257 lines
31 KiB
XML
257 lines
31 KiB
XML
<document ID-CLB-Dataset="24300" ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="862098811AD37CD4171CB6D7174F1C33" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole boribora Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="EDD6BF4F-581F-4246-B931-502AB27EC1BB" docUuidSource="ZooBank" docVersion="7" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1732649601836" updateUser="ExternalLinkService">
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<mods:titleInfo id="EA79375359514884D36D7BE396ABEF8F">
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<mods:title id="02F7A5F7084A2E326624CE08B91FC24F">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:namePart id="E8AE1211CDB10323152C1B6F0DDE3E5B">Salata, Sebastian</mods:namePart>
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<mods:namePart id="96C009519EFA51E4503EA0EEE73B73A9">Fisher, Brian L.</mods:namePart>
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<mods:typeOfResource id="2FA1182448645BAE70D79CBA43EC82BB">text</mods:typeOfResource>
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<mods:title id="A9781CAF4929F79CEE184ACA1C75666B">ZooKeys</mods:title>
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<mods:part id="6B5F1A0E75E97EBE4EB778FC8B3F5DDC">
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<mods:date id="8FF81A968DA79E2754AB44140E602153">2020</mods:date>
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<mods:number id="357E4CC527396C7234ED26C2155B1A04">905</mods:number>
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<mods:end id="080AE779BBEDAEB61DB38F5FADA5B0BA">235</mods:end>
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<mods:url id="E75BB9EEAF70BB3572E35671D153670A">http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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<mods:classification id="58DB68010EABF9CB4110E88FE277C0F7">journal article</mods:classification>
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<mods:identifier id="10842BE22174F1BD9EDF50A5BB8C8383" type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier id="B54FA5D5D582372F6C923C50F8918E24" type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<treatment id="862098811AD37CD4171CB6D7174F1C33" ID-GBIF-Taxon="161777020" LSID="urn:lsid:zoobank.org:act:EDD6BF4F-581F-4246-B931-502AB27EC1BB" httpUri="http://treatment.plazi.org/id/862098811AD37CD4171CB6D7174F1C33" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="13F30B478BF7B4E4D1EAA815D7323D31" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="01E87B46F73AC12A70BDB7169ABD572D" pageId="0" pageNumber="1">
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||
<taxonomicName id="01D00ECDA6F64B0936C57CEC1D3D45B8" ID-CoL="CMKX7" LSID="http://zoobank.org/EDD6BF4F-581F-4246-B931-502AB27EC1BB" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole boribora" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="boribora" status="sp. nov.">Pheidole boribora</taxonomicName>
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<taxonomicNameLabel id="C3B0083A3440D158DC9E79E786C60DA7" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="93B65A8CE80F4F423658EAC35B451C8D" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">Figs 73A-F</figureCitation>
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<figureCitation id="CD79A0A32BEE53DE4F22BC64737B70F9" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84J</figureCitation>
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<figureCitation id="1CAE89106F42B2E4C6CDFBDA00C97DE3" captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86J</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="E940025A1592B6F19655EB79EF90695B" pageId="0" pageNumber="1" type="type material">
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<paragraph id="4DBB5B7C86D24832AFCD08F10D5C9E43" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="DDB5A8F5329AE502DC947D4F31D77E86" pageId="0" pageNumber="1">
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<emphasis id="FDF8A0CF39813DE8A6DFA5DBEDB964F6" bold="true" pageId="0" pageNumber="1">
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<emphasis id="38904358F1E23BC007B77C36A93A2F18" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Toliara; Parc National
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<normalizedToken id="851FF7D1746BD68F09B5BBC684392176" originalValue="d’Andohahela">d'Andohahela</normalizedToken>
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, Col du Sedro, 3.8 km 113°ESE Mahamavo, 37.6 km 341°NNW Tolagnaro;
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<geoCoordinate id="53B94AA6AD825EDB310B55D883A2E8EC" degrees="24.76389" direction="south" orientation="latitude" precision="1" value="-24.76389">-24.76389</geoCoordinate>
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,
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<geoCoordinate id="050E2B5DDE213B60354B163E2DACDA62" degrees="46.75167" direction="east" orientation="longitude" precision="1" value="46.75167">46.75167</geoCoordinate>
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; alt. 900 m; 21 Jan 2002; B.L. Fisher et al. leg.; BLF05130, CASENT0456013 (CASC).
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<emphasis id="C52F041089E3D436F69D4500ED148AC1" bold="true" pageId="0" pageNumber="1">
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<emphasis id="0001965DCF6DFBA1B86BFF794577C579" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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Madagascar. •4w.; same data as for holotype; CASENT0456014-CASENT0456016, CASENT0872187-CASENT0872190 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="1A6FF19BC5D14DF6864AD089AAA6ECC5" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="94ADFC4DDE4908EA8DAAD6877EFE5BCE" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="98CF1BA386900A664778BD8320B393CA" pageId="0" pageNumber="1">
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<emphasis id="10C83449D1B942F45BF962E0BC0F0067" bold="true" pageId="0" pageNumber="1">
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<emphasis id="71D4DA31B89405190DBA8F64FA401400" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Head in full-face view oval, relatively as long as wide, anterior and posterior of eyes moderately convex; sides of the head with moderately dense, long, erect pilosity; genae smooth and shiny; inner hypostomal teeth distinct, distinct, low, thick, and triangular, with rounded apex; outer hypostomal teeth approximately the same size as inner hypostomal teeth, thick, dentate, with rounded tips; propodeal spines small, triangular; first gastral tergite finely shagreened; body reddish brown.
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<emphasis id="B99C4A36233BDFABDAF2E75BF67B0F30" bold="true" pageId="0" pageNumber="1">
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<emphasis id="91AF7BD1BFA9B638C11CD794997AE09F" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Head foveolate; promesonotum low, short, slightly convex, with relatively steep posterior declivity; propodeal spines small; mesosoma foveolate, foveolate, mesonotum, anepisternum, katepisternum and lateral surfaces of propodeum smooth; body yellow.
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</paragraph>
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</subSubSection>
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<subSubSection id="CE1B743B8389382392B79537B9898B2A" pageId="0" pageNumber="1" type="description">
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<paragraph id="842F07AF9992AC091B493DCC23BC69B3" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="C6D460B565134285623FC3F500F07941" pageId="0" pageNumber="1">
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<emphasis id="ADECF89E389CC88139339CA6DFBC8B3E" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis id="AEA4279D56D89E8B66A10FE60A74E2F7" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 1): HL: 1.5; HW: 1.4; SL: 0.66; EL: 0.17; WL: 1.07; PSL: 0.17; MTL: 0.66; PNW: 0.58; PTW: 0.19; PPW: 0.41; CI: 93.4; SI: 47.2; PSLI: 11.2; PPI: 45.5; PNI: 41.6; MTI: 46.9.
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<emphasis id="72BF1621D7074FF12A41A764B6DB4AB1" bold="true" pageId="0" pageNumber="1">
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<emphasis id="4D2AA46610AC01978C6074680B2C1DEB" italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view oval, anterior and posterior of eyes moderately convex (Fig.
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<figureCitation id="2278D05A445E35C2E603CDECEA981FB6" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73B</figureCitation>
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). In lateral view sub-rectangular; ventral and dorsal faces finely convex; dorsal face finely depressed posteriorly, forming shallow transverse depression between frons and occipital lobes; inner hypostomal teeth invisible. Sides of the head with moderately dense, long, erect pilosity; whole head with moderately dense, short, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae. Occipital lobes shiny, with thick, sparse, irregular rugae, interspaces smooth to finely foveolate; frons with thick, moderately dense, longitudinal rugae, interspaces smooth to finely foveolate; lateral sides of head foveolate, with additional thin, sparse, longitudinal rugae; malar area with dense and thin longitudinal rugulae; genae smooth and shiny. Clypeus shiny and smooth, with thin, longitudinal rugulae on the lateral sides; median notch present, narrow, and moderately deep; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, reaching the midlength of head; pilosity decumbent to erect (Fig.
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<figureCitation id="1C033D4951D80F5953BFB6E0F9633526" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73B, D</figureCitation>
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). Inner hypostomal teeth distinct, low, thick, and triangular, with rounded apex; outer hypostomal teeth approximately the same size as inner hypostomal teeth, thick, dentate, with rounded tips (Fig.
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<figureCitation id="C4DDC3F56DA95447516653F54F2B454E" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84J</figureCitation>
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).
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<emphasis id="3FCA5D09F328EC0C911DB75CDC26914A" bold="true" pageId="0" pageNumber="1">
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<emphasis id="1DEA4B08A18C7CA33C6BD0689C3ED046" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum short, angular, and relatively low, posterior mesonotum steep, with moderately large, tubercle-like projections; promesonotal groove absent; metanotal groove absent or indistinct; propodeal spines small, triangular, with base wide, apex rounded; humeral area with small and flat tubercles (Fig.
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<figureCitation id="FDD8242DCD531546324B7BF3516F3C8C" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73D</figureCitation>
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). Surface shiny, with very sparse, transverse to irregular thin rugae, pronotum and propodeum with additional indistinct and sparse foveolae. Pilosity moderately dense, long, and erect (Fig.
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<figureCitation id="83543FE051458BC3816889F9A4CC87FC" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73D, F</figureCitation>
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).
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<emphasis id="67CD16A954D14E5B51F20BD33BFC5F85" bold="true" pageId="0" pageNumber="1">
|
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<emphasis id="5C5556CB4F8EF16CAECA65EF872BBE79" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny, finely shagreened to smooth; peduncle short, with small, rounded, horizontal lobes on its basal part; node moderately high and triangular, with convex apex, in rear view node relatively straight; pilosity moderately sparse and erect (Fig.
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<figureCitation id="35DACAFB0C0EAD5521EE6F08C9BDD3D6" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73D, F</figureCitation>
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).
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||
<emphasis id="64B3B3CCCE66565BE4565E25262EA4E1" bold="true" pageId="0" pageNumber="1">
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<emphasis id="08CDDFC52D02157E75BD67F443F8E9A5" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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||
</emphasis>
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Shiny and finely shagreened; short and rounded; in dorsal view sides with short, acute, triangular projections; pilosity long, moderately dense, and erect (Fig.
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<figureCitation id="0C15A14E9CB94E7DCCF9B33A7C73D938" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73D, F</figureCitation>
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||
).
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<emphasis id="8CE7BCCF2B2EAC84F5E8BB7759FCBCE3" bold="true" pageId="0" pageNumber="1">
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<emphasis id="8A998CC064F24439DE104FF59AD52A75" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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First gastral tergite shiny and finely shagreened; pilosity dense, long and erect (Fig.
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<figureCitation id="3672E481EEFF172A45721CD6D78FB773" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73D, F</figureCitation>
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).
|
||
<emphasis id="83A5AE2EED6E09CC453B12E762FCCFEE" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="9118DAD9EB938A8FA24B1249F7E91D1C" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Head reddish brown; mesosoma and gaster yellowish brown to brown; legs dark yellow (Fig.
|
||
<figureCitation id="500DFFCB9BD1BDBB2DB6F3314F1B9118" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="486D6A68BC43EF5B462D358CD7C59A9A" doi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1" start="Figure 73" startId="F73">
|
||
<paragraph id="5B07E6C76FDFFF0DC9C9674AC665BE52" pageId="0" pageNumber="1">
|
||
<emphasis id="E4CF330AD44111A62A2501893DCF2ED2" bold="true" pageId="0" pageNumber="1">Figure 73.</emphasis>
|
||
<taxonomicName id="6F0C1377656B6108A9F62DA317501D16" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole boribora" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="boribora">
|
||
<emphasis id="A8CF4BED4D8C3068B01E267B48F2FDB0" italics="true" pageId="0" pageNumber="1">Pheidole boribora</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., full-face view (
|
||
<emphasis id="91A83904EDF040ED434456D805A75940" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
), profile (
|
||
<emphasis id="61C7B48FFDFF70644B556A84FC55CCDE" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
), and dorsal view (
|
||
<emphasis id="19DE931ABD57996711080F0DAF0A3A62" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
) of paratype minor worker (CASENT0456014) and full-face view (
|
||
<emphasis id="5B437C5A567541553FABB925A78F538B" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
), profile (
|
||
<emphasis id="11BC53B148AD82144ABB503B40A13D84" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
), and dorsal view (
|
||
<emphasis id="32D84054116DDC12B221C4A5EFAFC6F4" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
) of holotype major worker (CASENT0456013).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C766ED1272E4D87979752B307A46595C" pageId="0" pageNumber="1">
|
||
<emphasis id="1DE38B10F01ACE1C8C37F627CF6165D2" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="4CDF664638FE30C255231B0C43273688" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 3): HL: 0.55-0.58 (0.56); HW: 0.49-0.5 (0.49); SL: 0.57-0.58 (0.57); EL: 0.11-0.11 (0.11); WL: 0.68-0.76 (0.74); PSL: 0.07-0.1 (0.08); MTL: 0.42-0.46 (0.43); PNW: 0.31-0.32 (0.31); PTW: 0.06-0.08 (0.07); PPW: 0.12-0.14 (0.13); CI: 85.8-89.4 (87.5); SI: 116.2-116.8 (116.4); PSLI: 12.0-17.0 (14.6); PPI: 46.0-59.8 (53.3); PNI: 62.9-64.2 (63.5); MTI: 85.0-91.2 (88.0).
|
||
<emphasis id="3CC745816AE24FE0A322D3A079D21F9F" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="488863343C6C4EE912802FEB64C7033A" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
Occipital margin straight or indistinctly concave; occipital carina indistinct, weakly developed (Fig.
|
||
<figureCitation id="24F1876F66096EBF9560DE71531EFDD5" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73A</figureCitation>
|
||
). Pilosity moderately dense, long, erect. Whole head foveolate. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity erect (Fig.
|
||
<figureCitation id="26E4AEAFDA4774EE629666B63D1DD946" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73A, C</figureCitation>
|
||
).
|
||
<emphasis id="309CB468ABC21AB7866191BB20659601" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="86A4AB80D076F3258CBBE32E78D9F376" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
In lateral view, promesonotum low, short, slightly convex, with relatively steep posterior declivity; promesonotal groove absent; metanotal groove absent; propodeal spines small, triangular, apex acute (Fig.
|
||
<figureCitation id="8D5AEA194CA624994122D55CA07D61A5" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73C</figureCitation>
|
||
). Sculpture foveolate; katepisternum, anepisternum and mesonotum smooth. Pilosity sparse, long, and erect (Fig.
|
||
<figureCitation id="F7F0515BB1B5A79A4C1C012C26CB2F3F" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73C, E</figureCitation>
|
||
).
|
||
<emphasis id="42690524478D79D48ADB7AB1FE4E18F2" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="67F99ACFA982598559C17215B56F97BD" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Peduncle short and thin with ventral face slightly convex; node low, triangular, and small; with few moderately long, erect setae (Fig.
|
||
<figureCitation id="A7BFC16A19C82AD6AEA697F9CDD06D55" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73C, E</figureCitation>
|
||
).
|
||
<emphasis id="D1CC892650E71DD3CC1FDB7AAB9EF710" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="06E4BAC218C508FF8B31D77D84ADC0D7" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Short, low, and convex; with few moderately long, erect setae (Fig.
|
||
<figureCitation id="0D6F233C953131C4B526ABF92D5FAB06" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73C, E</figureCitation>
|
||
).
|
||
<emphasis id="5E8C5327B611E92BAB710F5E9F87B754" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="D6381446F188F8A2A7DA1B489DE53A16" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
With moderately sparse, erect pilosity (Fig.
|
||
<figureCitation id="81F4C3640EC7D7C9335AB723E4E2BE7F" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73C, E</figureCitation>
|
||
).
|
||
<emphasis id="6170CF917C64097D0E698A77EFA665B4" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="0D08DB226755CB92A99A0FF076B2E9C6" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Unicolourous, yellow (Fig.
|
||
<figureCitation id="1E395E3189F57A0E79CDB967739BB6FC" captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Pheidole boribora sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0456014) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0456013)." figureDoi="10.3897/zookeys.905.39592.figure73" httpUri="https://binary.pensoft.net/fig/373120" pageId="0" pageNumber="1">73C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E646357674B6A3C5E758147AA38CBEFB" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="77B53BABE45ED39F53E7867738A9C2A7" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="476AE6E493FD7F03B6F885ABA880D0DF" pageId="0" pageNumber="1">Malagasy for oval, in reference to the head shape of major workers.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B272E689DAE15F129D2D2D5D4C22A15F" pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph id="85DCE9CF5157B7BEEAE5A8F8D48D4477" pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph id="3E1C09EBA8E8B4F09ECC3A3C6C79595D" pageId="0" pageNumber="1">The species was collected at 900 m in elevation, in montane rainforest. Nest was located in a rotten log.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13E4CE5EDF631330F5588D70BEC69E3C" pageId="0" pageNumber="1" type="comments">
|
||
<paragraph id="14706505F5F3BE98BA5AA5EBFF0A3663" pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph id="73A072DD5FF6EFF84A2AEC79800C7419" pageId="0" pageNumber="1">
|
||
<emphasis id="FE76D59A6F6F957CC005D06A5A154D06" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="14BDC98A1F3D813B13A9FB4A217CE140" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="55F9F0D422476364036DC20BB074D8D4" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole boribora" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="boribora">
|
||
<emphasis id="D8FD2A5EBD69ED72744B4E99B3D0A64E" italics="true" pageId="0" pageNumber="1">Pheidole boribora</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most similar to
|
||
<taxonomicName id="EA5842D18B4931DFF1C651B8A97C9B48" lsidName="P. miramila" pageId="0" pageNumber="1" rank="species" species="miramila">
|
||
<emphasis id="D3763E34DCBDE756B705224282A0D322" italics="true" pageId="0" pageNumber="1">P. miramila</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and differs from it by presence of foveolae on head, dense and long pilosity of sides of head, dentate inner hypostomal teeth, and small propodeal spines.
|
||
<emphasis id="5217B9FCBA41BCD6270BCA999E196437" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="B9CE1509C326F766F8CF8F4DC822AE99" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="9CC274C7C2A668E3D6049D7B8BDBAFCE" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole boribora" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="boribora">
|
||
<emphasis id="E3015EA1C3DB2F944315896057E0CE6B" italics="true" pageId="0" pageNumber="1">Pheidole boribora</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most similar to
|
||
<taxonomicName id="6FA8BDD62086BA201A526FB8AFC242C9" lsidName="P. petax" pageId="0" pageNumber="1" rank="species" species="petax">
|
||
<emphasis id="14361FCCFF595B42BF0FA19EC719C88A" italics="true" pageId="0" pageNumber="1">P. petax</emphasis>
|
||
</taxonomicName>
|
||
and differs from it by smooth mesonotum, anepisternum, and lateral surfaces of propodeum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |