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<document id="6A585D01A3EDBB0AFB476B1D1249BD07" ID-DOI="10.11646/zootaxa.5296.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7973065" ID-ZooBank="624273A6-3028-42C2-ABE2-A18BBF828156" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685085730506" checkinUser="plazi" docAuthor="Reyes-Gómez, Adriana, Vargas-Ponce, Ofelia, Galván-Villa, Cristian, Salgado-Barragán, José, Esqueda-González, Ma. Del Carmen &amp; Ríos-Jara, Eduardo" docDate="2023" docId="795287BFE45B5519D0F1714772BFFCF2" docLanguage="en" docName="zootaxa.5296.2.2.pdf" docOrigin="Zootaxa 5296 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidozona subtilis Berry 1956" docType="treatment" docVersion="3" lastPageNumber="160" masterDocId="856BFFC7E4575514D06676457344FFAF" masterDocTitle="Inventory of chiton species (Polyplacophora) from the rocky intertidal of the Northern Gulf of California, with an illustrated taxonomic key" masterLastPageNumber="178" masterPageNumber="147" pageNumber="159" updateTime="1685088964505" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="37D4A819A5FF4EA02EED350A86D0B2B4">Inventory of chiton species (Polyplacophora) from the rocky intertidal of the Northern Gulf of California, with an illustrated taxonomic key</mods:title>
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<mods:namePart id="52B76E15E9E44F63002ACCABD1602C27">Reyes-Gómez, Adriana</mods:namePart>
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<mods:affiliation id="49ED87051DD0336ACD1F6EA8F2B51FE3">Departamento de Ecología Aplicada, Centro Universitario de Ciencias Biológicas y Agropecuarias, Universidad de Guadalajara, Camino Ing. Ramón Padilla Sánchez 2100, Zapopan, Jalisco, 45200, México.</mods:affiliation>
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<mods:namePart id="2DD48B0681DCF22D744D53BBD41E8DD4">Salgado-Barragán, José</mods:namePart>
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<mods:namePart id="2B862F2F21E5837132E722FBEC027FAC">Esqueda-González, Ma. Del Carmen</mods:namePart>
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<mods:namePart id="9AEAEA360E5DA688ACF2DACD043E67AB">Ríos-Jara, Eduardo</mods:namePart>
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<paragraph id="F14436A9E45B5518D0F17147715EF8B3" blockId="12.[151,538,1794,1857]" box="[151,538,1794,1821]" pageId="12" pageNumber="159">
<heading id="AA0C81C5E45B5518D0F17147715EF8B3" bold="true" box="[151,538,1794,1821]" fontSize="11" level="1" pageId="12" pageNumber="159" reason="1">
<taxonomicName id="36FB4D2AE45B5518D0F17147715EF8B3" ID-CoL="6PBNL" authority="Berry, 1956" authorityName="Berry" authorityYear="1956" box="[151,538,1794,1821]" class="Polyplacophora" family="Ischnochitonidae" genus="Lepidozona" kingdom="Animalia" order="Chitonida" pageId="12" pageNumber="159" phylum="Mollusca" rank="species" species="subtilis">
<emphasis id="C38FEABBE45B5518D0F17147715EF8B3" bold="true" box="[151,538,1794,1821]" pageId="12" pageNumber="159">
<emphasis id="C38FEABBE45B5518D0F1714772C5F8B2" bold="true" box="[151,385,1794,1821]" italics="true" pageId="12" pageNumber="159">Lepidozona subtilis</emphasis>
<bibRefCitation id="956A4B58E45B5518D1EE7146715EF8B3" author="Berry, S. S." box="[392,538,1794,1821]" pageId="12" pageNumber="159" pagination="71 - 74" refId="ref16326" refString="Berry, S. S. (1956) Diagnoses of new eastern Pacific chitons. Leaflets in Malacology, 1, 71 - 74." type="journal article" year="1956">Berry, 1956</bibRefCitation>
</emphasis>
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<paragraph id="F14436A9E45B5518D0F171637231F8EE" blockId="12.[151,538,1794,1857]" box="[151,373,1830,1857]" pageId="12" pageNumber="159">
(
<figureCitation id="69C02A2CE45B5518D0F971637264F8EE" box="[159,288,1830,1857]" captionStart="FIGURE 2" captionStartId="5.[151,250,1791,1815]" captionTargetBox="[151,1436,181,1767]" captionTargetId="figure-21@5.[151,1436,181,1768]" captionTargetPageId="5" captionText="FIGURE 2. Chiton species obtained from the Northern Gulf of California. A. Dorsal and lateral views of Ischnochiton guatemalensis CUCBACM 00450. B. Dorsal and lateral views of Ischnochiton tridentatus CUCBACM 00453. C. Dorsal and lateral views of Stenoplax limaciformis CUCBACM 00455. D. Dorsal and lateral views of Stenoplax mariposa CUCBACM 00459. E. Dorsal and lateral views of Stenoplax sonorana CUCBACM 00462. F. Dorsal view of Lepidozona clathrata CUCBACM 00464. G. Dorsal view of Lepidozona serrata CUCBACM 00471. H. Dorsal and lateral views of Lepidozona subtilis CUCBACM 00475. I. Dorsal and lateral views of Callistochiton elenensis CUCBACM 00476." figureDoi="http://doi.org/10.5281/zenodo.7973081" httpUri="https://zenodo.org/record/7973081/files/figure.png" pageId="12" pageNumber="159">Figures 2H</figureCitation>
,
<figureCitation id="69C02A2CE45B5518D14A71627229F8EE" box="[300,365,1831,1857]" captionStart="FIGURE 7" captionStartId="14.[151,250,1478,1502]" captionTargetBox="[151,1436,151,1455]" captionTargetId="figure-24@14.[151,1436,151,1455]" captionTargetPageId="14" captionText="FIGURE 7. AF. Scanning electron microscope images of Lepidozona subtilis CUCBACM 00475. A. Dorsal view of head valve. B. Dorsal view of half of intermediate valve IV. C. Dorsal view of tail valve. D. Valve IV aesthetes detail of intermediate area. E, F. Radular teeth details. GK. Scanning electron microscope images of Callistochiton elenensis CUCBACM 00476. G. Dorsal view of head valve. H. Dorsal view of half of intermediate valve IV. I. Dorsal view of tail valve. J. Dorsal girdle scales detail. K. Radular teeth. L. Radular teeth of Chiton virgulatus CUCBACM 00491 of Chiton virgulatus CUCBACM 00491." figureDoi="http://doi.org/10.5281/zenodo.7973378" httpUri="https://zenodo.org/record/7973378/files/figure.png" pageId="12" pageNumber="159">7AF</figureCitation>
)
</paragraph>
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<paragraph id="F14436A9E45B5518D0F1712E7733F82C" blockId="12.[151,1143,1899,1923]" box="[151,1143,1899,1923]" pageId="12" pageNumber="159">
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<treatmentCitation id="705A10B8E45B5518D0F1712E7163F82C" author="Berry, S. S." box="[151,551,1899,1923]" page="74" pageId="12" pageNumber="159" year="1956">
<taxonomicName id="36FB4D2AE45B5518D0F1712E7163F82C" ID-CoL="6PBNL" authority="Berry, 1956: 71 - 74" authorityName="Berry" authorityPageNumber="71 - 74" authorityYear="1956" box="[151,551,1899,1923]" class="Polyplacophora" family="Ischnochitonidae" genus="Lepidozona" kingdom="Animalia" order="Chitonida" pageId="12" pageNumber="159" phylum="Mollusca" rank="species" species="subtilis">
<emphasis id="C38FEABBE45B5518D0F1712E7219F82C" box="[151,349,1899,1923]" italics="true" pageId="12" pageNumber="159">Lepidozona subtilis</emphasis>
<bibRefCitation id="956A4B58E45B5518D105712E7163F82C" author="Berry, S. S." box="[355,551,1899,1923]" pageId="12" pageNumber="159" pagination="71 - 74" refId="ref16326" refString="Berry, S. S. (1956) Diagnoses of new eastern Pacific chitons. Leaflets in Malacology, 1, 71 - 74." type="journal article" year="1956">Berry, 1956: 7174</bibRefCitation>
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. Chresonymy and synonymy in
<bibRefCitation id="956A4B58E45B5518D317712E7730F82C" author="Kaas, P. &amp; Van Belle, R." box="[881,1140,1899,1923]" pageId="12" pageNumber="159" refId="ref18612" refString="Kaas, P. &amp; Van Belle, R. (1987) Monograph of Living Chitons (Mollusca: Polyplacophora). Vol. 3. Suborder Ischnochitonidae. Chaetopleurinae and Ischnochitoninae. E. J. Brill, Leiden, 302 pp. https: // doi. org / 10.1163 / 9789004431690" type="book" year="1987">Kaas &amp; Van Belle (1987)</bibRefCitation>
.
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</paragraph>
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<paragraph id="F14436A9E45B5518D0F171F47746F840" blockId="12.[151,1026,1969,2032]" pageId="12" pageNumber="159">
<emphasis id="C38FEABBE45B5518D0F171F47201F864" bold="true" box="[151,325,1969,1995]" pageId="12" pageNumber="159">Type material.</emphasis>
<materialsCitation id="41933CF4E45B5518D12A71F47746F840" collectionCode="SBMNH" country="Mexico" location="Puerto Penasco" pageId="12" pageNumber="159" specimenCode="SBMNH 34432" specimenCount="1" stateProvince="Sonora" typeStatus="holotype">
<typeStatus id="2E40880BE45B5518D12A71F472F3F864" box="[332,439,1969,1995]" pageId="12" pageNumber="159">Holotype</typeStatus>
,
<specimenCode id="A15D9ED2E45B5518D1A471F47137F864" box="[450,627,1969,1995]" collectionCode="SBMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="12" pageNumber="159" type="Museum">SBMNH 34432</specimenCode>
ex. “Berry collection 15132”.
<emphasis id="C38FEABBE45B5518D0A171907222F840" bold="true" box="[199,358,2005,2031]" pageId="12" pageNumber="159">Type locality.</emphasis>
<location id="F4246072E45B5518D10A7193715FF85F" LSID="urn:lsid:plazi:treatment:795287BFE45B5519D0F1714772BFFCF2:F4246072E45B5518D10A7193715FF85F" box="[364,539,2006,2032]" country="Mexico" name="Puerto Penasco" pageId="12" pageNumber="159" stateProvince="Sonora">Puerto Peñasco</location>
,
<collectingRegion id="333FF84BE45B5518D2407190713CF840" box="[550,632,2005,2031]" country="Mexico" name="Sonora" pageId="12" pageNumber="159">Sonora</collectingRegion>
,
<collectingCountry id="89EC7639E45B5518D2E27190719EF840" box="[644,730,2005,2031]" name="Mexico" pageId="12" pageNumber="159">México</collectingCountry>
(
<bibRefCitation id="956A4B58E45B5518D28E719070B1F840" author="Kaas, P. &amp; Van Belle, R." box="[744,1013,2005,2032]" pageId="12" pageNumber="159" refId="ref18612" refString="Kaas, P. &amp; Van Belle, R. (1987) Monograph of Living Chitons (Mollusca: Polyplacophora). Vol. 3. Suborder Ischnochitonidae. Chaetopleurinae and Ischnochitoninae. E. J. Brill, Leiden, 302 pp. https: // doi. org / 10.1163 / 9789004431690" type="book" year="1987">Kaas &amp; Van Belle 1987</bibRefCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="F14436A9E45A5519D0A176D37075FF1E" blockId="13.[151,1437,150,861]" box="[199,817,150,177]" pageId="13" pageNumber="160">
<emphasis id="C38FEABBE45A5519D0A176D372F5FF1F" bold="true" box="[199,433,150,176]" pageId="13" pageNumber="160">Material examined.</emphasis>
<specimenCount id="E7FDFD20E45A5519D1D176D27124FF1E" box="[439,608,151,177]" pageId="13" pageNumber="160" type="generic">284 specimens</specimenCount>
, BL
<quantity id="36039B4CE45A5519D2F376D27069FF1E" box="[661,813,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.2899999999999998" metricValueMax="1.97" metricValueMin="0.61" pageId="13" pageNumber="160" unit="mm" value="12.899999999999999" valueMax="19.7" valueMin="6.1">6.119.7 mm</quantity>
.
</paragraph>
<paragraph id="F14436A9E45A5519D0A176FF7603FEB2" blockId="13.[151,1437,150,861]" pageId="13" pageNumber="160">
<emphasis id="C38FEABBE45A5519D0A176FF726EFF7B" bold="true" box="[199,298,186,212]" pageId="13" pageNumber="160">Habitat.</emphasis>
On semihumid large and mediumsized rocks buried in sand with brown and red crusty algae patches exposed during low tide. At Playa Cementerio and Playa Gringa in the
<collectingRegion id="333FF84BE45A5519D3D3769A70B2FF56" box="[949,1014,223,249]" country="Brazil" name="Bahia" pageId="13" pageNumber="160">Bahía</collectingRegion>
de los Ángeles,
<collectingRegion id="333FF84BE45A5519D4D7769A761AFF56" box="[1201,1374,223,249]" country="Mexico" name="Baja California" pageId="13" pageNumber="160">Baja California</collectingRegion>
(BC) it was found on a substrate with abundant anemones and ascidians, asteroids, ophiuroids, and holothuroids.
</paragraph>
</subSubSection>
<subSubSection id="B9E16522E45A5519D0A1776272BFFCF2" pageId="13" pageNumber="160" type="discussion">
<paragraph id="F14436A9E45A5519D0A177627229FE26" blockId="13.[151,1437,150,861]" pageId="13" pageNumber="160">
<emphasis id="C38FEABBE45A5519D0A17762727FFEEE" bold="true" box="[199,315,295,321]" pageId="13" pageNumber="160">Remarks.</emphasis>
This species has a nodular radial sculpture (
<figureCitation id="69C02A2CE45A5519D342776270E3FEEE" box="[804,935,295,321]" captionStart="FIGURE 7" captionStartId="14.[151,250,1478,1502]" captionTargetBox="[151,1436,151,1455]" captionTargetId="figure-24@14.[151,1436,151,1455]" captionTargetPageId="14" captionText="FIGURE 7. AF. Scanning electron microscope images of Lepidozona subtilis CUCBACM 00475. A. Dorsal view of head valve. B. Dorsal view of half of intermediate valve IV. C. Dorsal view of tail valve. D. Valve IV aesthetes detail of intermediate area. E, F. Radular teeth details. GK. Scanning electron microscope images of Callistochiton elenensis CUCBACM 00476. G. Dorsal view of head valve. H. Dorsal view of half of intermediate valve IV. I. Dorsal view of tail valve. J. Dorsal girdle scales detail. K. Radular teeth. L. Radular teeth of Chiton virgulatus CUCBACM 00491 of Chiton virgulatus CUCBACM 00491." figureDoi="http://doi.org/10.5281/zenodo.7973378" httpUri="https://zenodo.org/record/7973378/files/figure.png" pageId="13" pageNumber="160">Figs. 7AC</figureCitation>
) and lateral areas with 46 fine grooves (
<figureCitation id="69C02A2CE45A5519D509776273F8FECA" captionStart="FIGURE 7" captionStartId="14.[151,250,1478,1502]" captionTargetBox="[151,1436,151,1455]" captionTargetId="figure-24@14.[151,1436,151,1455]" captionTargetPageId="14" captionText="FIGURE 7. AF. Scanning electron microscope images of Lepidozona subtilis CUCBACM 00475. A. Dorsal view of head valve. B. Dorsal view of half of intermediate valve IV. C. Dorsal view of tail valve. D. Valve IV aesthetes detail of intermediate area. E, F. Radular teeth details. GK. Scanning electron microscope images of Callistochiton elenensis CUCBACM 00476. G. Dorsal view of head valve. H. Dorsal view of half of intermediate valve IV. I. Dorsal view of tail valve. J. Dorsal girdle scales detail. K. Radular teeth. L. Radular teeth of Chiton virgulatus CUCBACM 00491 of Chiton virgulatus CUCBACM 00491." figureDoi="http://doi.org/10.5281/zenodo.7973378" httpUri="https://zenodo.org/record/7973378/files/figure.png" pageId="13" pageNumber="160">Fig. 7B</figureCitation>
). Aesthetes with 69 microaesthetes (
<figureCitation id="69C02A2CE45A5519D20A770E718DFECA" box="[620,713,331,357]" captionStart="FIGURE 7" captionStartId="14.[151,250,1478,1502]" captionTargetBox="[151,1436,151,1455]" captionTargetId="figure-24@14.[151,1436,151,1455]" captionTargetPageId="14" captionText="FIGURE 7. AF. Scanning electron microscope images of Lepidozona subtilis CUCBACM 00475. A. Dorsal view of head valve. B. Dorsal view of half of intermediate valve IV. C. Dorsal view of tail valve. D. Valve IV aesthetes detail of intermediate area. E, F. Radular teeth details. GK. Scanning electron microscope images of Callistochiton elenensis CUCBACM 00476. G. Dorsal view of head valve. H. Dorsal view of half of intermediate valve IV. I. Dorsal view of tail valve. J. Dorsal girdle scales detail. K. Radular teeth. L. Radular teeth of Chiton virgulatus CUCBACM 00491 of Chiton virgulatus CUCBACM 00491." figureDoi="http://doi.org/10.5281/zenodo.7973378" httpUri="https://zenodo.org/record/7973378/files/figure.png" pageId="13" pageNumber="160">Fig. 7D</figureCitation>
). Radula with bicuspid major lateral and bilobate minor lateral tooth (
<figureCitation id="69C02A2CE45A5519D086772A7225FE26" box="[224,353,367,393]" captionStart="FIGURE 7" captionStartId="14.[151,250,1478,1502]" captionTargetBox="[151,1436,151,1455]" captionTargetId="figure-24@14.[151,1436,151,1455]" captionTargetPageId="14" captionText="FIGURE 7. AF. Scanning electron microscope images of Lepidozona subtilis CUCBACM 00475. A. Dorsal view of head valve. B. Dorsal view of half of intermediate valve IV. C. Dorsal view of tail valve. D. Valve IV aesthetes detail of intermediate area. E, F. Radular teeth details. GK. Scanning electron microscope images of Callistochiton elenensis CUCBACM 00476. G. Dorsal view of head valve. H. Dorsal view of half of intermediate valve IV. I. Dorsal view of tail valve. J. Dorsal girdle scales detail. K. Radular teeth. L. Radular teeth of Chiton virgulatus CUCBACM 00491 of Chiton virgulatus CUCBACM 00491." figureDoi="http://doi.org/10.5281/zenodo.7973378" httpUri="https://zenodo.org/record/7973378/files/figure.png" pageId="13" pageNumber="160">Figs. 7EF</figureCitation>
).
</paragraph>
<paragraph id="F14436A9E45A5519D0A177D671C5FDCE" blockId="13.[151,1437,150,861]" pageId="13" pageNumber="160">
<bibRefCitation id="956A4B58E45A5519D0A177D67235FE02" author="Ferreira, A. J." box="[199,369,403,429]" pageId="13" pageNumber="160" pagination="162 - 180" refId="ref17385" refString="Ferreira, A. J. (1974) The genus Lepidozona in the Panamic Province, with the description of two new species. The Veliger, 17 (2), 162 - 180." type="journal article" year="1974">Ferreira (1974)</bibRefCitation>
proposed that
<taxonomicName id="36FB4D2AE45A5519D27377D671C6FE02" authorityName="Berry" authorityYear="1956" box="[533,642,403,429]" class="Polyplacophora" family="Ischnochitonidae" genus="Lepidozona" kingdom="Animalia" order="Chitonida" pageId="13" pageNumber="160" phylum="Mollusca" rank="species" species="subtilis">
<emphasis id="C38FEABBE45A5519D27377D67161FE03" box="[533,549,403,428]" italics="true" pageId="13" pageNumber="160">L</emphasis>
.
<emphasis id="C38FEABBE45A5519D25777D671C6FE02" box="[561,642,403,429]" italics="true" pageId="13" pageNumber="160">subtilis</emphasis>
</taxonomicName>
is distributed from
<geoCoordinate id="94CF506EE45A5519D33B77D670DDFE02" box="[861,921,403,429]" degrees="26" direction="north" orientation="latitude" pageId="13" pageNumber="160" precision="55555" value="26.0">26°N</geoCoordinate>
in the Gulf of California up to Puerto Peñasco including Puerto Lobos, Bahía
<collectingCountry id="89EC7639E45A5519D19D77F27136FE7E" box="[507,626,439,465]" name="Saint Pierre and Miquelon" pageId="13" pageNumber="160">San Pedro</collectingCountry>
, Guaymas, San Felipe, Puertecitos, Bahía San Luis Gonzaga, Bahía de los Ángeles, and Bahía San Francisquito. Most of this distribution accounts were based on field observations, specimens from the collection of the California Academy of Sciences, and private collections (Allyn G. Smith and George Hanselman). We recorded this species from Bahía Kino to Bahía de los Ángeles, BC which coincides with the distribution reported by
<bibRefCitation id="956A4B58E45A5519D1B674027139FDCE" author="Ferreira, A. J." box="[464,637,583,609]" pageId="13" pageNumber="160" pagination="162 - 180" refId="ref17385" refString="Ferreira, A. J. (1974) The genus Lepidozona in the Panamic Province, with the description of two new species. The Veliger, 17 (2), 162 - 180." type="journal article" year="1974">Ferreira (1974)</bibRefCitation>
.
</paragraph>
<paragraph id="F14436A9E45A5519D0A1742E72BFFCF2" blockId="13.[151,1437,150,861]" pageId="13" pageNumber="160">
This species was the most numerous in our samples, representing 40% of the total number of specimens, and was present at almost all sampling sites, except
<collectingRegion id="333FF84BE45A5519D2AD74CA7048FD06" box="[715,780,655,681]" country="Brazil" name="Bahia" pageId="13" pageNumber="160">Bahía</collectingRegion>
San Carlos,
<collectingRegion id="333FF84BE45A5519D3C074CA70BDFD06" box="[934,1017,655,681]" country="Mexico" name="Sonora" pageId="13" pageNumber="160">Sonora</collectingRegion>
. The studied specimens have varied coloration patterns of red, orange, yellow, olive green, dark green, and beige with combinations of these colors in spots or lines as was also reported by
<bibRefCitation id="956A4B58E45A5519D20E74927053FD5E" author="Ferreira, A. J." box="[616,791,727,753]" pageId="13" pageNumber="160" pagination="162 - 180" refId="ref17385" refString="Ferreira, A. J. (1974) The genus Lepidozona in the Panamic Province, with the description of two new species. The Veliger, 17 (2), 162 - 180." type="journal article" year="1974">Ferreira (1974)</bibRefCitation>
. We also observed a variable number of ribs in the adult individuals and no apparent relationship between rib number and body length. These variations were also pointed out by
<bibRefCitation id="956A4B58E45A5519D082755A72D4FC96" author="Ferreira, A. J." box="[228,400,799,825]" pageId="13" pageNumber="160" pagination="162 - 180" refId="ref17385" refString="Ferreira, A. J. (1974) The genus Lepidozona in the Panamic Province, with the description of two new species. The Veliger, 17 (2), 162 - 180." type="journal article" year="1974">Ferreira (1974)</bibRefCitation>
, who found that
<taxonomicName id="36FB4D2AE45A5519D22C755A71FCFC96" authorityName="Berry" authorityYear="1956" box="[586,696,799,825]" class="Polyplacophora" family="Ischnochitonidae" genus="Lepidozona" kingdom="Animalia" order="Chitonida" pageId="13" pageNumber="160" phylum="Mollusca" rank="species" species="subtilis">
<emphasis id="C38FEABBE45A5519D22C755A71FCFC96" box="[586,696,799,825]" italics="true" pageId="13" pageNumber="160">L. subtilis</emphasis>
</taxonomicName>
differs from its congeners by the absence of a protuberance (horn) on the apical edge of the scales.
</paragraph>
</subSubSection>
</treatment>
</document>