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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.234.3417" ID-GBIF-Dataset="40136982-78c7-4944-8d18-45c64fb915f1" ID-PMC="PMC3496914" ID-Pensoft-Pub="1313-2970-234-67" ID-PubMed="23372409" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-234-67" ModsDocOrigin="ZooKeys 234" ModsDocTitle="A revision of the Afrotropical spider genus Cambalida Simon, 1909 (Araneae, Corinnidae)" checkinTime="1451248622220" checkinUser="pensoft" docAuthor="Haddad, Charles Richard" docDate="2012" docId="8A73098F3246C95CAE93673B7E298C89" docLanguage="en" docName="ZooKeys 234: 67-119" docOrigin="ZooKeys 234" docSource="http://dx.doi.org/10.3897/zookeys.234.3417" docTitle="Cambalida dippenaarae Haddad, 2012, sp. n." docType="treatment" docVersion="4" lastPageNumber="91" masterDocId="60126347FF99FF86D631C93DB97DC527" masterDocTitle="A revision of the Afrotropical spider genus Cambalida Simon, 1909 (Araneae, Corinnidae)" masterLastPageNumber="119" masterPageNumber="67" pageNumber="87" updateTime="1668154629947" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the Afrotropical spider genus Cambalida Simon, 1909 (Araneae, Corinnidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Haddad, Charles Richard</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>234</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>67</mods:start>
<mods:end>119</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.234.3417</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.234.3417</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-234-67</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152037550" LSID="urn:lsid:zoobank.org:act:D1093D7A-46A6-403E-ACE4-09E66565DAFD" httpUri="http://treatment.plazi.org/id/8A73098F3246C95CAE93673B7E298C89" lastPageId="24" lastPageNumber="91" pageId="20" pageNumber="87">
<subSubSection pageId="20" pageNumber="87" type="nomenclature">
<paragraph pageId="20" pageNumber="87">
<taxonomicName LSID="urn:lsid:zoobank.org:act:D1093D7A-46A6-403E-ACE4-09E66565DAFD" class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida dippenaarae" order="Araneae" pageId="20" pageNumber="87" phylum="Arthropoda" rank="species" species="dippenaarae">
<pageBreakToken pageId="20" pageNumber="87" start="start">Cambalida</pageBreakToken>
dippenaarae
</taxonomicName>
<taxonomicNameLabel pageId="20" pageNumber="87">sp. n.</taxonomicNameLabel>
Figures 1-4, 14-39, 42, 44-49, 53, 70-73
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="87" type="type material">
<paragraph pageId="20" pageNumber="87">Type material.</paragraph>
<paragraph pageId="20" pageNumber="87">
Holotype male, together with one paratype male. ZAMBIA: Livingstone, Quarry near Livingstone Airport,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-17.799967">17°47.998'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="25.776466">25°46.588'E</geoCoordinate>
, leg. C. Haddad &amp; J. Parau, 1.XII.2006 (leaf litter) (NCA 2007/625).
</paragraph>
</subSubSection>
<subSubSection lastPageId="22" lastPageNumber="89" pageId="20" pageNumber="87" type="paratypes">
<paragraph pageId="20" pageNumber="87">Paratypes.</paragraph>
<paragraph pageId="20" pageNumber="87">
SOUTH AFRICA:Eastern Cape Province:Kei Mouth,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-32.686768">32°41.206'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="28.37495">28°22.497'E</geoCoordinate>
, leg. C. Haddad, 8.XII.2005 (leaf litter, coastal forest), 1♂ 2♀ (NCA 2006/1290). Free State Province: Bloemfontein, Free State National Botanical Gardens,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-29.05">29°03'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="26.216667">26°13'E</geoCoordinate>
, leg. C. Haddad, 3.I.2011 (sifting leaf litter), 1♀ (NMBA 16157); Same locality,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-29.033333">29°02'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="26.2">26°12'E</geoCoordinate>
, leg. V. Butler, 25.XI.2009 (
<taxonomicName class="Magnoliopsida" family="Araliaceae" genus="Cussonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cussonia paniculata" order="Apiales" pageId="20" pageNumber="87" phylum="Tracheophyta" rank="species" species="paniculata">Cussonia paniculata</taxonomicName>
leaf litter), 1♂ (NMBA 15727); Same data, 18.IX.2009, 2♀ (NMBA 15666); Same locality,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-29.133333">29°08'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="26.166666">26°10'E</geoCoordinate>
, leg. L. Lotz, XI.2006 (pitfall traps, next to ridge under tree), 1♂ (NMBA 10941); Brandfort district, Florisbad Research Station,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-28.766666">28°46'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="26.083334">26°05'E</geoCoordinate>
, 1250m a.s.l., leg. L.N. Lotz, 1-15.II.1988 (pitfall traps), 1♀ (NMBA 3829). KwaZulu-Natal Province: Ndumo Game Reserve, Crocodile farm,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-26.9071">26°54.426'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="32.31975">32°19.185'E</geoCoordinate>
, leg. C. Haddad, 17.I.2006 (on ground surface), 1♂ 2♀ (NCA 2006/423); Same locality, Environmental Centre,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-26.915916">26°54.955'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="32.306267">32°18.376'E</geoCoordinate>
, leg. C. Haddad &amp; V. Swart, 6.XII.2009 (base of grass tussocks, broadleaf woodland), 1♀ (TMSA 23628); Tembe Elephant Park, Sparse woodland,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-26.95">26°57'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="32.383335">32°23'E</geoCoordinate>
, leg. C. Haddad, 6.I.2002 (searching, leaf litter), 1♀ (NCA 2007/3543); Same locality,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-27.016666">27°01'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="32.4">32°24'E</geoCoordinate>
, leg. C. Haddad, 5.I.2002 (leaf litter, deep sand forest), 5♀ (NCA 2002/378); Umziki Pan, near Hluhluwe [
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-28.033333">28°02'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="32.316666">32°19'E</geoCoordinate>
], leg. P. Reavell, 12.II.1990 (at night on tree trunk), 1♀ (NMSA). North-West Province: Marikana, Buffelspoort,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-25.75">25°45'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="27.483334">27°29'E</geoCoordinate>
, leg. A.S. Honiball, 30.XI.2006, 1♂ 1♀ (NCA 2007/1155); Vryburg district, Weltevrede Farm,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-27.416267">27°24.976'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="24.498434">24°29.906'E</geoCoordinate>
, leg. R. Lyle, R. Fourie, D. du Plessis &amp; J. Adendorff, 8-12.I.2008 (garden, active collecting), 1♂ 2♀ (NCA 2009/3676). ZAMBIA: Wildlives Game Farm, near Choma, open savanna,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-16.9829">16°58.974'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="26.649567">26°38.974'E</geoCoordinate>
, leg. C. Haddad, 4.XII.2006 (leaf litter), 3♂ 7♀ (NCA 2007/552); Same locality, Nabuyani River,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-16.993584">16°59.615'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="26.634884">26°38.093'E</geoCoordinate>
, leg. C. Haddad, 3.XII.2006 (leaf litter), 7♂ 16♀ (NCA 2007/1126).
</paragraph>
<paragraph lastPageId="22" lastPageNumber="89" pageId="20" pageNumber="87">
Other material examined. BOTSWANA: Okavango Delta, near Shakawe, Lesideng Research Camp,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.430367">18°25.822'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="21.896183">21°53.771'E</geoCoordinate>
, leg. C. Haddad, 25.XI.2006 (leaf litter), 1imm. 5♂ (NCA 2007/937); Same locality, leg. C. Haddad, 26-29.XI.2006 (night collecting), 1♂ 2♀ (NCA 2007/974); Same locality, leg. C. Haddad, 26.
<normalizedToken originalValue="XI">XI-</normalizedToken>
11.XII.2006 (pitfalls, riverine forest), 1♀ (NCA 2007/1116); Samochima lagoon, Shakawe Fishing Camp,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.42915">18°25.749'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="21.900583">21°54.035'E</geoCoordinate>
, leg. C. Haddad, 10.XII.2006 (leaf litter), 1imm. 2♀ (NCA 2007/1050). MOZAMBIQUE: Morrungulo Resort, 12m a.s.l.,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-23.23305">23°13.983'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="35.49312">35°29.587'E</geoCoordinate>
, leg. C. Haddad, R. Lyle &amp; R. Fourie, 6.XII.2007 (leaf litter, dune forest), 2imm. 3♂ 1♀ (NCA 2008/188). NAMIBIA: Caprivi strip, 34km E of Divun
<pageBreakToken pageId="21" pageNumber="88" start="start">gu</pageBreakToken>
,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.049067">18°02.944'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="21.910183">21°54.611'E</geoCoordinate>
, leg. C. Haddad, 30.XI.2006 (under rocks), 2♀ (NCA 2007/907). SOUTH AFRICA: Eastern Cape Province: Cwebe Nature Reserve, The Haven,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-32.241615">32°14.497'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="28.910883">28°54.653'E</geoCoordinate>
, leg. C. Haddad, 30.X.2006 (grassy litter behind dunes), 1imm. 1♂ (NCA 2007/243); Great Fish River Reserve, at Selbourne,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-33.133335">33°08'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="26.65">26°39'E</geoCoordinate>
, leg. M. Burger, 5.XII.1993 (pitfall traps), 1♀ (NCA 96/58); Sundays River Valley,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-33.383335">33°23'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="25.433332">25°26'E</geoCoordinate>
, leg. H. Potgieter, 23.XI.1999 (pitfall traps, citrus orchard), 4♂ (NCA 2000/236). Gauteng Province:Kloofendal Nature Reserve, near Roodepoort,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-26.133333">26°08'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="27.866667">27°52'E</geoCoordinate>
, leg. A. Leroy, 9.I.1988 (pitfall traps), 1♀ (NCA 89/151). KwaZulu-Natal Province: 15km N of Richard's Bay,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-28.783333">28°47'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="32.1">32°06'E</geoCoordinate>
, leg. T. Wassenaar, 10.XII.1996 (pitfall traps, rehabilitated coastal forest), 1♂ (NCA 97/841); iSimangaliso [Greater St Lucia] Wetlands Park, Eastern Shores Nature Reserve,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-29.095434">29°05.726'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="26.15725">26°09.435'E</geoCoordinate>
, leg. C. Haddad, 3.VII.2007 (leaf litter), 1imm. 1♀ (NCA 2007/2899); iSimangaliso [Greater St Lucia] Wetlands Park, Hell's Gate,
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-28.038334">28°02.3'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="92" value="32.433334">32°26.0'E</geoCoordinate>
, leg. J. Esterhuizen, 19.I.2003 (tsetse fly traps), 1♂ (NCA 2004/795); Ithala Game Reserve, Ngubhu loop, near ruins,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-27.513617">27°30.817'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="31.2384">31°14.304'E</geoCoordinate>
, leg. C. Haddad, R. Fourie &amp; D. du Plessis, 1.VII.2007 (under rocks),6imm. 1♂ (NCA 2007/2814); Same locality, Doornkraal Camp,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-27.51225">27°30.735'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="31.20385">31°12.231'E</geoCoordinate>
, leg. C. Haddad &amp; R. Fourie, 29.VI.2007 (sifting leaf litter), 14imm. 2♀ (NCA 2007/2875); Kosi Bay Nature Reserve,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-26.962784">26°57.767'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="32.81635">32°48.981'E</geoCoordinate>
, leg. C. Haddad, 15.IV.2006 (leaf litter, coastal forest), 1imm. 1♂ (NCA 2006/757); Ophathe Game Reserve, Montane grassland,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-28.4224">28°25.344'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="31.399282">31°23.957'E</geoCoordinate>
, 897m a.s.l., leg. C. Haddad, 4.X.2008 (sifting leaf litter), 6imm. 1♂ (NCA 2008/3900); Tembe
<pageBreakToken pageId="22" pageNumber="89" start="start">Elephant</pageBreakToken>
Park,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-27.016666">27°01'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="32.4">32°24'E</geoCoordinate>
, leg. C. Haddad, 5.I.2002 (leaf litter, deep sand forest), 2♂ 1♀ (NCA 2002/523). Limpopo Province: Klein Kariba, near Warmbaths [Bela-Bela],
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-24.866667">24°52'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="28.333334">28°20'E</geoCoordinate>
, 1140m a.s.l., leg. A. Leroy, 27.XI.1996, 1♀ (NCA 2004/830); Little Leigh,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.9485">22°56.910'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="29.869617">29°52.177'E</geoCoordinate>
, leg. I. Sinthumule, 22.XI.2005 (gallery forest), 1♂ (NCA 2009/2041); Same locality, leg. B. van der Waal, 22.XI.2005 (gallery forest), 2♂ (NCA 2010/3324); Mabula Lodge, near Warmbaths [Bela-Bela],
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-24.833334">24°50'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="27.95">27°57'E</geoCoordinate>
, leg. J. Loubser, 16.VII.1989 (running with ants in leaf litter), 1♀ (NCA 91/431); Same locality, leg. J. Leroy, 16.XII.1989 (running with ants in leaf litter), 1♀ (NCA 91/434); Nylsvlei Nature Reserve, Naboomspruit,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-24.65">24°39'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="28.666666">28°40'E</geoCoordinate>
, leg. G. Ferreira, 27.X.1975 (pitfalls), 1♂ (NCA 87/265); Roedtan, between Settlers and Tuinplaas (Springbokvlake), leg. M. van Jaarsveld, 26.III.2003 (pitfall traps, grass), 1♀ (NCA 2003/1336); Soutpansberg Mountains, Lajuma Mountain Retreat, Island 4,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-23.031567">23°01.894'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="29.435383">29°26.123'E</geoCoordinate>
, leg. M. Mafadza, 5.XII.2004 (woodland litter sifting), 1♂ 2♀ (NCA 2006/961); Same locality, Island 4, 23°01.894'S 29°26.123'E, leg. M. Mafadza, 28.XI.2004 (sifting leaf litter, sample 2), 1♀ (NCA 2005/1888); Same locality, leg. M. Mafadza, 28.XI.2004 (active searching), 1♂ (NCA 2005/2025); Same locality, leg. C. Haddad, 6.II.2008 (base of grass tussocks), 4♀ (NCA 2008/508); Tshulu,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-22.583334">22°35'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="30.8">30°48'E</geoCoordinate>
, leg. S. Foord, 21.II.2008 (pitfall traps), 1♀ (NCA 2008/2876). Mpumulanga Province: Nelspruit, Agricultural College,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-25.35">25°21'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="31.766666">31°46'E</geoCoordinate>
, leg. P. Stephen, 22.XII.1998 (pitfall traps, grapefruit orchard), 1♀ (NCA 99/133); Same locality, leg. P. Stephen, 12.XI.1999 (pitfall traps, citrus orchard), 2♂ 1♀ (NCA 2000/184). North West Province: Magaliesberg, Hartebeespoort,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-25.716667">25°43'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="27.833334">27°50'E</geoCoordinate>
, leg. A. Honiball, 10.XII.2008, 1♀ (NCA 2011/837). ZAMBIA: Kafue National Park, Near Namwala, Chibila Camp,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-15.7772665">15°46.636'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="26.00675">26°00.405'E</geoCoordinate>
, leg. C. Haddad &amp; J. Parau, 7.XII.2006 (leaf litter), 1♂ 1♀ (NCA 2007/572); Wildlives Game Farm, near Choma, Hunter's Camp,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-16.982616">16°58.957'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="26.616217">26°36.973'E</geoCoordinate>
, leg. C. Haddad, J. Parau &amp; F. Jordaan, 3.XII.2006 (leaf litter), 1imm. 5♀ (NCA 2007/474); Same locality, Campsite,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-17.05">17°03'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="26.5">26°30'E</geoCoordinate>
, leg. F. Nyathi, 9-14.XII.1994 (drift fence pitfall trap), 1♂ (NMZ 11896). ZIMBABWE: Sengwa Wildlife Research Area,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-18.166666">18°10'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="28.233334">28°14'E</geoCoordinate>
, leg. M.S. Cumming, 15.I.2007, 1♂ (NCA 2007/1307).
</paragraph>
<caption pageId="22" pageNumber="89">
<paragraph pageId="22" pageNumber="89">
Figures 44-49. Scanning electron microscope photographs of
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida dippenaarae" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="dippenaarae">Cambalida dippenaarae</taxonomicName>
sp. n.female (44-46) and male (47-49) spinneret morphology: 44, 47 anterior lateral spinneret 45, 48 posterior median spinneret 46, 49 posterior lateral spinneret. Abbreviations: Ac aciniform gland spigot(s) Cy cylindrical gland spigot(s) MAmp major ampullate gland spigot(s) mAmp minor ampullate gland spigot(s) n nubbin(s) Pi piriform gland spigot(s) ta tartipore.
</paragraph>
</caption>
<caption pageId="22" pageNumber="89">
<paragraph pageId="22" pageNumber="89">
Figures 50-56. Digital microscope photographs of emboli of Afrotropical
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="genus">Cambalida</taxonomicName>
species in ventral view: 50
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida compressa" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="compressa">Cambalida compressa</taxonomicName>
sp. n. 51
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida coriacea" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="coriacea">Cambalida coriacea</taxonomicName>
Simon, 1909 52
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida deminuta" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="deminuta">Cambalida deminuta</taxonomicName>
(Simon, 1909) 53
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida dippenaarae" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="dippenaarae">Cambalida dippenaarae</taxonomicName>
sp. n. 54
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida fulvipes" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="fulvipes">Cambalida fulvipes</taxonomicName>
(Simon, 1896) 55
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida griswoldi" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="griswoldi">Cambalida griswoldi</taxonomicName>
sp. n. 56
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida loricifera" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="loricifera">Cambalida loricifera</taxonomicName>
(Simon, 1885). Scale bars = 0.1mm.
</paragraph>
</caption>
<caption pageId="22" pageNumber="89">
<paragraph pageId="22" pageNumber="89">
Figures 57-60. Genitalic morphology of
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida compressa" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="compressa">Cambalida compressa</taxonomicName>
sp. n.: 57 female epigyne, ventral view 58 same, dorsal view 59 male palp, ventral view 60 same, retrolateral view. Scale bars = 0.25mm.
</paragraph>
</caption>
<caption pageId="22" pageNumber="89">
<paragraph pageId="22" pageNumber="89">
Figures 61-64. Genitalic morphology of
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida coriacea" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="coriacea">Cambalida coriacea</taxonomicName>
Simon, 1909: 61 female epigyne, ventral view 62 same, dorsal view 63 male palp, ventral view 64 same, retrolateral view. Scale bars = 0.25mm.
</paragraph>
</caption>
<caption pageId="22" pageNumber="89">
<paragraph pageId="22" pageNumber="89">
Figure 65. Distribution of
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida compressa" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="compressa">Cambalida compressa</taxonomicName>
sp. n. (open circles) and
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida coriacea" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="coriacea">Cambalida coriacea</taxonomicName>
Simon, 1909 (stars) in the Afrotropical Region.
</paragraph>
</caption>
<caption pageId="22" pageNumber="89">
<paragraph pageId="22" pageNumber="89">
Figures 66-69. Genitalic morphology of
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida deminuta" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="deminuta">Cambalida deminuta</taxonomicName>
(Simon, 1909): 66 female epigyne, ventral view 67 same, dorsal view 68 male palp, ventral view 69 same, retrolateral view. Scale bars = 0.25mm.
</paragraph>
</caption>
<caption pageId="22" pageNumber="89">
<paragraph pageId="22" pageNumber="89">
Figures 70-73. Genitalic morphology of
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida dippenaarae" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="dippenaarae">Cambalida dippenaarae</taxonomicName>
sp. n.: 70 female epigyne, ventral view 71 same, dorsal view 72 male palp, ventral view 73 same, retrolateral view. Scale bars = 0.25mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="22" pageNumber="89" type="diagnosis">
<paragraph pageId="22" pageNumber="89">Diagnosis.</paragraph>
<paragraph pageId="22" pageNumber="89">
Females are closely related to
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida fulvipes" order="Araneae" pageId="22" pageNumber="89" phylum="Arthropoda" rank="species" species="fulvipes">Cambalida fulvipes</taxonomicName>
but can be recognised by the subtriangular rather than round ST II and the narrower epigynal ridges (compare Fig. 70 with Fig. 80). Males can be recognised by the nearly parallel-sided basal section and the narrowly coiled distal section of the embolus.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="89" type="etymology">
<paragraph pageId="22" pageNumber="89">Etymology.</paragraph>
<paragraph pageId="22" pageNumber="89">The species name is a patronym in honour of Ansie Dippenaar-Schoeman, in recognition of her contributions to the study and promotion of research on African arachnids.</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="90" pageId="22" pageNumber="89" type="female">
<paragraph pageId="22" pageNumber="89">Female</paragraph>
<paragraph pageId="22" pageNumber="89">
(holotype, Livingstone, NCA 2007/625). Measurements: CL 2.53, CW 1.85, AL 3.15, AW 2.07, TL 5.75 (4.90-5.95), FL 0.22, SL 1.16, SW 1.11,
<normalizedToken originalValue="AMEAME">AME-AME</normalizedToken>
0.06,
<normalizedToken originalValue="AMEALE">AME-ALE</normalizedToken>
0.02,
<normalizedToken originalValue="ALEALE">ALE-ALE</normalizedToken>
0.35,
<normalizedToken originalValue="PMEPME">PME-PME</normalizedToken>
0.11,
<normalizedToken originalValue="PMEPLE">PME-PLE</normalizedToken>
0.04,
<normalizedToken originalValue="PLEPLE">PLE-PLE</normalizedToken>
0.53, PERW 0.83, MOQAW 0.33, MOQPW 0.49, MOQL 0.48.
</paragraph>
<paragraph pageId="22" pageNumber="89">Length of leg segments (sequence from femur to tarsus, and total): I 1.78 + 0.80 + 1.50 + 1.40 + 0.95 = 6.43; II 1.63 + 0.74 + 1.28 + 1.25 + 0.84 = 5.74; III 1.48 + 0.71 + 1.13 + 1.36 + 0.68 = 5.36; IV 2.18 + 0.88 + 2.03 + 2.28 + 0.92 = 8.29.</paragraph>
<paragraph pageId="23" pageNumber="90">
<pageBreakToken pageId="23" pageNumber="90" start="start">Carapace</pageBreakToken>
deep red-brown with black mottling, clypeus dark brown medially, yellow-brown laterally, eye region nearly black; faint black striae radiating from fovea towards palps and leg coxae; surface granulate, sparsely covered in white plumose setae. All eyes with black rings; AER procurved, ALE much larger than AME; AME separated by distance slightly less than
<normalizedToken originalValue="½">1/2</normalizedToken>
their diameter, AME separated from ALE by 16 AME diameter; clypeus height slightly larger than AME diameter; PER procurved, PLE very slightly larger than PME; PME separated by distance slightly less than 35 their diameter, PME separated from PLE by distance equal to 15 PLE diameter; CW:PERW = 2.23:1. Chelicerae dark orange-brown, orange proximally and along prolateral distal margin; three teeth on promargin, median tooth largest, distal tooth slightly smaller than proximal tooth, distal tooth situated closest to median tooth; two teeth on retromargin, distal tooth slightly larger than proximal tooth, closer to fang base than promarginal teeth; endites dark brown, fading to yellow and cream prolaterally; labium dark brown, creamy-yellow distally; sternum deep orange-brown with dark brown mottling. Legs finely granulate; femora
<normalizedToken originalValue="IIV">I-IV</normalizedToken>
dark brown, with slightly paler lines dorsally, bright yellow dorsally at distal end; patellae
<normalizedToken originalValue="IIII">I-III</normalizedToken>
yellow with faint black mottling ventrally, black around patellar indentation; patellae IV yellow-orange with faint black mottling laterally and ventrally, black around patellar indentation; tibiae, metatarsi and tarsi
<normalizedToken originalValue="IIII">I-III</normalizedToken>
bright yellow-orange; tibiae IV orange with dense black mottling, except proximally and distally; metatarsi IV orange with faint black mottling; tarsi IV yellow with faint black mottling. Leg spination: femora: I pl 1 do 3, II do 3, III pl 1 do 3 rl 1, IV pl 1 do 3 rl 1; patellae with do 1 long distal seta; tibiae: I and II spineless, III pl 2 rl 2 plv 2 vt 2, IV pl 2 do 1 rl 2 plv 2 vt 2; metatarsi: I plv 2 rlv 2, II plv 2 rlv 2, III pl 2 rl 2 plv 2 rlv 2 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3. Palpal spination: femora do 2, patellae pl 1 spine do 2 setae, tibiae pl 1 do 1 plv 1, tarsi pl 1 plv 3 rlv 1. Abdomen dark grey, with dark red-brown dorsal scutum extending slightly more than
<normalizedToken originalValue="¼">1/4</normalizedToken>
abdomen length, with fine cream chevrons posteriorly and small white spot of dense plumose setae just above spinnerets; venter mottled creamy-grey, slightly darker towards spinnerets; epigastric scutum bright yellow-orange with black mottling, inframamillary sclerite yellow-brown. Epigyne with lateral copulatory openings situated within small curved epigynal ridges (Fig. 70); copulatory ducts initially directed anterolaterally, looping posteriorly, then transversely and anteriorly, entering ST II posteromedially; ST II somewhat triangular, with sharply angled lateral margins, joined narrowly to kidney-shaped posterior ST I (Fig. 71).
</paragraph>
</subSubSection>
<subSubSection lastPageId="24" lastPageNumber="91" pageId="23" pageNumber="90" type="male">
<paragraph pageId="23" pageNumber="90">Male</paragraph>
<paragraph pageId="23" pageNumber="90">
(paratype, Livingstone, NCA 2007/625). Measurements: CL 2.40, CW 1.75, AL 2.90, AW 1.48, TL 5.45 (4.47-5.45), FL 0.19, SL 1.11, SW 0.99,
<normalizedToken originalValue="AMEAME">AME-AME</normalizedToken>
0.06,
<normalizedToken originalValue="AMEALE">AME-ALE</normalizedToken>
0.02,
<normalizedToken originalValue="ALEALE">ALE-ALE</normalizedToken>
0.31,
<normalizedToken originalValue="PMEPME">PME-PME</normalizedToken>
0.10,
<normalizedToken originalValue="PMEPLE">PME-PLE</normalizedToken>
0.06,
<normalizedToken originalValue="PLEPLE">PLE-PLE</normalizedToken>
0.51, PERW 0.76, MOQAW 0.30, MOQPW 0.43, MOQL 0.44.
</paragraph>
<paragraph pageId="23" pageNumber="90">Length of leg segments (sequence from femur to tarsus, and total): I 1.85 + 0.70 + 1.65 + 1.60 + 1.10 = 6.90; II 1.64 + 0.68 + 1.30 + 1.40 + 0.90 = 5.92; III 1.52 + 0.66 + 1.15 + 1.45 + 0.77 = 5.55; IV 2.30 + 0.83 + 2.11 + 2.55 + 1.06 = 8.85.</paragraph>
<paragraph lastPageId="24" lastPageNumber="91" pageId="23" pageNumber="90">
Carapace dark red-brown with black mottling, clypeus slightly paler laterally, eye region darker; faint black striae radiating from fovea towards palps and leg coxae; surface finely granulate, densely covered in white plumose setae. All eyes with black rings;
<pageBreakToken pageId="24" pageNumber="91" start="start">AER</pageBreakToken>
procurved, ALE much larger than AME; AME separated by distance slightly less than
<normalizedToken originalValue="½">1/2</normalizedToken>
their diameter, AME separated from ALE by distance equal to ⅛ AME diameter; clypeus height slightly larger than
<normalizedToken originalValue="1½">11/2</normalizedToken>
AME diameter; PER procurved, PLE very slightly larger than PME; PME separated by distance slightly more than 35 their diameter, PME separated from PLE by distance equal to ⅓ PME diameter; CW:PERW = 2.30:1. Chelicerae brown with black mottling on anterior surface, paler proximally, yellow along prolateral distal margin; three teeth on promargin, median tooth largest, distal tooth smaller than proximal tooth, distal tooth situated closest to median tooth; two slightly separated subequal teeth on retromargin, closer to fang base than promarginal teeth; endites dark yellow-brown with dark brown mottling, fading to yellow and cream prolaterally; labium orange-brown proximally, creamy-yellow distally; sternum deep orange-brown with dense black mottling. Legs finely granulate; femora I and II pale yellow-brown, yellow distally; femora III and IV dark orange-brown, yellow distally; patellae I and II yellow, and III and IV yellow-orange, all with faint black lateral mottling, black around patellar indentation; tibiae, metatarsi and tarsi I and II yellow with faint black mottling laterally; tibiae, metatarsi and tarsi III yellow-orange with faint black mottling laterally; tibiae IV deep orange with black mottling, yellow distally; metatarsi IV yellow-orange with black mottling, absent proximally and distally; tarsi IV yellow. Leg spination: femora: I pl 1 do 3, II do 3, III pl 1 do 3 rl 1, IV pl 1 do 3 rl 1; patellae with do 1 long distal seta; tibiae: I plv 0-1, II rlv 0-1, III pl 2 rl 2 plv 2 vt 2, IV pl 2 do 1 rl 2 plv 2 vt 2; metatarsi: I plv 2 rlv 2, II plv 2 rlv 2, III pl 2 rl 2 plv 1 rlv 1 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3. Palpal spination: femora do 2, patellae pl 1 spine do 2 setae, tibiae pl 1 plv 1, tarsi pl 2 plv 2. Abdomen with deep red-brown, nearly black, dorsal scutum covering entire dorsum, with small white spot of dense plumose setae just above spinnerets; sides of abdomen mottled dark grey; venter pale grey, epigastric sclerite, post-epigastric sclerites, ventral sclerite and inframamillary sclerite red-brown with black mottling. Palps yellow-brown with black mottling, cymbium orange-brown; embolus with parallel-sided basal section, distal section of coil transverse, bent at right angle to distally-directed tip (Figs 53, 72); embolic coil relatively narrow (Fig. 73).
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="91" type="distribution">
<paragraph pageId="24" pageNumber="91">Distribution.</paragraph>
<paragraph pageId="24" pageNumber="91">
Widespread throughout southern Africa (Fig. 79); largely sympatric with
<taxonomicName class="Arachnida" family="Corinnidae" genus="Cambalida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambalida fulvipes" order="Araneae" pageId="24" pageNumber="91" phylum="Arthropoda" rank="species" species="fulvipes">Cambalida fulvipes</taxonomicName>
across its range (Fig. 95).
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="91" type="biology">
<paragraph pageId="24" pageNumber="91">Biology.</paragraph>
<paragraph pageId="24" pageNumber="91">A fairly common ground-dwelling spider collected mainly by litter sifting and pitfall traps in forest and savanna habitats. Occasionally collected from the Grassland Biome in South Africa.</paragraph>
</subSubSection>
</treatment>
</document>