treatments-xml/data/80/22/75/802275FDE49A519D985FCA1C85E4DBF6.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1150.93760" ID-Pensoft-Pub="1313-2970-1150-1" ID-Pensoft-UUID="2D0F861C78665B9BABB241437CA5ED53" ID-ZooBank="A38C5FB69F664F858788AAA53D21704D" ModsDocID="1313-2970-1150-1" checkinTime="1677664009133" checkinUser="pensoft" docAuthor="Crews, Sarah C." docDate="2023" docId="802275FDE49A519D985FCA1C85E4DBF6" docLanguage="en" docName="ZooKeys 1150: 1-189" docOrigin="ZooKeys 1150" docPubDate="2023-02-27" docSource="http://dx.doi.org/10.3897/zookeys.1150.93760" docTitle="Karaops keithlongbottomi Crews &amp; Harvey 2011" docType="treatment" docVersion="1" id="2D0F861C78665B9BABB241437CA5ED53" lastPageNumber="1" masterDocId="2D0F861C78665B9BABB241437CA5ED53" masterDocTitle="But wait, there's more! Descriptions of new species and undescribed sexes of flattie spiders (Araneae, Selenopidae, Karaops) from Australia" masterLastPageNumber="189" masterPageNumber="1" pageNumber="1" updateTime="1677664009133" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>But wait, there's more! Descriptions of new species and undescribed sexes of flattie spiders (Araneae, Selenopidae, Karaops) from Australia</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Crews, Sarah C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9360-6236</mods:nameIdentifier>
<mods:affiliation>California Academy of Sciences, Department of Entomology, 55 Music Concourse Drive, San Francisco, CA, 94118, USA</mods:affiliation>
<mods:nameIdentifier type="email">screwsemail@gmail.com</mods:nameIdentifier>
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<mods:date>2023</mods:date>
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<mods:number>2023-02-27</mods:number>
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<mods:number>1150</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1150.93760</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1150.93760</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:802275FDE49A519D985FCA1C85E4DBF6" httpUri="http://treatment.plazi.org/id/802275FDE49A519D985FCA1C85E4DBF6" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="6NK8V" LSID="802275FD-E49A-519D-985F-CA1C85E4DBF6" authority="Crews &amp; Harvey, 2011" authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops keithlongbottomi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keithlongbottomi">Karaops keithlongbottomi Crews &amp; Harvey, 2011</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 17" captionStartId="F24" captionText="Figure 17. Members of the Kimberley species group A Karaops sp., Kelly's Knob, Western Australia B Karaops sp., Kelly's Knob, Western Australia C Karaops sp., Keep River National Park, Northern Territory (sel _ 1320, MAGNT A 004876) D Karaops sp. eating a fly, Kelly's Knob, Western Australia E Karaops keithlongbottomi, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244, WAM T 155620)." figureDoi="10.3897/zookeys.1150.93760.figure17" httpUri="https://binary.pensoft.net/fig/816404" pageId="0" pageNumber="1">Figs 17E</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F25" captionText="Figure 18. Members of the Kimberley species group A Karaops sp., Keep River National Park, Northern Territory (sel _ 1315, MAGNT A 004871) B habitat, Keep River National Park, Northern Territory C Karaops keithlongbottomi, holotype male, Drysdale River Station, Western Australia (WAM T 55002) D Karaops keithlongbottomi, palp, dorsal, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244, WAM T 155620) E Karaops keithlongbottomi, holotype male, palp, dorsal, Drysdale River Station, Western Australia (WAM T 55002) F Karaops dejongi sp. nov., holotype female, RAAF Boab Quarry, Western Australia (sel _ 1273, WAM T 155649) G Karaops umiida, holotype female, Irvine Island, Western Australia (WAM T 110393) H habitat, Keep River National Park, Northern Territory I Karaops keithlongbottomi, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244 WAM T 155620) J same, face showing marks on chelicerae. Scale bars: 0.5 mm (D, E); 1 mm (A, C, F, J); 5 mm (G, I)." figureDoi="10.3897/zookeys.1150.93760.figure18" httpUri="https://binary.pensoft.net/fig/816405" pageId="0" pageNumber="1">, 18C-E</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F25" captionText="Figure 18. Members of the Kimberley species group A Karaops sp., Keep River National Park, Northern Territory (sel _ 1315, MAGNT A 004871) B habitat, Keep River National Park, Northern Territory C Karaops keithlongbottomi, holotype male, Drysdale River Station, Western Australia (WAM T 55002) D Karaops keithlongbottomi, palp, dorsal, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244, WAM T 155620) E Karaops keithlongbottomi, holotype male, palp, dorsal, Drysdale River Station, Western Australia (WAM T 55002) F Karaops dejongi sp. nov., holotype female, RAAF Boab Quarry, Western Australia (sel _ 1273, WAM T 155649) G Karaops umiida, holotype female, Irvine Island, Western Australia (WAM T 110393) H habitat, Keep River National Park, Northern Territory I Karaops keithlongbottomi, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244 WAM T 155620) J same, face showing marks on chelicerae. Scale bars: 0.5 mm (D, E); 1 mm (A, C, F, J); 5 mm (G, I)." figureDoi="10.3897/zookeys.1150.93760.figure18" httpUri="https://binary.pensoft.net/fig/816405" pageId="0" pageNumber="1">, 18I, J</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F26" captionText="Figure 19. Members of the Kimberley species group A Karaops keithlongbottomi, palp, ventral, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244, WAM T 155620) B same, retrolateral C Karaops sp., Mimbi Caves, Western Australia (sel _ 1276, WAM T 155652) D Karaops dejongi sp. nov., paratype male, RAAF Boab Quarry, Western Australia (sel _ 1272, WAM T 155648) E Karaops sp., Geike Gorge, Western Australia (sel _ 1267, WAM T 155643) F habitat, Mimbi Caves, Western Australia. Scale bars: 0.5 mm (A, B); 1 mm (C-E)." figureDoi="10.3897/zookeys.1150.93760.figure19" httpUri="https://binary.pensoft.net/fig/816406" pageId="0" pageNumber="1">, 19A, B</figureCitation>
<figureCitation captionStart="Map 1" captionStartId="F3" captionText="Map 1. Map of Australia showing areas that are expanded in subsequent figures. CD = Central Desert group, D = Karaops dawara group, F = Karaops francesae group, K = Kimberley group, UA = species that are not assigned to a group, PG = Pilbara-Gascoyne group." figureDoi="10.3897/zookeys.1150.93760.map1" httpUri="https://binary.pensoft.net/fig/816383" pageId="0" pageNumber="1">, Maps 1</figureCitation>
<figureCitation captionStart="Map 7" captionStartId="F23" captionText="Map 7. Species of the Kimberley species group. Diamonds (juveniles): light purple = Mimbi Caves; blue = Geike Gorge, green = Adcock Gorge, teal = Augustus Island, pink = Little Mertens Falls, yellow = Emma Gorge, orange = Kelly's Knob, red = Keep River National Park; Circles: green = Karaops dejongi sp. nov., blue = Karaops jenniferae, red = Karaops dalmanyi sp. nov., purple = Karaops sp. adult male (will be described in subsequent publication), white = Karaops conilurus sp. nov., pink = Karaops umiida, yellow = Karaops alanlongbottomi, teal = Karaops keithlongbottomi, mauve = Karaops larryoo, orange = Karaops garyodwyeri sp. nov., black = Karaops malumbu sp. nov. Juveniles are treated as distinct species based on molecular data." figureDoi="10.3897/zookeys.1150.93760.map7" httpUri="https://binary.pensoft.net/fig/816403" pageId="0" pageNumber="1">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops keithlongbottomi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keithlongbottomi">Karaops keithlongbottomi</taxonomicName>
Crews &amp; Harvey, 2011: 34, figs 19-20 (♂, examined).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" location="Western Australia" specimenCount="♂" stateProvince="Western Australia">
<collectingRegion country="Australia" name="Western Australia">Western Australia</collectingRegion>
• 
<specimenCount>♂</specimenCount>
(reared in captivity)
</materialsCitation>
, 
<materialsCitation collectingDate="2016-05-21" collectorName="J. DeJong" county="Mitchell River National Park" latitude="-14.82218" location="Little Merten's Falls" longLatPrecision="5" longitude="125.7131" municipality="Bujani" specimenCount="6" stateProvince="Western Australia">
<specimenCount type="generic">6 imm.</specimenCount>
; 
<collectingCounty>Mitchell River National Park</collectingCounty>
, 
<collectingMunicipality>Bujani</collectingMunicipality>
(
<location LSID="urn:lsid:plazi:treatment:802275FDE49A519D985FCA1C85E4DBF6:E8EA8ADDAB97DDA8A0F69BA312611D79" county="Mitchell River National Park" latitude="-14.82218" longLatPrecision="5" longitude="125.7131" municipality="Bujani" name="Little Merten's Falls" stateProvince="Western Australia">
Little 
<normalizedToken originalValue="Merten’s">Merten's</normalizedToken>
Falls
</location>
); 
<geoCoordinate degrees="14.82218" direction="south" orientation="latitude" precision="1" value="-14.82218">-14.82218</geoCoordinate>
, 
<geoCoordinate degrees="125.7131" direction="east" orientation="longitude" precision="5" value="125.7131">125.7131</geoCoordinate>
; 
<collectingDate value="2016-05-21">21 May 2016</collectingDate>
; 
<collectorName>J. DeJong</collectorName>
leg.; on rocks at night; sel_1244-1245; SCC16_047; (WAM T155620-T155621)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="emended diagnosis">
<paragraph pageId="0" pageNumber="1">Emended diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Given the number of the new, morphologically similar species described here, the diagnosis is updated, and new illustrations are provided for ease of identification. The male is similar to other members of the group but is most easily differentiated by the RTA (Figs 
<figureCitation captionStart="Figure 18" captionStartId="F25" captionText="Figure 18. Members of the Kimberley species group A Karaops sp., Keep River National Park, Northern Territory (sel _ 1315, MAGNT A 004871) B habitat, Keep River National Park, Northern Territory C Karaops keithlongbottomi, holotype male, Drysdale River Station, Western Australia (WAM T 55002) D Karaops keithlongbottomi, palp, dorsal, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244, WAM T 155620) E Karaops keithlongbottomi, holotype male, palp, dorsal, Drysdale River Station, Western Australia (WAM T 55002) F Karaops dejongi sp. nov., holotype female, RAAF Boab Quarry, Western Australia (sel _ 1273, WAM T 155649) G Karaops umiida, holotype female, Irvine Island, Western Australia (WAM T 110393) H habitat, Keep River National Park, Northern Territory I Karaops keithlongbottomi, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244 WAM T 155620) J same, face showing marks on chelicerae. Scale bars: 0.5 mm (D, E); 1 mm (A, C, F, J); 5 mm (G, I)." figureDoi="10.3897/zookeys.1150.93760.figure18" httpUri="https://binary.pensoft.net/fig/816405" pageId="0" pageNumber="1">18D, E</figureCitation>
, 
<figureCitation captionStart="Figure 19" captionStartId="F26" captionText="Figure 19. Members of the Kimberley species group A Karaops keithlongbottomi, palp, ventral, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244, WAM T 155620) B same, retrolateral C Karaops sp., Mimbi Caves, Western Australia (sel _ 1276, WAM T 155652) D Karaops dejongi sp. nov., paratype male, RAAF Boab Quarry, Western Australia (sel _ 1272, WAM T 155648) E Karaops sp., Geike Gorge, Western Australia (sel _ 1267, WAM T 155643) F habitat, Mimbi Caves, Western Australia. Scale bars: 0.5 mm (A, B); 1 mm (C-E)." figureDoi="10.3897/zookeys.1150.93760.figure19" httpUri="https://binary.pensoft.net/fig/816406" pageId="0" pageNumber="1">19A, B</figureCitation>
). In ventral view, the vRTA is somewhat shaped like the tube and bell of a sousaphone, rather than the oblong or spoon-shaped vRTA of other species. The dRTA is short and plicate.
</paragraph>
<caption doi="10.3897/zookeys.1150.93760.map7" httpUri="https://binary.pensoft.net/fig/816403" pageId="0" pageNumber="1" start="Map 7" startId="F23">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Map 7.</emphasis>
Species of the Kimberley species group. Diamonds (juveniles): light purple = Mimbi Caves; blue = Geike Gorge, green = Adcock Gorge, teal = Augustus Island, pink = Little Mertens Falls, yellow = Emma Gorge, orange = 
<normalizedToken originalValue="Kelly’s">Kelly's</normalizedToken>
Knob, red = Keep River National Park; Circles: green = 
<taxonomicName authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops dejongi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dejongi">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops dejongi</emphasis>
</taxonomicName>
sp. nov., blue = 
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops jenniferae" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="jenniferae">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops jenniferae</emphasis>
</taxonomicName>
, red = 
<taxonomicName authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops dalmanyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dalmanyi">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops dalmanyi</emphasis>
</taxonomicName>
sp. nov., purple = 
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops</emphasis>
</taxonomicName>
sp. adult male (will be described in subsequent publication), white = 
<taxonomicName authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops conilurus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="conilurus">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops conilurus</emphasis>
</taxonomicName>
sp. nov., pink = 
<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops umiida" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="umiida">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops umiida</emphasis>
</taxonomicName>
, yellow = 
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops alanlongbottomi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="alanlongbottomi">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops alanlongbottomi</emphasis>
</taxonomicName>
, teal = 
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops keithlongbottomi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keithlongbottomi">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops keithlongbottomi</emphasis>
</taxonomicName>
, mauve = 
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops larryoo" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="larryoo">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops larryoo</emphasis>
</taxonomicName>
, orange = 
<taxonomicName authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops garyodwyeri" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="garyodwyeri">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops garyodwyeri</emphasis>
</taxonomicName>
sp. nov., black = 
<taxonomicName authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops malumbu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="malumbu">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops malumbu</emphasis>
</taxonomicName>
sp. nov. Juveniles are treated as distinct species based on molecular data.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
The description is updated here based on the recently collected additional male. The full description of the male can be found in 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.99.723" author="Crews, SC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 103" refId="B14" refString="Crews, SC, Harvey, MS, 2011. The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental Region. ZooKeys 99: 1 - 103, DOI: https://doi.org/10.3897/zookeys.99.723" title="The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental Region." url="https://doi.org/10.3897/zookeys.99.723" volume="99" year="2011">Crews and Harvey (2011)</bibRefCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Male</emphasis>
(sel_1244, WAM T155620). Color (in life Fig. 
<figureCitation captionStart="Figure 17" captionStartId="F24" captionText="Figure 17. Members of the Kimberley species group A Karaops sp., Kelly's Knob, Western Australia B Karaops sp., Kelly's Knob, Western Australia C Karaops sp., Keep River National Park, Northern Territory (sel _ 1320, MAGNT A 004876) D Karaops sp. eating a fly, Kelly's Knob, Western Australia E Karaops keithlongbottomi, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244, WAM T 155620)." figureDoi="10.3897/zookeys.1150.93760.figure17" httpUri="https://binary.pensoft.net/fig/816404" pageId="0" pageNumber="1">17E</figureCitation>
/preserved Fig. 
<figureCitation captionStart="Figure 18" captionStartId="F25" captionText="Figure 18. Members of the Kimberley species group A Karaops sp., Keep River National Park, Northern Territory (sel _ 1315, MAGNT A 004871) B habitat, Keep River National Park, Northern Territory C Karaops keithlongbottomi, holotype male, Drysdale River Station, Western Australia (WAM T 55002) D Karaops keithlongbottomi, palp, dorsal, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244, WAM T 155620) E Karaops keithlongbottomi, holotype male, palp, dorsal, Drysdale River Station, Western Australia (WAM T 55002) F Karaops dejongi sp. nov., holotype female, RAAF Boab Quarry, Western Australia (sel _ 1273, WAM T 155649) G Karaops umiida, holotype female, Irvine Island, Western Australia (WAM T 110393) H habitat, Keep River National Park, Northern Territory I Karaops keithlongbottomi, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244 WAM T 155620) J same, face showing marks on chelicerae. Scale bars: 0.5 mm (D, E); 1 mm (A, C, F, J); 5 mm (G, I)." figureDoi="10.3897/zookeys.1150.93760.figure18" httpUri="https://binary.pensoft.net/fig/816405" pageId="0" pageNumber="1">18C, I, J</figureCitation>
): Carapace: pale tan-yellow with two dark marks mediolaterally on either side of fovea and a small, dark mark posteromedially/orangish brown, darker markings not as conspicuous, with dark, slender setae, darker where two dark marks located, distributed somewhat densely but integument still visible beneath, a few dark, stiff setae posteriorly. Chelicerae: yellowish tan, paturon with dark edges (Fig. 
<figureCitation captionStart="Figure 18" captionStartId="F25" captionText="Figure 18. Members of the Kimberley species group A Karaops sp., Keep River National Park, Northern Territory (sel _ 1315, MAGNT A 004871) B habitat, Keep River National Park, Northern Territory C Karaops keithlongbottomi, holotype male, Drysdale River Station, Western Australia (WAM T 55002) D Karaops keithlongbottomi, palp, dorsal, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244, WAM T 155620) E Karaops keithlongbottomi, holotype male, palp, dorsal, Drysdale River Station, Western Australia (WAM T 55002) F Karaops dejongi sp. nov., holotype female, RAAF Boab Quarry, Western Australia (sel _ 1273, WAM T 155649) G Karaops umiida, holotype female, Irvine Island, Western Australia (WAM T 110393) H habitat, Keep River National Park, Northern Territory I Karaops keithlongbottomi, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244 WAM T 155620) J same, face showing marks on chelicerae. Scale bars: 0.5 mm (D, E); 1 mm (A, C, F, J); 5 mm (G, I)." figureDoi="10.3897/zookeys.1150.93760.figure18" httpUri="https://binary.pensoft.net/fig/816405" pageId="0" pageNumber="1">18J</figureCitation>
)/orangish with dark marks less conspicuous than in life. Abdomen: pale brown with dark brown spots anteriorly and laterally, slightly dark in cardiac area, two dark patches medially, chevrons posteriorly, black, undulate mark posteriorly/orangish yellow, with spots and markings less conspicuous, long, dark, slender setae. Legs: pale yellowish white, Cx with dark marks prolaterally, Tr, Fm with small black dots at Tr-Fm joint, dark marks paler centrally, Fm with dark annulation at Fm-Pt joint, Pt with dark annulation at Pt-Ti joint, Ti with dark annulation mid-Ti and one closer to the Ti-Mt joint, Mt with annulation at both ends, Ta dusky distally/orangish brown, dark marks less conspicuous; spination leg I Ti v 2-2-2-2-2-2, Mt v 2-2; leg II Ti v 2-2-2-2-2, Mt v 2-2-2.
</paragraph>
<caption doi="10.3897/zookeys.1150.93760.figure17" httpUri="https://binary.pensoft.net/fig/816404" pageId="0" pageNumber="1" start="Figure 17" startId="F24">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 17.</emphasis>
Members of the Kimberley species group 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops</emphasis>
</taxonomicName>
sp., 
<normalizedToken originalValue="Kelly’s">Kelly's</normalizedToken>
Knob, Western Australia 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops</emphasis>
</taxonomicName>
sp., 
<normalizedToken originalValue="Kelly’s">Kelly's</normalizedToken>
Knob, Western Australia 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops</emphasis>
</taxonomicName>
sp., Keep River National Park, Northern Territory (sel_1320, MAGNT A004876) 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops</emphasis>
</taxonomicName>
sp. eating a fly, 
<normalizedToken originalValue="Kelly’s">Kelly's</normalizedToken>
Knob, Western Australia 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops keithlongbottomi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keithlongbottomi">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops keithlongbottomi</emphasis>
</taxonomicName>
, Bujani (Little 
<normalizedToken originalValue="Merten’s">Merten's</normalizedToken>
Falls), Mitchell River National Park, Western Australia (sel_1244, WAM T155620).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1150.93760.figure18" httpUri="https://binary.pensoft.net/fig/816405" pageId="0" pageNumber="1" start="Figure 18" startId="F25">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 18.</emphasis>
Members of the Kimberley species group 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops</emphasis>
</taxonomicName>
sp., Keep River National Park, Northern Territory (sel_1315, MAGNT A004871) 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
habitat, Keep River National Park, Northern Territory 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops keithlongbottomi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keithlongbottomi">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops keithlongbottomi</emphasis>
</taxonomicName>
, holotype male, Drysdale River Station, Western Australia (WAM T55002) 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops keithlongbottomi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keithlongbottomi">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops keithlongbottomi</emphasis>
</taxonomicName>
, palp, dorsal, Bujani (Little 
<normalizedToken originalValue="Merten’s">Merten's</normalizedToken>
Falls), Mitchell River National Park, Western Australia (sel_1244, WAM T155620) 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops keithlongbottomi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keithlongbottomi">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops keithlongbottomi</emphasis>
</taxonomicName>
, holotype male, palp, dorsal, Drysdale River Station, Western Australia (WAM T55002) 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops dejongi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dejongi">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops dejongi</emphasis>
</taxonomicName>
sp. nov., holotype female, RAAF Boab Quarry, Western Australia (sel_1273, WAM T155649) 
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops umiida" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="umiida">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops umiida</emphasis>
</taxonomicName>
, holotype female, Irvine Island, Western Australia (WAM T110393) 
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
habitat, Keep River National Park, Northern Territory 
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops keithlongbottomi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keithlongbottomi">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops keithlongbottomi</emphasis>
</taxonomicName>
, Bujani (Little 
<normalizedToken originalValue="Merten’s">Merten's</normalizedToken>
Falls), Mitchell River National Park, Western Australia (sel_1244 WAM T155620) 
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
same, face showing marks on chelicerae. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">D, E</emphasis>
); 1 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A, C, F, J</emphasis>
); 5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">G, I</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1150.93760.figure19" httpUri="https://binary.pensoft.net/fig/816406" pageId="0" pageNumber="1" start="Figure 19" startId="F26">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 19.</emphasis>
Members of the Kimberley species group 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops keithlongbottomi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keithlongbottomi">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops keithlongbottomi</emphasis>
</taxonomicName>
, palp, ventral, Bujani (Little 
<normalizedToken originalValue="Merten’s">Merten's</normalizedToken>
Falls), Mitchell River National Park, Western Australia (sel_1244, WAM T155620) 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
same, retrolateral 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops</emphasis>
</taxonomicName>
sp., Mimbi Caves, Western Australia (sel_1276, WAM T155652) 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops dejongi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dejongi">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops dejongi</emphasis>
</taxonomicName>
sp. nov., paratype male, RAAF Boab Quarry, Western Australia (sel_1272, WAM T155648) 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops</emphasis>
</taxonomicName>
sp., Geike Gorge, Western Australia (sel_1267, WAM T155643) 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
habitat, Mimbi Caves, Western Australia. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
); 1 mm (
<emphasis bold="true" pageId="0" pageNumber="1">C-E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Known from Ngauwudu (Mitchell Plateau), Northern Kimberley, Western Australia (Map 
<figureCitation captionStart="Map 7" captionStartId="F23" captionText="Map 7. Species of the Kimberley species group. Diamonds (juveniles): light purple = Mimbi Caves; blue = Geike Gorge, green = Adcock Gorge, teal = Augustus Island, pink = Little Mertens Falls, yellow = Emma Gorge, orange = Kelly's Knob, red = Keep River National Park; Circles: green = Karaops dejongi sp. nov., blue = Karaops jenniferae, red = Karaops dalmanyi sp. nov., purple = Karaops sp. adult male (will be described in subsequent publication), white = Karaops conilurus sp. nov., pink = Karaops umiida, yellow = Karaops alanlongbottomi, teal = Karaops keithlongbottomi, mauve = Karaops larryoo, orange = Karaops garyodwyeri sp. nov., black = Karaops malumbu sp. nov. Juveniles are treated as distinct species based on molecular data." figureDoi="10.3897/zookeys.1150.93760.map7" httpUri="https://binary.pensoft.net/fig/816403" pageId="0" pageNumber="1">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="natural history">
<paragraph pageId="0" pageNumber="1">Natural history.</paragraph>
<paragraph pageId="0" pageNumber="1">
Very little is known about the type other than it was collected from Drysdale River Station, late 1995. The precise locality, time of year, and microhabitat are unknown. The abdomen of the type is damaged (Fig. 
<figureCitation captionStart="Figure 18" captionStartId="F25" captionText="Figure 18. Members of the Kimberley species group A Karaops sp., Keep River National Park, Northern Territory (sel _ 1315, MAGNT A 004871) B habitat, Keep River National Park, Northern Territory C Karaops keithlongbottomi, holotype male, Drysdale River Station, Western Australia (WAM T 55002) D Karaops keithlongbottomi, palp, dorsal, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244, WAM T 155620) E Karaops keithlongbottomi, holotype male, palp, dorsal, Drysdale River Station, Western Australia (WAM T 55002) F Karaops dejongi sp. nov., holotype female, RAAF Boab Quarry, Western Australia (sel _ 1273, WAM T 155649) G Karaops umiida, holotype female, Irvine Island, Western Australia (WAM T 110393) H habitat, Keep River National Park, Northern Territory I Karaops keithlongbottomi, Bujani (Little Merten's Falls), Mitchell River National Park, Western Australia (sel _ 1244 WAM T 155620) J same, face showing marks on chelicerae. Scale bars: 0.5 mm (D, E); 1 mm (A, C, F, J); 5 mm (G, I)." figureDoi="10.3897/zookeys.1150.93760.figure18" httpUri="https://binary.pensoft.net/fig/816405" pageId="0" pageNumber="1">18C</figureCitation>
). The new specimens were collected on rocks at night at Bujani (Little 
<normalizedToken originalValue="Merten’s">Merten's</normalizedToken>
Falls), Mitchell River National Park. Males and penultimate males are known from the drier, cooler part of the year. This species is known from the Mitchell subregion of the North Kimberley bioregion. The area has a tropical savannah climate, and it is very wet in the wet season, often rendering roads impassable (
<bibRefCitation author="Graham, G" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="1" publicationUrl="https://www.dpaw.wa.gov.au/images/documents/about/science/projects/waaudit/north_kimberley01_p497-505.pdf" refId="B24" refString="Graham, G, 2001b. North Kimberley 1 (NK1 - Mitchell subregion). In: Department of Conservation and Land Management A Biodiversity Audit of Western Australia's 53 Biogeographic Subregions in 2002, 497-505. https://www.dpaw.wa.gov.au/images/documents/about/science/projects/waaudit/north_kimberley01_p497-505.pdf" title="North Kimberley 1 (NK 1 - Mitchell subregion). In: Department of Conservation and Land Management A Biodiversity Audit of Western Australia's 53 Biogeographic Subregions in 2002, 497 - 505." url="https://www.dpaw.wa.gov.au/images/documents/about/science/projects/waaudit/north_kimberley01_p497-505.pdf" year="2001 b">Graham 2001b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
This specimen and the penultimate female are quite large - the largest 
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops</emphasis>
</taxonomicName>
species known. The length of the holotype is unknown due to abdominal damage, but the carapace is 3.3. The length of the newly collected specimen is 8.41, with a carapace length of 4.64. The female remains undescribed. Although we only know of two areas where it is found, the new data do help to provide a bit more knowledge of its range, highlighting the importance targeted collecting and rearing to adulthood (Suppl. material 2: tables S1, S6).
</paragraph>
</subSubSection>
</treatment>
</document>