / 0000 - 0002 - 2625 - 9024</mods:affiliation>
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<mods:affiliation id="2C19308A8009946118D2F9C124B4A465">B. P. Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii, 96817 - 0916 U. S. A.</mods:affiliation>
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<treatment id="8D1487CFFFA9F925FF665958FCA5FEF1" ID-DOI="http://doi.org/10.5281/zenodo.7320168" ID-GBIF-Taxon="204971490" ID-Zenodo-Dep="7320168" LSID="urn:lsid:plazi:treatment:8D1487CFFFA9F925FF665958FCA5FEF1" httpUri="http://treatment.plazi.org/id/8D1487CFFFA9F925FF665958FCA5FEF1" lastPageId="7" lastPageNumber="488" pageId="6" pageNumber="487">
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<heading id="5E4A81B5FFA9F924FF665958FD63FACF" bold="true" box="[151,729,1289,1316]" fontSize="11" level="1" pageId="6" pageNumber="487" reason="1">
<taxonomicName id="C2BD4D5AFFA9F924FF665958FD63FACF" ID-CoL="DMRL" authority="(H. Druce, 1873)" baseAuthorityName="H. Druce" baseAuthorityYear="1873" box="[151,729,1289,1316]" class="Insecta" family="Hesperiidae" genus="Ancistroides" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="487" phylum="Arthropoda" rank="subSpecies" species="armatus" subSpecies="armatus">
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<emphasis id="37C9EACBFFA9F924FF665958FDB8FAC8" bold="true" box="[151,514,1289,1316]" italics="true" pageId="6" pageNumber="487">Ancistroides armatus armatus</emphasis>
(H. Druce, 1873)
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<paragraph id="050236D9FFA9F924FF66597FFDF9FAA4" blockId="6.[151,729,1289,1353]" box="[151,579,1326,1353]" pageId="6" pageNumber="487">
(
<figureCitation id="9D862A5CFFA9F924FF6E597FFF5FFAA4" box="[159,229,1326,1352]" captionStart="FIGURE 2" captionStartId="3.[151,250,1026,1050]" captionTargetBox="[157,1432,425,996]" captionTargetId="figure-144@3.[151,1436,417,1003]" captionTargetPageId="3" captionText="FIGURE 2. Adults of Ancistroides species. Codes correspond to the specimen IDs in Table 1 and Figure 1." figureDoi="http://doi.org/10.5281/zenodo.7318567" httpUri="https://zenodo.org/record/7318567/files/figure.png" pageId="6" pageNumber="487">Fig. 2</figureCitation>
: A95, An8;
<figureCitation id="9D862A5CFFA9F924FE9A597FFE0AFAA4" box="[363,432,1326,1352]" captionStart="FIGURE 4" captionStartId="4.[151,250,1403,1427]" captionTargetBox="[159,1430,385,1379]" captionTargetId="figure-123@4.[151,1435,382,1379]" captionTargetPageId="4" captionText="FIGURE 4. Female genitalia of Ancistroides species. Codes correspond to the specimen IDs in Table 1 and Figure 1. A. lateral view; B. ventral view; C. lamella postvaginalis, front view." figureDoi="http://doi.org/10.5281/zenodo.7318573" httpUri="https://zenodo.org/record/7318573/files/figure.png" pageId="6" pageNumber="487">Fig. 4</figureCitation>
: An8;
<figureCitation id="9D862A5CFFA9F924FE09597FFD81FAA4" box="[504,571,1326,1352]" captionStart="FIGURE 7" captionStartId="7.[151,250,907,931]" captionTargetBox="[159,1433,310,882]" captionTargetId="figure-74@7.[151,1435,309,883]" captionTargetPageId="7" captionText="FIGURE 7. Male genitalia of Ancistroides armatus (A95). A. ring, lateral view; B. tegumen, dorsal view; C. gnathos, ventral view; D. uncus, dorsal view; E. left valva, inner side; F. left valva, dorsal view; G. aedeagus, dorsal view; H. aedeagus, lateral view; I. juxta." figureDoi="http://doi.org/10.5281/zenodo.7318581" httpUri="https://zenodo.org/record/7318581/files/figure.png" pageId="6" pageNumber="487">Fig. 7</figureCitation>
)
</paragraph>
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<taxonomicName id="C2BD4D5AFFA9F924FF66593DFDF3FA6F" ID-CoL="95ZM3" authority="H. Druce, 1873: 359" authorityName="H. Druce" authorityPageNumber="359" authorityYear="1873" box="[151,585,1387,1412]" class="Insecta" family="Hesperiidae" genus="Astictopterus" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="487" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="37C9EACBFFA9F924FF66593DFECCFA6F" box="[151,374,1388,1411]" italics="true" pageId="6" pageNumber="487">Astictopterus armatus</emphasis>
H. Druce,1873: 359
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.
<typeStatus id="DA06887BFFA9F924FDA3593DFD3EFA68" box="[594,644,1388,1412]" pageId="6" pageNumber="487">Type</typeStatus>
locality: Borneo.
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<taxonomicName id="C2BD4D5AFFA9F924FF6659DAFDDCFA4E" authority=": Elwes &amp; Edwards 1897: 221" authorityName="Elwes &amp; Edwards" authorityPageNumber="221" authorityYear="1897" box="[151,614,1418,1442]" class="Insecta" family="Hesperiidae" genus="Kerana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="487" phylum="Arthropoda" rank="species" species="armata">
<emphasis id="37C9EACBFFA9F924FF6659DAFE89FA4E" box="[151,307,1419,1442]" italics="true" pageId="6" pageNumber="487">Kerana armata</emphasis>
:
<bibRefCitation id="612C4B28FFA9F924FEB159DBFDDCFA4E" author="Elwes, H. J. &amp; Edwards, J." box="[320,614,1418,1442]" pageId="6" pageNumber="487" pagination="101 - 324" refId="ref5524" refString="Elwes, H. J. &amp; Edwards, J. (1897) A revision of the Oriental Hesperiidae. Transactions of the Zoological Society of London, 14 (4), 101 - 324, pls. 18 - 27. https: // doi. org / 10.1111 / j. 1096 - 3642.1897. tb 00059. x" type="journal article" year="1897">Elwes &amp; Edwards 1897: 221</bibRefCitation>
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.
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<treatmentCitation id="841C10C8FFA9F924FF6659F8FD16FA2D" author="Maruyama, K." box="[151,684,1449,1473]" page="29" pageId="6" pageNumber="487" year="1991">
<taxonomicName id="C2BD4D5AFFA9F924FF6659F8FD16FA2D" ID-CoL="DMRL" authority=": Maruyama 1991: 29" authorityName=": Maruyama" authorityPageNumber="29" authorityYear="1991" box="[151,684,1449,1473]" class="Insecta" family="Hesperiidae" genus="Ancistroides" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="487" phylum="Arthropoda" rank="subSpecies" species="armatus" subSpecies="armatus">
<emphasis id="37C9EACBFFA9F924FF6659F8FE70FA2D" box="[151,458,1449,1473]" italics="true" pageId="6" pageNumber="487">Ancistroides armatus armatus</emphasis>
:
<bibRefCitation id="612C4B28FFA9F924FE2859F8FD16FA2D" author="Maruyama, K." box="[473,684,1449,1473]" pageId="6" pageNumber="487" refId="ref6060" refString="Maruyama, K. (1991) Butterflies of Borneo. Vol. 2, No. 2, Hesperiidae. Tobishima Co., Tokyo, xiii + 89 pp., 48 pls." type="book" year="1991">Maruyama 1991: 29</bibRefCitation>
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;
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<bibRefCitation id="612C4B28FFA9F924FD4659F8FCE1FA2D" author="Eliot, J. N." box="[695,859,1449,1473]" pageId="6" pageNumber="487" refId="ref5470" refString="Eliot, J. N. (1992) The Butterflies of the Malay Peninsula. 4 th Revised Edition. The Malayan Nature Society, Kuala Lumpur, x + 595 pp., 69 pls. [originally by Corbet A. S. &amp; Pendlebury, H. M.]" type="book" year="1992">Eliot 1992: 356</bibRefCitation>
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;
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<bibRefCitation id="612C4B28FFA9F924FC9759F8FBACFA2D" author="Evans, W. H." box="[870,1046,1449,1473]" pageId="6" pageNumber="487" refId="ref5587" refString="Evans, W. H. (1949) A catalogue of the Hesperiidae from Europe, Asia &amp; Australia in the British Museum (Natural History). The British Museum, London, 502 pp. https: // doi. org / 10.5962 / bhl. title. 105941" type="book" year="1949">Evans 1949: 281</bibRefCitation>
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;
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;
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The length of forewing is about
<quantity id="C2459B3CFFA9F924FE0D5A56FDF0F9CE" box="[508,586,1543,1570]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="487" unit="mm" value="26.0">26 mm</quantity>
(n = 2) in both sexes. Wing patterns are prominently different to those of the preceding species recognized by clades A to C (
<figureCitation id="9D862A5CFFA9F924FD425A7AFD4CF9A9" box="[691,758,1579,1605]" captionStart="FIGURE 1" captionStartId="2.[151,250,1403,1427]" captionTargetBox="[164,1423,462,1368]" captionTargetId="figure-136@2.[152,1435,453,1379]" captionTargetPageId="2" captionText="FIGURE 1. NJ tree based on Kimura-2-Parameter distances for the partial mitochondrial COI sequences of Ancistroides, values at nodes represent the bootstrap support (BP) values." figureDoi="http://doi.org/10.5281/zenodo.7318563" httpUri="https://zenodo.org/record/7318563/files/figure.png" pageId="6" pageNumber="487">Fig. 1</figureCitation>
): in male, there is a large velvety black patch of specialized scales from vein 1b to the middle of the cell on the ventral side of the forewing, and a similar patch filling the cell on the dorsal side of the hindwing.
</paragraph>
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<subSubSection id="4DA76552FFA9F924FF365AC6FBCBF819" pageId="6" pageNumber="487" type="description">
<paragraph id="050236D9FFA9F924FF365AC6FB0FF866" blockId="6.[151,1437,1543,2038]" pageId="6" pageNumber="487">
<emphasis id="37C9EACBFFA9F924FF365AC6FECBF95D" bold="true" box="[199,369,1687,1713]" pageId="6" pageNumber="487">Male genitalia</emphasis>
(
<figureCitation id="9D862A5CFFA9F924FE715AC6FE7EF95D" box="[384,452,1687,1713]" captionStart="FIGURE 7" captionStartId="7.[151,250,907,931]" captionTargetBox="[159,1433,310,882]" captionTargetId="figure-74@7.[151,1435,309,883]" captionTargetPageId="7" captionText="FIGURE 7. Male genitalia of Ancistroides armatus (A95). A. ring, lateral view; B. tegumen, dorsal view; C. gnathos, ventral view; D. uncus, dorsal view; E. left valva, inner side; F. left valva, dorsal view; G. aedeagus, dorsal view; H. aedeagus, lateral view; I. juxta." figureDoi="http://doi.org/10.5281/zenodo.7318581" httpUri="https://zenodo.org/record/7318581/files/figure.png" pageId="6" pageNumber="487">Fig. 7</figureCitation>
). In lateral view, uncus shorter than tegumen, basal part raised, distal end pointed, lower branch short and triangular; gnathos arm-like, transverse part a little longer than longitudinal part; saccus about 2/5 the length of vinculum. In dorsal view, uncus tapered, distal half bifid and narrowly U-shaped, tip of both branches rounded. In lateral view of valva, dorsal margin flat, distal part bifid into a protrudent shoulder and a sickle shaped branch which bears sawteech on inner edge; harpe triangular, vestigial. Aedeagus remarkably robust, left half of dorsal edge with a wide and deep gap; in lateral view, suprazonal sheath slightly curved downwards, about 3/5 the length of subzonal sheath. Juxta shield-shaped, with a pair of slender and pointed dorsal arms.
</paragraph>
<paragraph id="050236D9FFA9F924FF365BC2FBCBF819" blockId="6.[151,1437,1543,2038]" pageId="6" pageNumber="487">
<emphasis id="37C9EACBFFA9F924FF365BC2FE30F841" bold="true" box="[199,394,1939,1965]" pageId="6" pageNumber="487">Female genitalia</emphasis>
(
<figureCitation id="9D862A5CFFA9F924FE665BC2FE66F841" box="[407,476,1939,1965]" captionStart="FIGURE 4" captionStartId="4.[151,250,1403,1427]" captionTargetBox="[159,1430,385,1379]" captionTargetId="figure-123@4.[151,1435,382,1379]" captionTargetPageId="4" captionText="FIGURE 4. Female genitalia of Ancistroides species. Codes correspond to the specimen IDs in Table 1 and Figure 1. A. lateral view; B. ventral view; C. lamella postvaginalis, front view." figureDoi="http://doi.org/10.5281/zenodo.7318573" httpUri="https://zenodo.org/record/7318573/files/figure.png" pageId="6" pageNumber="487">Fig. 4</figureCitation>
: An8). In lateral view, lamella antevaginal nearly oval, strongly developed; antrum very robust. In ventral view, a wide gap between two strong arms of lamella antevaginal, ostium bursae well sclerotized. Lamella postvaginalis trapezoidal, middle of distal edge with a shallow V-shaped notch.
</paragraph>
</subSubSection>
<subSubSection id="4DA76552FFA8F925FF365CC7FCA5FEF1" pageId="7" pageNumber="488" type="distribution">
<paragraph id="050236D9FFA8F925FF365CC7FB49FF5D" blockId="7.[151,1437,150,285]" box="[199,1267,150,177]" pageId="7" pageNumber="488">
<emphasis id="37C9EACBFFA8F925FF365CC7FEDBFF5C" bold="true" box="[199,353,150,176]" pageId="7" pageNumber="488">Distribution.</emphasis>
S.
<collectingCountry id="7DAA7649FFA8F925FE745CC6FE4EFF5D" box="[389,500,151,177]" name="Myanmar" pageId="7" pageNumber="488">Myanmar</collectingCountry>
, Peninsular
<collectingCountry id="7DAA7649FFA8F925FD8A5CC6FD5BFF5D" box="[635,737,151,177]" name="Thailand" pageId="7" pageNumber="488">Thailand</collectingCountry>
, W.
<collectingCountry id="7DAA7649FFA8F925FCE35CC6FCC1FF5D" box="[786,891,151,177]" name="Malaysia" pageId="7" pageNumber="488">Malaysia</collectingCountry>
, Sumatra, Batoe, Sipora, Borneo.
</paragraph>
<paragraph id="050236D9FFA8F925FF365CEAFCA5FEF1" blockId="7.[151,1437,150,285]" pageId="7" pageNumber="488">
<emphasis id="37C9EACBFFA8F925FF365CEAFE81FF39" bold="true" box="[199,315,187,213]" pageId="7" pageNumber="488">Remarks.</emphasis>
According to
<bibRefCitation id="612C4B28FFA8F925FE2B5CEAFDC8FF39" author="Evans, W. H." box="[474,626,187,213]" pageId="7" pageNumber="488" refId="ref5587" refString="Evans, W. H. (1949) A catalogue of the Hesperiidae from Europe, Asia &amp; Australia in the British Museum (Natural History). The British Museum, London, 502 pp. https: // doi. org / 10.5962 / bhl. title. 105941" type="book" year="1949">Evans (1949)</bibRefCitation>
, there is the other subspecies, viz.
<taxonomicName id="C2BD4D5AFFA8F925FC055CEAFF63FF15" authority="(Evans, 1926)" baseAuthorityName="Evans" baseAuthorityYear="1926" class="Insecta" family="Hesperiidae" genus="Ancistroides" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="488" phylum="Arthropoda" rank="subSpecies" species="armatus" subSpecies="niasana">
<emphasis id="37C9EACBFFA8F925FC055CEAFAFBFF38" box="[1012,1345,187,213]" italics="true" pageId="7" pageNumber="488">Ancistroides armatus niasana</emphasis>
(Evans, 1926)
</taxonomicName>
which is endemic to Nias. The band on the dorsal side of the forewing is obsolete in male, and reduced to separated suffused red spots in female (
<bibRefCitation id="612C4B28FFA8F925FDA15D52FCA8FEF1" author="Evans, W. H." box="[592,786,259,285]" pageId="7" pageNumber="488" refId="ref5587" refString="Evans, W. H. (1949) A catalogue of the Hesperiidae from Europe, Asia &amp; Australia in the British Museum (Natural History). The British Museum, London, 502 pp. https: // doi. org / 10.5962 / bhl. title. 105941" type="book" year="1949">Evans 1949: 281</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>