119 lines
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119 lines
14 KiB
XML
<document id="6CF9F6A7212C67D49E8FBC2556870F25" ID-CLB-Dataset="33935" ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak" docDate="2011" docId="7D2EE251673097B211B91B09C7E1DEA8" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha fluminoris Hoffman 1977" docType="treatment" docVersion="8" lastPageNumber="68" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="65" updateTime="1732923565725" updateUser="ExternalLinkService">
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<mods:title id="C55889C815BE228E6DE37D3F5D27E838">Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:name id="0B8138A29540A7670865DE611AFBB990" type="personal">
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<mods:roleTerm id="C149EE429A7B39C26C3699ADF24AB154">Author</mods:roleTerm>
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<mods:namePart id="EFB248D431E6B213B2ABEF51C4DA662E">Likhitrakarn, Natdanai</mods:namePart>
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<mods:namePart id="141E045065EA471C3956024A89C7135E">Golovatch, Sergei I.</mods:namePart>
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<mods:roleTerm id="AC0FFA217E5FD1533A3B29BDF42184CC">Author</mods:roleTerm>
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<mods:namePart id="CD799B1108B45E1097E13D224F0C0562">Panha, Somsak</mods:namePart>
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<mods:typeOfResource id="367C2F8A400EDEEEE4459514414A4F21">text</mods:typeOfResource>
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<mods:title id="68A134D11C54DEEF9E19DCC305CBDD1F">ZooKeys</mods:title>
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<mods:part id="B9053EB899E3C064E42E82975C4EA17F">
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<mods:date id="0F1F7B1A3749F65F35FC935A59170CA4">2011</mods:date>
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<mods:detail id="73787D74D53FE04D8411B6D8EEC24C74" type="volume">
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<mods:number id="1985D374731B45689146F76B09E31E00">131</mods:number>
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<mods:start id="DBE190A04446E4509BA725B971C1291D">1</mods:start>
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<mods:location id="7D29AD4869B539E984D2F90A23AC0620">
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<mods:url id="9074AC8BCA28FE73093DDDAB408BA2C4">http://dx.doi.org/10.3897/zookeys.131.1921</mods:url>
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<mods:classification id="AA810CD033B5B1847FD64DE2F99DA23D">journal article</mods:classification>
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<mods:identifier id="9AE6D8AF51C628FCE7E33A9CE1AEB15B" type="DOI">http://dx.doi.org/10.3897/zookeys.131.1921</mods:identifier>
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<mods:identifier id="E8B5D048B6431BB14FF3F8ABEEB16784" type="Pensoft-Pub">1313-2970-131-1</mods:identifier>
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<treatment id="7D2EE251673097B211B91B09C7E1DEA8" ID-GBIF-Taxon="152031129" LSID="urn:lsid:plazi:treatment:7D2EE251673097B211B91B09C7E1DEA8" httpUri="http://treatment.plazi.org/id/7D2EE251673097B211B91B09C7E1DEA8" lastPageId="67" lastPageNumber="68" pageId="64" pageNumber="65" scope_class="Diplopoda" scope_family="Paradoxosomatidae" scope_order="Polydesmida">
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<subSubSection id="79D6435AC982D183B468763C6772FA98" pageId="64" pageNumber="65" type="nomenclature">
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<paragraph id="4F10DB02350BE141620FAAD4E1DD67E1" pageId="64" pageNumber="65">
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<taxonomicName id="E0676A581B7B6EA653363E4F9D2F51E4" ID-CoL="758N8" LSID="http://species-id.net/wiki/Orthomorpha_fluminoris" authority="Hoffman, 1977" authorityName="Hoffman" authorityYear="1977" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha fluminoris" order="Polydesmida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="fluminoris">Orthomorpha fluminoris Hoffman, 1977</taxonomicName>
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Figs 5556
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</paragraph>
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</subSubSection>
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<subSubSection id="0337B673297D8523E2BAA0595E7DDC1D" pageId="64" pageNumber="65" type="reference_group">
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<paragraph id="A47E2DF35FF6F01274343AC041BD7AB5" pageId="64" pageNumber="65">
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<taxonomicName id="6BAFE99E12B89472388F16C823ECA4F5" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha fluminoris" order="Polydesmida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="fluminoris">Orthomorpha fluminoris</taxonomicName>
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<bibRefCitation id="ACF4BCA1CBB046DC4131DC4ADF858512" author="Hoffman, RL" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="699 - 719" title="Diplopoda from Malayan caves, collected by M. Pierre Strinati." volume="84" year="1977">Hoffman 1977</bibRefCitation>
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: 701 (D).
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</paragraph>
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<paragraph id="9F95E02D265651D28EC93C04B4387841" pageId="64" pageNumber="65">
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<taxonomicName id="F263431477E793252493CA954196C42B" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha fluminoris" order="Polydesmida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="fluminoris">Orthomorpha fluminoris</taxonomicName>
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-
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<bibRefCitation id="9F21337831D000B3C51BE41ACE7ACA2F" pageId="64" pageNumber="65">Golovatch 1998</bibRefCitation>
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: 42 (D, M).
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</paragraph>
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</subSubSection>
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<subSubSection id="A1DE0F88F34711719D6EE55A721AD57F" pageId="64" pageNumber="65" type="holotype">
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<paragraph id="793DEA5B7F26C7D163C64780F3DDE107" pageId="64" pageNumber="65">Holotype.</paragraph>
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<paragraph id="94BD51D70AEA5A78AEA06B7876D1F234" pageId="64" pageNumber="65">♂ (MHNG), Malaysia, environs of Kuala Lumpur, Batu Caves, 24.02.1975, leg. P. Strinati.</paragraph>
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</subSubSection>
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<subSubSection id="15EBA2AE9BEEB84B894D21CD45E8E584" lastPageId="67" lastPageNumber="68" pageId="64" pageNumber="65" type="redescription">
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<paragraph id="898B977367DE64C98AF83C2D55DBC9F1" pageId="64" pageNumber="65">Redescription.</paragraph>
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<paragraph id="7DB5FFBD6D5569C1F518B57451C9282F" pageId="64" pageNumber="65">Length 39 mm, width of midbody pro- and metazona 2.9 and 4.4 mm, respectively. Coloration of alcohol material upon long-term preservation dark castaneous brown with contrasting yellowish paraterga; venter, legs yellownish and tip of epiproct light red-brown, antennae also light red-brown, but antenomere 6 slightly infuscate, brownish, antenomere 7 dark brown, and tip of antennae pallid (Fig. 55).</paragraph>
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<paragraph id="19E9CC62B9B55B5A9E603D514785D2B9" lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65">
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Clypeolabral region densely setose, vertex sparsely so, epicranial suture distinct. Antennae rather short, clavate (antennomere 6 broadest), extending behind body segment 2 dorsally. Head in width <collum <segment 2 <3 <4 <5-16; thereafter body gently and gradually tapering. Collum with three transverse rows of setae, 4+4 anterior, 2+2 intermediate, and 3+3 posterior; caudal corner of paraterga nearly pointed, slightly declined ventrally and continuing
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<normalizedToken id="2F45BC52319C25CFDB6C4CF1CCD73335" originalValue="collum’s">collum's</normalizedToken>
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convexity (Fig. 55A & B); paraterga acutangular (ca 50°), slightly extending behind tergal margin, posterior
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<pageBreakToken id="1E11DDC51131AB6BAF1F347F489E1BB9" pageId="65" pageNumber="66" start="start">edge</pageBreakToken>
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slightly concave (Fig. 55A & B). Tegument smooth and poorly shining, prozona very finely shagreened, metazona leathery, rugulose, below paraterga microgranular. Metaterga 2-19 with two transverse rows of setae: 2+2 in anterior (pre-sulcus) row and 3+3 in posterior (postsulcus) one, all mostly abraded, but still traceable as insertion points. Tergal setae simple, moderately long, about 1/4 metatergal length. Axial line traceable. Paraterga very well developed (Fig. 55A-G), all lying below dorsum (at about 1/4 body height), slightly upturned to subhorizontal, in lateral view moderately strongly enlarged on pore-bearing segments, thinner on poreless ones; shoulders always present, broadly rounded and narrowly bordered, fused to callus; anterior edge of paraterga 2 straight, following paraterga rounded, caudal corner of all paraterga rounded, extending invariably and increasingly beyond rear tergal margin. Calluses delimited by a sulcus both dorsally and ventrally, rather broad. Posterior edge of paraterga evidently concave, especially strongly so on segments 16-19. Ozopores evident, lateral, lying in an ovoid groove at about 1/4 in front of caudal corner. Transverse sulcus complete on metaterga 5-18, incomplete on segments 4 and 19, very narrow, shallow, not reaching bases of paraterga, very finely ribbed at bottom (Fig. 55A, C
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<pageBreakToken id="ED7D4138EE0BC92B740C668F95128DC1" pageId="66" pageNumber="67" start="start">&</pageBreakToken>
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amp; F). Stricture between pro- and metazona narrow and shallow, faintly beaded at bottom down to base of paraterga (Fig. 55A-F). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2-4, a low and interrupted swelling supplied with a small tooth caudally on segments 5-7, retained only as small denticles on segments 8-14, thereafter missing (Fig. 55B, D & E). Epiproct (Fig. 55E-G) conical, flattened dorsoventrally, with two small apical papillae directed ventrocaudally, subtruncate at tip; pre-apical papillae small, lying close to tip. Hypoproct (Fig. 55G) nearly semi-circular, caudal tip very broadly rounded, setiferous knobs at caudal edge very small and well-separated.
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</paragraph>
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<paragraph id="8ACFCA6E0C07CA34CC970E4924C08455" pageId="66" pageNumber="67">Sterna sparsely setose, without modifications; cross-impressions shallow; a pair of strongly separated, anteroventrally directed, narrowly rounded, setose cones between ♂ coxae 4 (Fig. 55H & I). A paramedian pair of small tubercles in front of gonopod aperture. Legs long and slender, midbody ones ca 1.3-1.5 times as long as body height, prefemora without modifications, ♂ tarsal brushes present until on legs of segment 17.</paragraph>
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<paragraph id="A9DC360DF51099DAF99DF1A0A483414E" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67">
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Gonopods (Fig. 56) a little more complex. Coxa long and slender, with several strong setae distodorsally. Prefemur densely setose, about half the length of femorite +
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<normalizedToken id="9ECCD6166C792BE76E2F998B10A213F8" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part. Femorite slender, slightly curved and nearly not enlarged distad, with a small, but conspicuous ventral knob near base of lamina lateralis, with a
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<pageBreakToken id="3C8242424416EBD5D19BA2373C0C5C0F" pageId="67" pageNumber="68" start="start">"</pageBreakToken>
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postfemoral" part demarcated by a distinct, oblique, lateral sulcus; solenophore with a tridentate tip, middle denticle shorter than terminal tooth, but longer than a small subterminal lobule; solenomere long, flagelliform, a short tip exposed.
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</paragraph>
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<caption id="1F726A7B93166F419F502F82B164A78C" pageId="67" pageNumber="68">
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<paragraph id="CBEA33294961FEEE2FD3D771BF2DEC2B" pageId="67" pageNumber="68">
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Figure 55.
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<taxonomicName id="336B3299BF13CC0BD52382B6B292126E" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha fluminoris" order="Polydesmida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="fluminoris">Orthomorpha fluminoris</taxonomicName>
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Hoffman, 1977, ♂ holotype. A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively
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<normalizedToken id="23DB1F3867A90E2E7B9CAF01B6DB7E51" originalValue="E–G">E-G</normalizedToken>
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posterior part of body, lateral, dorsal and ventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
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</paragraph>
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</caption>
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<caption id="2A5B43E46AB9C01E6222199EA5E89B7C" pageId="67" pageNumber="68">
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<paragraph id="06A090F00E3493546507FE7AED0A847D" pageId="67" pageNumber="68">
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Figure 56.
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<taxonomicName id="572D608517C85422254C3A9F8863F1C0" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha fluminoris" order="Polydesmida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="fluminoris">Orthomorpha fluminoris</taxonomicName>
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Hoffman, 1977, ♂ holotype.
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<normalizedToken id="26726E4CCFBD6A5957C7BCB4FB7BCE0D" originalValue="A–C">A-C</normalizedToken>
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right gonopod, lateral, mesal and subcaudal views, respectively. Scale bar: 0.2 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="D981D578AC635D3F2A012AFEB3B03EBF" pageId="67" pageNumber="68" type="remarks">
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<paragraph id="12210BEA1017F6EA4DA4B945E5724FF5" pageId="67" pageNumber="68">Remarks.</paragraph>
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<paragraph id="2CFC419461CD2090A45C6EA3D4184385" pageId="67" pageNumber="68">
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The original description is fairly complete, also illustrated in due detail (
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<bibRefCitation id="0ACDF636DF1F3FB7B9B5ED0F55DCDDB9" author="Hoffman, RL" journalOrPublisher="Journal of Natural History" pageId="114" pageNumber="115" pagination="361 - 389" title="Descriptions and allocation of new or poorly known genera and species of Paradoxosomatidae from south-eastern Asia (Diplopoda: Polydesmida)." url="doi: 10.1080/00222937300770281" volume="7" year="1973">Hoffman 1973</bibRefCitation>
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). The above redescription, also based on the holotype, is only meant to bring it to our standard and to present additional illustrations.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |