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<document id="BDD9551809B485E50AEA992EB34CE924" ID-CLB-Dataset="26770" ID-DOI="10.11646/zootaxa.4608.2.11" ID-GBIF-Dataset="f155c619-ee05-44b6-a8b1-feb851ca1af5" ID-ISSN="1175-5326" ID-Zenodo-Dep="3055376" ID-ZooBank="533A0CA9-0364-4593-947C-F709062AF188" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1558350051275" checkinUser="plazi" docAuthor="Maghradze, Eteri, Faille, Arnaud, Barjadze, Shalva &amp; Hlaváč, Peter" docDate="2019" docId="6751AF6BFF896B2AFF4D2FE0FE908E01" docLanguage="en" docName="zootaxa.4608.2.11.pdf" docOrigin="Zootaxa 4608 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bergrothia barbakadzei Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč 2019, sp. nov." docType="treatment" docVersion="7" lastPageNumber="378" masterDocId="9B68D713FF886B2DFFDA2A2DFFCE8A2C" masterDocTitle="A new cavernicolous species of the genus Bergrothia Reitter, 1884 (Coleoptera, Staphylinidae, Pselaphinae) from Georgia" masterLastPageNumber="379" masterPageNumber="371" pageNumber="372" updateTime="1698728757977" updateUser="ExternalLinkService">
<mods:mods id="738662123510A46DB0D541869557E843" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DE73EFEF3EA7AB37CAAAFB2371781B34">
<mods:title id="3F721066FA899FF7456F74F1D0C04A97">A new cavernicolous species of the genus Bergrothia Reitter, 1884 (Coleoptera, Staphylinidae, Pselaphinae) from Georgia</mods:title>
</mods:titleInfo>
<mods:name id="25BA9F0C8287A8A0731433A4E2BE9398" type="personal">
<mods:role id="31C1BAF5CDAC33C34EDB5950588D7EFF">
<mods:roleTerm id="AEB790AA4EEBBEE8D61A3BACA2255C32">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E026CA3CD7E54D8ED7FE1392E156737">Maghradze, Eteri</mods:namePart>
</mods:name>
<mods:name id="936B30A1BEAF6A1681FE83D974F95112" type="personal">
<mods:role id="6BDE708EE60691DC7410C58EAB4F9A04">
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<mods:namePart id="AE9461C2B96043AC9D2788F09B0D17FB">Faille, Arnaud</mods:namePart>
</mods:name>
<mods:name id="6F91CC0DC0088EF2520608605FD8DEC0" type="personal">
<mods:role id="88DDC0462C5D5C01A09C72C91A0FFA8B">
<mods:roleTerm id="CCBF04EA1D5D12554601B6BC66A8F597">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F2552C91A3C15B107D81B30A1DE83644">Barjadze, Shalva</mods:namePart>
</mods:name>
<mods:name id="8D183617433FDB901BD36CEF737A4FA0" type="personal">
<mods:role id="B344AA643F0635310DE01F81BEB5C96B">
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</mods:role>
<mods:namePart id="6BEF8C089DC4318DB9233BD8A0AAEFD2">Hlaváč, Peter</mods:namePart>
</mods:name>
<mods:typeOfResource id="391838B06AC57C691AE4D51E7ADD4A11">text</mods:typeOfResource>
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<mods:title id="112FB2B1B6D8DACD2E65FA86F5707D1F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="87C19E996ECE8F8A7F13B02BB17B7663">
<mods:date id="DBFEF38779569A51E1CDFA63E4AD5F85">2019</mods:date>
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<mods:number id="39B49105B46C3DC60EAF0FA20FCF68AC">2019-05-20</mods:number>
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<mods:detail id="5E45A807D2464F06EB2895BF0DE19E6C" type="volume">
<mods:number id="4E77BA57190EB7FDC459B5874EBE1FD4">4608</mods:number>
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<mods:identifier id="B32AFC4019BEAD1076DD14A64C3C44FB" type="CLB-Dataset">26770</mods:identifier>
<mods:identifier id="EA19561080524FEEC8E1A0B1F4147618" type="DOI">10.11646/zootaxa.4608.2.11</mods:identifier>
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<treatment id="6751AF6BFF896B2AFF4D2FE0FE908E01" ID-GBIF-Taxon="157232198" LSID="urn:lsid:plazi:treatment:6751AF6BFF896B2AFF4D2FE0FE908E01" httpUri="http://treatment.plazi.org/id/6751AF6BFF896B2AFF4D2FE0FE908E01" lastPageId="7" lastPageNumber="378" pageId="1" pageNumber="372">
<subSubSection id="A7E24DF6FF896B2CFF4D2FE0FBCB8FCB" box="[151,1029,1484,1512]" pageId="1" pageNumber="372" type="nomenclature">
<paragraph id="EF471E7DFF896B2CFF4D2FE0FBCB8FCB" blockId="1.[151,1029,1484,1547]" box="[151,1029,1484,1512]" pageId="1" pageNumber="372">
<heading id="B40FA911FF896B2CFF4D2FE0FBCB8FCB" bold="true" box="[151,1029,1484,1512]" fontSize="11" level="1" pageId="1" pageNumber="372" reason="1">
<emphasis id="DD8CC26FFF896B2CFF4D2FE0FBCB8FCB" bold="true" box="[151,1029,1484,1512]" pageId="1" pageNumber="372">
<taxonomicName id="28F865FEFF896B2CFF4D2FE0FC6E8FCB" ID-CoL="8XVPV" authority="Maghradze, Faille, Barjadze &amp; Hlavac" authorityName="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč" authorityYear="2019" box="[151,928,1484,1512]" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="species" species="barbakadzei" status="sp. nov.">
<emphasis id="DD8CC26FFF896B2CFF4D2FE0FE788FC4" bold="true" box="[151,438,1485,1512]" italics="true" pageId="1" pageNumber="372">Bergrothia barbakadzei</emphasis>
Maghradze, Faille, Barjadze &amp; Hlaváč
</taxonomicName>
<taxonomicNameLabel id="C6BF7F14FF896B2CFC722FE0FBCB8FCB" box="[936,1029,1485,1511]" pageId="1" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A7E24DF6FF896B2CFF4D2FDCFED78C27" box="[151,281,1521,1547]" pageId="1" pageNumber="372" type="description">
<paragraph id="EF471E7DFF896B2CFF4D2FDCFED78C27" blockId="1.[151,1029,1484,1547]" box="[151,281,1521,1547]" pageId="1" pageNumber="372">
(
<figureCitation id="77C302F8FF896B2CFF452FDCFEDF8C27" box="[159,273,1521,1547]" captionStart-0="FIGURES 78" captionStart-1="FIGURES 914" captionStart-2="FIGURES 1518" captionStart-3="FIGURES 1923" captionStartId-0="4.[151,264,1089,1114]" captionStartId-1="5.[151,264,1399,1424]" captionStartId-2="6.[151,264,1953,1978]" captionStartId-3="7.[151,264,782,807]" captionTargetBox-0="[153,1435,177,1025]" captionTargetBox-1="[153,1435,251,1350]" captionTargetBox-2="[153,1435,177,1887]" captionTargetBox-3="[161,1431,182,687]" captionTargetId-0="figure@4.[153,1438,177,1025]" captionTargetId-1="figure@5.[153,1438,270,1350]" captionTargetId-2="figure@6.[153,1438,177,1887]" captionTargetId-3="figure@7.[153,1438,177,687]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURES 78. Bergrothia barbakadzei sp. nov., habitus 7, dorsal view (female); 8, ventral view (male)." captionText-1="FIGURES 914. Bergrothia barbakadzei sp. nov. 9, head, dorsal view (female); 10, head dorsal view with (10a) sensilla on segment IV of maxillary palpus (female); 11, antenna (female); 12, antennal segment XI (female); 13, head, ventral view (male); 14, pronotum, dorsal view (female)." captionText-2="FIGURES 1518. Bergrothia barbakadzei sp. nov. 15, elytra (female); 16, median leg (male); 17, abdominal tergites IIVII, dorsal view (female); 18, abdominal tergites IIVII, ventral view (male)." captionText-3="FIGURES 1923. Mesotibiae of Bergrothia barbakadzei sp. nov. (scale bar: 250 μm). 19, male; 20, female; aedeagus of Bergrothia barbakadzei sp. nov. 21, lateral view (scale bar: 200 μm); 22, dorsal view (scale bar: 200 μm); aedeagus of Bergrothia mingrelica (Reitter). 23, dorsal view (Reitter) (scale bar: 200 μm). Redrawn from Hlaváč, 1999." httpUri-0="https://zenodo.org/record/3055394/files/figure.png" httpUri-1="https://zenodo.org/record/3055404/files/figure.png" httpUri-2="https://zenodo.org/record/3055410/files/figure.png" httpUri-3="https://zenodo.org/record/3055424/files/figure.png" pageId="1" pageNumber="372">Figs 722</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A7E24DF6FF896B2CFF4D2C15FE918D63" pageId="1" pageNumber="372" type="materials_examined">
<paragraph id="EF471E7DFF896B2CFF4D2C15FE918D63" blockId="1.[151,1436,1592,1979]" pageId="1" pageNumber="372">
<emphasis id="DD8CC26FFF896B2CFF4D2C15FE4D8C7F" bold="true" box="[151,387,1592,1619]" pageId="1" pageNumber="372">Material examined:</emphasis>
<materialsCitation id="5F901420FF896B2CFE502C14FD7C8C5B" ID-GBIF-Occurrence="2242680551" collectingDate="2018-07-05" collectionCode="SMNS" collectorName="E. Maghradze" country="Georgia" location="Datvis Cave" municipality="Sataplia-Tskaltubo" pageId="1" pageNumber="372" specimenCount="1" stateProvince="Imereti" typeStatus="holotype">
<typeStatus id="3043A0DFFF896B2CFE502C14FE3B8C7F" box="[394,501,1593,1619]" pageId="1" pageNumber="372" type="holotype">Holotype</typeStatus>
, ♁:
<collectingCountry id="97EF5EEDFF896B2CFDFF2C14FD688C7F" box="[549,678,1593,1619]" name="Georgia" pageId="1" pageNumber="372">GEORGIA</collectingCountry>
,
<collectingRegion id="2D3CD09FFF896B2CFD6B2C14FC9A8C7F" box="[689,852,1593,1619]" country="Georgia" name="Imereti" pageId="1" pageNumber="372">Imereti region</collectingRegion>
,
<collectingMunicipality id="0F238407FF896B2CFC852C14FBFB8C7F" box="[863,1077,1593,1619]" pageId="1" pageNumber="372">SatapliaTskaltubo</collectingMunicipality>
karst massif,
<location id="EA2748A6FF896B2CFB082C14FA918C7F" LSID="urn:lsid:plazi:treatment:6751AF6BFF896B2AFF4D2FE0FE908E01:EA2748A6FF896B2CFB082C14FA918C7F" box="[1234,1375,1593,1619]" country="Georgia" municipality="Sataplia-Tskaltubo" name="Datvis Cave" pageId="1" pageNumber="372" stateProvince="Imereti">Datvis Cave</location>
, dark zone,
<collectingDate id="8B02C155FF896B2CFF042C71FEAE8C5B" box="[222,352,1628,1655]" pageId="1" pageNumber="372" value="2018-07-05">
<date id="9B4638BDFF896B2CFF042C71FEAE8C5B" box="[222,352,1628,1655]" pageId="1" pageNumber="372" value="2018-07-05">05.vii.2018</date>
</collectingDate>
,
<collectorName id="420D7BABFF896B2CFEAA2C70FDDF8C5B" box="[368,529,1629,1655]" pageId="1" pageNumber="372">E. Maghradze</collectorName>
leg. (
<collectionCode id="89E986B8FF896B2CFD822C70FD648C5B" box="[600,682,1629,1655]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="372">SMNS</collectionCode>
)
</materialsCitation>
.
<typeStatus id="3043A0DFFF896B2CFD1A2C70FCE38C5B" box="[704,813,1629,1655]" pageId="1" pageNumber="372" type="paratype">Paratypes</typeStatus>
(6♁♁,
<specimenCount id="F9FED5F4FF896B2CFC562C70FC7E8C54" box="[908,944,1629,1656]" pageId="1" pageNumber="372" type="female">6♀</specimenCount>
):
<materialsCitation id="5F901420FF896B2CFC112C70FAA48C5B" ID-GBIF-Occurrence="2242680556" box="[971,1386,1628,1655]" collectingDate="2018-07-05" collectionCode="IZISU" collectorName="E. Maghradze" country="Georgia" location="Datvis Cave" municipality="Sataplia-Tskaltubo" pageId="1" pageNumber="372" specimenCount="1" stateProvince="Imereti" typeStatus="paratype">
same data as holotype, 1♁ (
<collectionCode id="89E986B8FF896B2CFACF2C70FAAC8C5B" box="[1301,1378,1629,1655]" pageId="1" pageNumber="372">IZISU</collectionCode>
)
</materialsCitation>
, 1♁ (
<collectionCode id="89E986B8FF896B2CFF452CACFF178CB7" box="[159,217,1665,1691]" pageId="1" pageNumber="372">CPH</collectionCode>
)
<materialsCitation id="5F901420FF896B2CFF332CACFE628CB7" ID-GBIF-Occurrence="2242680555" box="[233,428,1665,1691]" collectionCode="NMPC" country="Georgia" pageId="1" pageNumber="372" specimenCount="1" specimenCount-female="1" stateProvince="Imereti" typeStatus="paratype">
and
<specimenCount id="F9FED5F4FF896B2CFEC12CACFEF08CB7" box="[283,318,1665,1691]" pageId="1" pageNumber="372" type="female">1♀</specimenCount>
(
<collectionCode id="89E986B8FF896B2CFE942CACFE6A8CB7" box="[334,420,1665,1691]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="372">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5F901420FF896B2CFE632CACFB918CB7" ID-GBIF-Occurrence="2242680560" box="[441,1119,1664,1691]" collectingDate="2018-05-26" collectionCode="IZISU" collectorName="E. Maghradze" country="Georgia" location="Datvis Cave" municipality="Sataplia-Tskaltubo" pageId="1" pageNumber="372" specimenCount="1" specimenCount-female="1" stateProvince="Imereti" typeStatus="paratype">
same data as holotype, but
<collectingDate id="8B02C155FF896B2CFD292CACFCAD8CB7" box="[755,867,1665,1691]" pageId="1" pageNumber="372" value="2018-05-26">
<date id="9B4638BDFF896B2CFD292CACFCAD8CB7" box="[755,867,1665,1691]" pageId="1" pageNumber="372" value="2018-05-26">26.v.2018</date>
</collectingDate>
,
<specimenCount id="F9FED5F4FF896B2CFCAB2CACFC5A8CB7" box="[881,916,1665,1691]" pageId="1" pageNumber="372" type="female">1♀</specimenCount>
and 1♁, (
<collectionCode id="89E986B8FF896B2CFBD32CACFB998CB7" box="[1033,1111,1665,1691]" pageId="1" pageNumber="372">IZISU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5F901420FF896B2CFBB62CACFE798C93" ID-GBIF-Occurrence="2242680558" collectingDate="2018-07-29" collectionCode="IZISU" collectorName="E. Maghradze" country="Georgia" location="Datvis Cave" municipality="Sataplia-Tskaltubo" pageId="1" pageNumber="372" specimenCount="1" specimenCount-female="1" stateProvince="Imereti" typeStatus="paratype">
same data as holotype, but
<collectingDate id="8B02C155FF896B2CFF4D2C88FED78C93" box="[151,281,1701,1727]" pageId="1" pageNumber="372" value="2018-07-29">
<date id="9B4638BDFF896B2CFF4D2C88FED78C93" box="[151,281,1701,1727]" pageId="1" pageNumber="372" value="2018-07-29">29.vii.2018</date>
</collectingDate>
,
<specimenCount id="F9FED5F4FF896B2CFEFC2C88FE878C93" box="[294,329,1701,1727]" pageId="1" pageNumber="372" type="female">1♀</specimenCount>
, (
<collectionCode id="89E986B8FF896B2CFEBB2C88FE618C93" box="[353,431,1701,1727]" pageId="1" pageNumber="372">IZISU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5F901420FF896B2CFE1E2C88FBAE8C93" ID-GBIF-Occurrence="2242680550" box="[452,1120,1700,1727]" collectingDate="2018-09-03" collectionCode="SMNS" collectorName="E. Maghradze" country="Georgia" location="Datvis Cave" municipality="Sataplia-Tskaltubo" pageId="1" pageNumber="372" specimenCount="1" stateProvince="Imereti" typeStatus="holotype">
same data as data as
<typeStatus id="3043A0DFFF896B2CFD6F2C88FCD78C93" box="[693,793,1701,1727]" pageId="1" pageNumber="372" type="holotype">holotype</typeStatus>
,but
<collectingDate id="8B02C155FF896B2CFC902C88FC0B8C93" box="[842,965,1701,1727]" pageId="1" pageNumber="372" value="2018-09-03">
<date id="9B4638BDFF896B2CFC902C88FC0B8C93" box="[842,965,1701,1727]" pageId="1" pageNumber="372" value="2018-09-03">03.ix.2018</date>
</collectingDate>
, 1♁ (
<collectionCode id="89E986B8FF896B2CFBDC2C88FB968C93" box="[1030,1112,1701,1727]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="372">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5F901420FF896B2CFBB72C88FE7F8CCF" ID-GBIF-Occurrence="2242680557" collectingDate="2018-09-28" collectionCode="NMPC" collectorName="E. Maghradze" country="Georgia" location="Datvis Cave" municipality="Sataplia-Tskaltubo" pageId="1" pageNumber="372" specimenCount="1" stateProvince="Imereti" typeStatus="paratype">
same data as holotype, but
<collectingDate id="8B02C155FF896B2CFF4D2CE4FEDC8CCF" box="[151,274,1737,1763]" pageId="1" pageNumber="372" value="2018-09-28">
<date id="9B4638BDFF896B2CFF4D2CE4FEDC8CCF" box="[151,274,1737,1763]" pageId="1" pageNumber="372" value="2018-09-28">28.ix.2018</date>
</collectingDate>
, 1♁, (
<collectionCode id="89E986B8FF896B2CFE8C2CE4FE678CCF" box="[342,425,1737,1763]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="372">NMPC</collectionCode>
)
</materialsCitation>
<materialsCitation id="5F901420FF896B2CFE622CE4FD918CCF" ID-GBIF-Occurrence="2242680554" box="[440,607,1737,1763]" collectionCode="CPH" country="Georgia" pageId="1" pageNumber="372" specimenCount="1" specimenCount-female="1" stateProvince="Imereti" typeStatus="paratype">
and
<specimenCount id="F9FED5F4FF896B2CFE322CE4FDC58CCF" box="[488,523,1737,1763]" pageId="1" pageNumber="372" type="female">1♀</specimenCount>
(
<collectionCode id="89E986B8FF896B2CFDC02CE4FD988CCF" box="[538,598,1737,1763]" pageId="1" pageNumber="372">CPH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5F901420FF896B2CFDB12CE4FC8E8D2B" ID-GBIF-Occurrence="2242680552" collectingDate="2018-07-05" collectionCode="CPH" collectorName="E. Maghradze" country="Georgia" location="Prometheus Cave" municipality="Sataplia-Tskaltubo" pageId="1" pageNumber="372" specimenCount="1" stateProvince="Imereti" typeStatus="paratype">
<collectingCountry id="97EF5EEDFF896B2CFDB12CE4FD228CCF" box="[619,748,1737,1763]" name="Georgia" pageId="1" pageNumber="372">GEORGIA</collectingCountry>
,
<collectingRegion id="2D3CD09FFF896B2CFD2D2CE4FC578CCF" box="[759,921,1737,1763]" country="Georgia" name="Imereti" pageId="1" pageNumber="372">Imereti region</collectingRegion>
,
<collectingMunicipality id="0F238407FF896B2CFC7F2CE4FBB58CCF" box="[933,1147,1737,1763]" pageId="1" pageNumber="372">SatapliaTskaltubo</collectingMunicipality>
karst massif,
<location id="EA2748A6FF896B2CFAC22CE4FF1D8D2B" LSID="urn:lsid:plazi:treatment:6751AF6BFF896B2AFF4D2FE0FE908E01:EA2748A6FF896B2CFAC22CE4FF1D8D2B" country="Georgia" municipality="Sataplia-Tskaltubo" name="Prometheus Cave" pageId="1" pageNumber="372" stateProvince="Imereti">Prometheus Cave</location>
, dark zone,
<collectingDate id="8B02C155FF896B2CFE872CC1FE118D2B" box="[349,479,1772,1799]" pageId="1" pageNumber="372" value="2018-07-05">
<date id="9B4638BDFF896B2CFE872CC1FE118D2B" box="[349,479,1772,1799]" pageId="1" pageNumber="372" value="2018-07-05">05.vii.2018</date>
</collectingDate>
, 1♁,
<collectorName id="420D7BABFF896B2CFDC72CC0FD758D2B" box="[541,699,1773,1799]" pageId="1" pageNumber="372">E. Maghradze</collectorName>
leg. (
<collectionCode id="89E986B8FF896B2CFD262CC0FCF68D2B" box="[764,824,1773,1799]" pageId="1" pageNumber="372">CPH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5F901420FF896B2CFC962CC0FC048D07" ID-GBIF-Occurrence="2242680559" collectingDate="2018-05-12" collectionCode="SMNS" collectorName="A. Faille" country="Georgia" location="Melouri Cave" municipality="Sataplia-Tskaltubo" pageId="1" pageNumber="372" specimenCount="1" specimenCount-female="1" stateProvince="Imereti" typeStatus="paratype">
<collectingCountry id="97EF5EEDFF896B2CFC962CC0FC038D2B" box="[844,973,1773,1799]" name="Georgia" pageId="1" pageNumber="372">GEORGIA</collectingCountry>
,
<collectingRegion id="2D3CD09FFF896B2CFC032CC0FBB28D2B" box="[985,1148,1773,1799]" country="Georgia" name="Imereti" pageId="1" pageNumber="372">Imereti region</collectingRegion>
,
<collectingMunicipality id="0F238407FF896B2CFB522CC0FA908D2B" box="[1160,1374,1773,1799]" pageId="1" pageNumber="372">SatapliaTskaltubo</collectingMunicipality>
karst massif,
<location id="EA2748A6FF896B2CFF352D3CFE438D07" LSID="urn:lsid:plazi:treatment:6751AF6BFF896B2AFF4D2FE0FE908E01:EA2748A6FF896B2CFF352D3CFE438D07" box="[239,397,1809,1835]" country="Georgia" municipality="Sataplia-Tskaltubo" name="Melouri Cave" pageId="1" pageNumber="372" stateProvince="Imereti">Melouri Cave</location>
, dark zone,
<specimenCount id="F9FED5F4FF896B2CFDCD2D3CFDF48D07" box="[535,570,1809,1835]" pageId="1" pageNumber="372" type="female">1♀</specimenCount>
,
<collectingDate id="8B02C155FF896B2CFD932D3CFD0C8D07" box="[585,706,1809,1835]" pageId="1" pageNumber="372" value="2018-05-12">
<date id="9B4638BDFF896B2CFD932D3CFD0C8D07" box="[585,706,1809,1835]" pageId="1" pageNumber="372" value="2018-05-12">12.v. 2018</date>
</collectingDate>
,
<collectorName id="420D7BABFF896B2CFD162D3CFCE18D07" box="[716,815,1809,1835]" pageId="1" pageNumber="372">A. Faille</collectorName>
leg. (
<collectionCode id="89E986B8FF896B2CFCAA2D3CFC0C8D07" box="[880,962,1809,1835]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="372">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5F901420FF896B2CFC0D2D3CFE958D63" ID-GBIF-Occurrence="2242680553" collectingDate="2018-05-26" collectionCode="SMNS" collectorName="E. Maghradze" country="Georgia" location="Melouri Cave" municipality="Sataplia-Tskaltubo" pageId="1" pageNumber="372" specimenCount="1" specimenCount-female="1" stateProvince="Imereti" typeStatus="paratype">
same data, but
<collectingDate id="8B02C155FF896B2CFB5B2D3CFB3C8D07" box="[1153,1266,1809,1835]" pageId="1" pageNumber="372" value="2018-05-26">
<date id="9B4638BDFF896B2CFB5B2D3CFB3C8D07" box="[1153,1266,1809,1835]" pageId="1" pageNumber="372" value="2018-05-26">26.v.2018</date>
</collectingDate>
,
<collectorName id="420D7BABFF896B2CFB242D3CFA528D07" box="[1278,1436,1809,1835]" pageId="1" pageNumber="372">E. Maghradze</collectorName>
leg.,
<specimenCount id="F9FED5F4FF896B2CFF142D18FF3F8D63" box="[206,241,1845,1871]" pageId="1" pageNumber="372" type="female">1♀</specimenCount>
(
<collectionCode id="89E986B8FF896B2CFEDB2D18FE9D8D63" box="[257,339,1845,1871]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="372">SMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A7E24DF6FF896B28FF1D2D75FA918CF3" lastPageId="5" lastPageNumber="376" pageId="1" pageNumber="372" type="description">
<paragraph id="EF471E7DFF896B2CFF1D2D75FF1C8D97" blockId="1.[151,1436,1592,1979]" pageId="1" pageNumber="372">
<emphasis id="DD8CC26FFF896B2CFF1D2D75FE978D5F" bold="true" box="[199,345,1880,1907]" pageId="1" pageNumber="372">Description.</emphasis>
Body length, male:
<quantity id="2800B398FF896B2CFDE02D75FD108D5F" box="[570,734,1880,1907]" metricMagnitude="-3" metricUnit="m" metricValue="2.885" metricValueMax="3.02" metricValueMin="2.75" pageId="1" pageNumber="372" unit="mm" value="2.885" valueMax="3.02" valueMin="2.75">2.753.02 mm</quantity>
, female:
<quantity id="2800B398FF896B2CFC992D74FC278D5F" box="[835,1001,1881,1907]" metricMagnitude="-3" metricUnit="m" metricValue="2.815" metricValueMax="2.83" metricValueMin="2.8" pageId="1" pageNumber="372" unit="mm" value="2.815" valueMax="2.83" valueMin="2.8">2.802.83 mm</quantity>
; combined width of elytra, male:
<quantity id="2800B398FF896B2CFA872D74FF328DBB" metricMagnitude="-4" metricUnit="m" metricValue="7.65" metricValueMax="7.7" metricValueMin="7.6" pageId="1" pageNumber="372" unit="mm" value="0.765" valueMax="0.77" valueMin="0.76">0.76 0.77 mm</quantity>
, female:
<quantity id="2800B398FF896B2CFEBB2D51FDCB8DBB" box="[353,517,1916,1943]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" metricValueMax="7.9" metricValueMin="7.5" pageId="1" pageNumber="372" unit="mm" value="0.77" valueMax="0.79" valueMin="0.75">0.750.79 mm</quantity>
; colour reddishbrown, slightly shiny, whole body covered with golden setae (
<figureCitation id="77C302F8FF896B2CFAB02D50FF0A8D97" captionStart="FIGURES 78" captionStartId="4.[151,264,1089,1114]" captionTargetBox="[153,1435,177,1025]" captionTargetId="figure@4.[153,1438,177,1025]" captionTargetPageId="4" captionText="FIGURES 78. Bergrothia barbakadzei sp. nov., habitus 7, dorsal view (female); 8, ventral view (male)." httpUri="https://zenodo.org/record/3055394/files/figure.png" pageId="1" pageNumber="372">Figs 78</figureCitation>
).
</paragraph>
<caption id="BB874EF5FF8A6B2FFF4D2D84FB358DEE" box="[151,1275,1961,1986]" httpUri="https://zenodo.org/record/3055384/files/figure.png" pageId="2" pageNumber="373" startId="2.[151,264,1961,1986]" targetBox="[371,1216,185,1923]" targetPageId="2">
<paragraph id="EF471E7DFF8A6B2FFF4D2D84FB358DEE" blockId="2.[151,1275,1961,1986]" box="[151,1275,1961,1986]" pageId="2" pageNumber="373">
<emphasis id="DD8CC26FFF8A6B2FFF4D2D84FEF48DEE" bold="true" box="[151,314,1961,1986]" pageId="2" pageNumber="373">FIGURES 13.</emphasis>
Entrances of the caves sampled. 1, Datvis (Bear) Cave; 2, Prometheus Cave; 3, Melouri Cave.
</paragraph>
</caption>
<caption id="BB874EF5FF8B6B2EFF4D28D9FCD18901" httpUri="https://zenodo.org/record/3055388/files/figure.png" pageId="3" pageNumber="374" startId="3.[151,250,756,781]" targetBox="[153,1435,182,688]" targetPageId="3">
<paragraph id="EF471E7DFF8B6B2EFF4D28D9FCD18901" blockId="3.[151,1436,756,813]" pageId="3" pageNumber="374">
<emphasis id="DD8CC26FFF8B6B2EFF4D28D9FEDA8921" bold="true" box="[151,276,756,781]" pageId="3" pageNumber="374">FIGURE 4.</emphasis>
Distribution of
<taxonomicName id="28F865FEFF8B6B2EFE6628D8FD608921" authority="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč, 2019" authorityName="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč" authorityYear="2019" box="[444,686,757,781]" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="barbakadzei" status="sp. nov.">
<emphasis id="DD8CC26FFF8B6B2EFE6628D8FD608921" box="[444,686,757,781]" italics="true" pageId="3" pageNumber="374">Bergrothia barbakadzei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C6BF7F14FF8B6B2EFD6C28D8FCC68921" box="[694,776,757,781]" pageId="3" pageNumber="374" rank="species">
<emphasis id="DD8CC26FFF8B6B2EFD6C28D8FCC68921" bold="true" box="[694,776,757,781]" pageId="3" pageNumber="374">sp. nov.</emphasis>
</taxonomicNameLabel>
, Georgia, location of the Imereti region (4A); Studied localities (4B). 1, Melouri Cave; 2, Prometheus Cave and 3, Datvis Cave.
</paragraph>
</caption>
<caption id="BB874EF5FF8B6B2EFF4D2DB6FB588DF8" httpUri="https://zenodo.org/record/3055392/files/figure.png" pageId="3" pageNumber="374" startId="3.[151,264,1947,1972]" targetBox="[224,1297,873,1889]" targetPageId="3">
<paragraph id="EF471E7DFF8B6B2EFF4D2DB6FB588DF8" blockId="3.[151,1436,1947,2004]" pageId="3" pageNumber="374">
<emphasis id="DD8CC26FFF8B6B2EFF4D2DB6FEF58D98" bold="true" box="[151,315,1947,1972]" pageId="3" pageNumber="374">FIGURES 56.</emphasis>
5, Plan of Prometheus Cave (
<bibRefCitation id="8B69638CFF8B6B2EFDB72DB6FCAE8D98" author="Tatashidze, Z. K. &amp; Tsikarishvili, K. D. &amp; Jishkariani, J. M." box="[621,864,1947,1972]" pageId="3" pageNumber="374" refId="ref4318" refString="Tatashidze, Z. K., Tsikarishvili, K. D. &amp; Jishkariani, J. M. (2009 b) The Cadastre of the Karst Caves of Georgia. Petiti Publishing House, Tbilisi, 670 pp. [in Georgian]" type="book" year="2009">
Tatashidze
<emphasis id="DD8CC26FFF8B6B2EFD382DB1FCD88D9F" box="[738,790,1947,1971]" italics="true" pageId="3" pageNumber="374">et al.</emphasis>
2009b
</bibRefCitation>
); 6, Plan of Melouri Cave (
<bibRefCitation id="8B69638CFF8B6B2EFBA22DB6FAA98D9F" author="Tatashidze, Z. &amp; Tsikarishvili, K. &amp; Jishkariani, J. &amp; Jamrishvili, A. &amp; Geladze, G. &amp; Lominadze, G." box="[1144,1383,1947,1972]" pageId="3" pageNumber="374" refId="ref4268" refString="Tatashidze, Z., Tsikarishvili, K., Jishkariani, J., Jamrishvili, A., Geladze, G. &amp; Lominadze, G. (2009 a) Tskaltubo Cave System. Petiti Publishing House, Tbilisi, 72 pp. [in Georgian]" type="book" year="2009">
Tatashidze
<emphasis id="DD8CC26FFF8B6B2EFB372DB1FAEF8D9F" box="[1261,1313,1947,1971]" italics="true" pageId="3" pageNumber="374">et al.</emphasis>
2009a
</bibRefCitation>
) (the black circles show the place where the specimens of
<taxonomicName id="28F865FEFF8B6B2EFD742D96FC6C8DFF" authority="Barbakadzei" authorityName="Barbakadzei" box="[686,930,1979,2003]" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="genus">
<emphasis id="DD8CC26FFF8B6B2EFD742D96FC6C8DFF" box="[686,930,1979,2003]" italics="true" pageId="3" pageNumber="374">Bergrothia Barbakadzei</emphasis>
</taxonomicName>
<emphasis id="DD8CC26FFF8B6B2EFC732D96FC3A8DFF" bold="true" box="[937,1012,1979,2003]" pageId="3" pageNumber="374">sp. nov</emphasis>
. were sampled)
</paragraph>
</caption>
<caption id="BB874EF5FF8C6B29FF4D2E6CFB0A8E76" box="[151,1220,1089,1114]" httpUri="https://zenodo.org/record/3055394/files/figure.png" pageId="4" pageNumber="375" startId="4.[151,264,1089,1114]" targetBox="[153,1435,177,1025]" targetPageId="4">
<paragraph id="EF471E7DFF8C6B29FF4D2E6CFB0A8E76" blockId="4.[151,1220,1089,1114]" box="[151,1220,1089,1114]" pageId="4" pageNumber="375">
<emphasis id="DD8CC26FFF8C6B29FF4D2E6CFEF58E76" bold="true" box="[151,315,1089,1114]" pageId="4" pageNumber="375">FIGURES 78.</emphasis>
<taxonomicName id="28F865FEFF8C6B29FE9B2E6FFDFF8E76" authority="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč, 2019" authorityName="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč" authorityYear="2019" box="[321,561,1090,1114]" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="barbakadzei" status="sp. nov.">
<emphasis id="DD8CC26FFF8C6B29FE9B2E6FFDFF8E76" box="[321,561,1090,1114]" italics="true" pageId="4" pageNumber="375">Bergrothia barbakadzei</emphasis>
</taxonomicName>
<emphasis id="DD8CC26FFF8C6B29FDE22E6FFD418E76" bold="true" box="[568,655,1090,1114]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="C6BF7F14FF8C6B29FDE22E6FFD458E76" box="[568,651,1090,1114]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
habitus 7, dorsal view (female); 8, ventral view (male).
</paragraph>
</caption>
<paragraph id="EF471E7DFF8C6B29FF1D2EB6FDF58C6D" blockId="4.[151,1437,1179,2033]" pageId="4" pageNumber="375">
<emphasis id="DD8CC26FFF8C6B29FF1D2EB6FEC98E99" bold="true" box="[199,263,1179,1205]" pageId="4" pageNumber="375">Head</emphasis>
(
<figureCitation id="77C302F8FF8C6B29FEC32EB6FE418E99" box="[281,399,1179,1205]" captionStart="FIGURES 914" captionStartId="5.[151,264,1399,1424]" captionTargetBox="[153,1435,251,1350]" captionTargetId="figure@5.[153,1438,270,1350]" captionTargetPageId="5" captionText="FIGURES 914. Bergrothia barbakadzei sp. nov. 9, head, dorsal view (female); 10, head dorsal view with (10a) sensilla on segment IV of maxillary palpus (female); 11, antenna (female); 12, antennal segment XI (female); 13, head, ventral view (male); 14, pronotum, dorsal view (female)." httpUri="https://zenodo.org/record/3055404/files/figure.png" pageId="4" pageNumber="375">Figs 9, 13</figureCitation>
) slightly elongate, about 1.2 times as long as wide, surface between lateral carinae rough (
<figureCitation id="77C302F8FF8C6B29FF7A2E92FF2A8EF5" box="[160,228,1215,1241]" captionStart="FIGURES 914" captionStartId="5.[151,264,1399,1424]" captionTargetBox="[153,1435,251,1350]" captionTargetId="figure@5.[153,1438,270,1350]" captionTargetPageId="5" captionText="FIGURES 914. Bergrothia barbakadzei sp. nov. 9, head, dorsal view (female); 10, head dorsal view with (10a) sensilla on segment IV of maxillary palpus (female); 11, antenna (female); 12, antennal segment XI (female); 13, head, ventral view (male); 14, pronotum, dorsal view (female)." httpUri="https://zenodo.org/record/3055404/files/figure.png" pageId="4" pageNumber="375">Fig. 9</figureCitation>
); lateral vertexal carinae welldefined, reaching antennal tubercles at level of vertexal foveae and gradually weakening posteriorly; medial carina originating almost at level of constriction between head capsule and neck, and extending anteriorly to frontal excavation. Frontal excavation welldeveloped between Ushaped supraantennal tubercles; vertexal foveae naked and deep, located on each side of medial carina at midlength of it; lateral and ventrolateral setae long, 1.52.9 times longer than dorsal setae on frons and vertex; eyes strongly reduced, consisting of 34 small, pale ommatidia; clypeus large, rounded; lateral carinae from antennal tubercles to the middle of clypeus forming an angle of approximately 90°, lateral carinae of clypeus welldeveloped and reaching ocular spines; mandible stout, with five teeth; maxillary palpi (
<figureCitation id="77C302F8FF8C6B29FD252F96FC808FF9" box="[767,846,1467,1493]" captionStart="FIGURES 914" captionStartId="5.[151,264,1399,1424]" captionTargetBox="[153,1435,251,1350]" captionTargetId="figure@5.[153,1438,270,1350]" captionTargetPageId="5" captionText="FIGURES 914. Bergrothia barbakadzei sp. nov. 9, head, dorsal view (female); 10, head dorsal view with (10a) sensilla on segment IV of maxillary palpus (female); 11, antenna (female); 12, antennal segment XI (female); 13, head, ventral view (male); 14, pronotum, dorsal view (female)." httpUri="https://zenodo.org/record/3055404/files/figure.png" pageId="4" pageNumber="375">Fig. 13</figureCitation>
) small, shorter than head; palpomere I minuscule, ca. 0.20 times as long as palpomere II; palpomere II long and thickened distally, palpomere IV 2.124.00 times longer than palpomere III and 0.751.41 times longer than palpomere II. Dorsolateral sides of palpomere IV with three, welldefined sensillae (
<figureCitation id="77C302F8FF8C6B29FE7A2C0AFDE08C6D" box="[416,558,1575,1601]" captionStart="FIGURES 914" captionStartId="5.[151,264,1399,1424]" captionTargetBox="[153,1435,251,1350]" captionTargetId="figure@5.[153,1438,270,1350]" captionTargetPageId="5" captionText="FIGURES 914. Bergrothia barbakadzei sp. nov. 9, head, dorsal view (female); 10, head dorsal view with (10a) sensilla on segment IV of maxillary palpus (female); 11, antenna (female); 12, antennal segment XI (female); 13, head, ventral view (male); 14, pronotum, dorsal view (female)." httpUri="https://zenodo.org/record/3055404/files/figure.png" pageId="4" pageNumber="375">Figs 10, 10a</figureCitation>
).
</paragraph>
<paragraph id="EF471E7DFF8C6B29FF1D2C67FA418C81" blockId="4.[151,1437,1179,2033]" pageId="4" pageNumber="375">
<emphasis id="DD8CC26FFF8C6B29FF1D2C67FEF48C49" bold="true" box="[199,314,1610,1637]" pageId="4" pageNumber="375">Antennae</emphasis>
(
<figureCitation id="77C302F8FF8C6B29FE962C66FE538C49" box="[332,413,1611,1637]" captionStart="FIGURES 914" captionStartId="5.[151,264,1399,1424]" captionTargetBox="[153,1435,251,1350]" captionTargetId="figure@5.[153,1438,270,1350]" captionTargetPageId="5" captionText="FIGURES 914. Bergrothia barbakadzei sp. nov. 9, head, dorsal view (female); 10, head dorsal view with (10a) sensilla on segment IV of maxillary palpus (female); 11, antenna (female); 12, antennal segment XI (female); 13, head, ventral view (male); 14, pronotum, dorsal view (female)." httpUri="https://zenodo.org/record/3055404/files/figure.png" pageId="4" pageNumber="375">Fig. 11</figureCitation>
) long and slender; male:
<quantity id="2800B398FF8C6B29FD1E2C67FCA28C49" box="[708,876,1610,1637]" metricMagnitude="-3" metricUnit="m" metricValue="1.495" metricValueMax="1.52" metricValueMin="1.47" pageId="4" pageNumber="375" unit="mm" value="1.495" valueMax="1.52" valueMin="1.47">1.471.52 mm</quantity>
; female:
<quantity id="2800B398FF8C6B29FC0C2C67FBB08C49" box="[982,1150,1610,1637]" metricMagnitude="-3" metricUnit="m" metricValue="1.535" metricValueMax="1.59" metricValueMin="1.48" pageId="4" pageNumber="375" unit="mm" value="1.535" valueMax="1.59" valueMin="1.48">1.481.59 mm</quantity>
; all antennomeres longer than wide; scape and pedicel of subequal length, scape about 1.3 times as long as wide, pedicel about twice as long as wide; antennomere XI longer and thicker than all remaining antennomeres, 2.4 times as long as wide (
<figureCitation id="77C302F8FF8C6B29FAF52CBEFA4C8C81" box="[1327,1410,1683,1709]" captionStart="FIGURES 914" captionStartId="5.[151,264,1399,1424]" captionTargetBox="[153,1435,251,1350]" captionTargetId="figure@5.[153,1438,270,1350]" captionTargetPageId="5" captionText="FIGURES 914. Bergrothia barbakadzei sp. nov. 9, head, dorsal view (female); 10, head dorsal view with (10a) sensilla on segment IV of maxillary palpus (female); 11, antenna (female); 12, antennal segment XI (female); 13, head, ventral view (male); 14, pronotum, dorsal view (female)." httpUri="https://zenodo.org/record/3055404/files/figure.png" pageId="4" pageNumber="375">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="EF471E7DFF8C6B29FF1D2C9AFECE8D11" blockId="4.[151,1437,1179,2033]" pageId="4" pageNumber="375">
<emphasis id="DD8CC26FFF8C6B29FF1D2C9AFE8E8CFD" bold="true" box="[199,320,1719,1745]" pageId="4" pageNumber="375">Pronotum</emphasis>
(
<figureCitation id="77C302F8FF8C6B29FE952C9AFE6E8CFD" box="[335,416,1719,1745]" captionStart="FIGURES 914" captionStartId="5.[151,264,1399,1424]" captionTargetBox="[153,1435,251,1350]" captionTargetId="figure@5.[153,1438,270,1350]" captionTargetPageId="5" captionText="FIGURES 914. Bergrothia barbakadzei sp. nov. 9, head, dorsal view (female); 10, head dorsal view with (10a) sensilla on segment IV of maxillary palpus (female); 11, antenna (female); 12, antennal segment XI (female); 13, head, ventral view (male); 14, pronotum, dorsal view (female)." httpUri="https://zenodo.org/record/3055404/files/figure.png" pageId="4" pageNumber="375">Fig. 14</figureCitation>
) rounded at sides, about 1.11.2 times as long as wide; widest near middle, in middle about 1.4 times wider than anterior margin and 1.3 times wider than posterior margin; with median depression, four small antebasal foveae and welldefined lateral foveae; basal third of pronotum bearing pair of spinelike projections (
<figureCitation id="77C302F8FF8C6B29FF7A2D0EFF3D8D11" box="[160,243,1827,1853]" captionStart="FIGURES 914" captionStartId="5.[151,264,1399,1424]" captionTargetBox="[153,1435,251,1350]" captionTargetId="figure@5.[153,1438,270,1350]" captionTargetPageId="5" captionText="FIGURES 914. Bergrothia barbakadzei sp. nov. 9, head, dorsal view (female); 10, head dorsal view with (10a) sensilla on segment IV of maxillary palpus (female); 11, antenna (female); 12, antennal segment XI (female); 13, head, ventral view (male); 14, pronotum, dorsal view (female)." httpUri="https://zenodo.org/record/3055404/files/figure.png" pageId="4" pageNumber="375">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="EF471E7DFF8C6B29FF1D2D6AFAE78D85" blockId="4.[151,1437,1179,2033]" pageId="4" pageNumber="375">
<emphasis id="DD8CC26FFF8C6B29FF1D2D6AFEDD8D4D" bold="true" box="[199,275,1863,1889]" pageId="4" pageNumber="375">Elytra</emphasis>
(
<figureCitation id="77C302F8FF8C6B29FEFE2D6AFEB98D4D" box="[292,375,1862,1889]" captionStart="FIGURES 1518" captionStartId="6.[151,264,1953,1978]" captionTargetBox="[153,1435,177,1887]" captionTargetId="figure@6.[153,1438,177,1887]" captionTargetPageId="6" captionText="FIGURES 1518. Bergrothia barbakadzei sp. nov. 15, elytra (female); 16, median leg (male); 17, abdominal tergites IIVII, dorsal view (female); 18, abdominal tergites IIVII, ventral view (male)." httpUri="https://zenodo.org/record/3055410/files/figure.png" pageId="4" pageNumber="375">Fig. 15</figureCitation>
) smooth, slightly shorter than abdomen, 0.650.74 times as long as all visible abdominal tergites, slightly wider than long, about 1.3 times as long as pronotum; posterior margin truncate, posterolateral angle round; dorsally moderately convex; with three welldefined and deep basal foveae; suture welldefined.
</paragraph>
<paragraph id="EF471E7DFF8C6B28FF1D2D9FFF2F8AD7" blockId="4.[151,1437,1179,2033]" lastBlockId="5.[151,1436,153,251]" lastPageId="5" lastPageNumber="376" pageId="4" pageNumber="375">
<emphasis id="DD8CC26FFF8C6B29FF1D2D9FFEF28DE1" bold="true" box="[199,316,1970,1997]" pageId="4" pageNumber="375">Abdomen</emphasis>
(
<figureCitation id="77C302F8FF8C6B29FE942D9EFE1D8DE1" box="[334,467,1971,1997]" captionStart="FIGURES 1518" captionStartId="6.[151,264,1953,1978]" captionTargetBox="[153,1435,177,1887]" captionTargetId="figure@6.[153,1438,177,1887]" captionTargetPageId="6" captionText="FIGURES 1518. Bergrothia barbakadzei sp. nov. 15, elytra (female); 16, median leg (male); 17, abdominal tergites IIVII, dorsal view (female); 18, abdominal tergites IIVII, ventral view (male)." httpUri="https://zenodo.org/record/3055410/files/figure.png" pageId="4" pageNumber="375">Figs 17, 18</figureCitation>
) about 1.2 times longer than wide, about as wide as elytra; first visible tergite (IV) lacking basal carinae, about 1.5 times as wide as long, with wide, setose basal depression and two welldefined oblique lateral carinae; second and third visible tergites (VVI) subequal in length, fourth visible tergite (VII) about 1.7 times as long as second (V) and third (VI) visible tergites and more than twice as long as fifth visible tergite (VIII).
</paragraph>
<caption id="BB874EF5FF8D6B28FF4D2F5AFDCB8FFC" httpUri="https://zenodo.org/record/3055404/files/figure.png" pageId="5" pageNumber="376" startId="5.[151,264,1399,1424]" targetBox="[153,1435,251,1350]" targetPageId="5">
<paragraph id="EF471E7DFF8D6B28FF4D2F5AFDCB8FFC" blockId="5.[151,1436,1399,1488]" pageId="5" pageNumber="376">
<emphasis id="DD8CC26FFF8D6B28FF4D2F5AFE878FBC" bold="true" box="[151,329,1399,1424]" pageId="5" pageNumber="376">FIGURES 914.</emphasis>
<taxonomicName id="28F865FEFF8D6B28FE8A2F55FD8C8FBC" authority="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč, 2019" authorityName="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč" authorityYear="2019" box="[336,578,1400,1424]" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="barbakadzei" status="sp. nov.">
<emphasis id="DD8CC26FFF8D6B28FE8A2F55FD8C8FBC" box="[336,578,1400,1424]" italics="true" pageId="5" pageNumber="376">Bergrothia barbakadzei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C6BF7F14FF8D6B28FD902F55FD528FBC" box="[586,668,1400,1424]" pageId="5" pageNumber="376" rank="species">
<emphasis id="DD8CC26FFF8D6B28FD902F55FD528FBC" bold="true" box="[586,668,1400,1424]" pageId="5" pageNumber="376">sp. nov.</emphasis>
</taxonomicNameLabel>
9, head, dorsal view (female); 10, head dorsal view with (10a) sensilla on segment IV of maxillary palpus (female); 11, antenna (female); 12, antennal segment XI (female); 13, head, ventral view (male); 14, pronotum, dorsal view (female).
</paragraph>
</caption>
<paragraph id="EF471E7DFF8D6B28FF1D2C3CFF0D8C5F" blockId="5.[151,1437,1552,2011]" pageId="5" pageNumber="376">
<emphasis id="DD8CC26FFF8D6B28FF1D2C3CFF318C07" bold="true" box="[199,255,1553,1579]" pageId="5" pageNumber="376">Legs</emphasis>
short and slender, profemora bearing row of small and curved prone setae along subapical carinae. mesotrochanters of males with thick and strong tooth (
<figureCitation id="77C302F8FF8D6B28FD092C19FCEB8C63" box="[723,805,1588,1615]" captionStart="FIGURES 1518" captionStartId="6.[151,264,1953,1978]" captionTargetBox="[153,1435,177,1887]" captionTargetId="figure@6.[153,1438,177,1887]" captionTargetPageId="6" captionText="FIGURES 1518. Bergrothia barbakadzei sp. nov. 15, elytra (female); 16, median leg (male); 17, abdominal tergites IIVII, dorsal view (female); 18, abdominal tergites IIVII, ventral view (male)." httpUri="https://zenodo.org/record/3055410/files/figure.png" pageId="5" pageNumber="376">Fig. 16</figureCitation>
), mesotibiae with small spur in apical third in male (
<figureCitation id="77C302F8FF8D6B28FAB52C19FF788C5F" captionStart="FIGURES 1923" captionStartId="7.[151,264,782,807]" captionTargetBox="[161,1431,182,687]" captionTargetId="figure@7.[153,1438,177,687]" captionTargetPageId="7" captionText="FIGURES 1923. Mesotibiae of Bergrothia barbakadzei sp. nov. (scale bar: 250 μm). 19, male; 20, female; aedeagus of Bergrothia barbakadzei sp. nov. 21, lateral view (scale bar: 200 μm); 22, dorsal view (scale bar: 200 μm); aedeagus of Bergrothia mingrelica (Reitter). 23, dorsal view (Reitter) (scale bar: 200 μm). Redrawn from Hlaváč, 1999." httpUri="https://zenodo.org/record/3055424/files/figure.png" pageId="5" pageNumber="376">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="EF471E7DFF8D6B28FF1D2C51FD688CBB" blockId="5.[151,1437,1552,2011]" box="[199,678,1660,1687]" pageId="5" pageNumber="376">
<emphasis id="DD8CC26FFF8D6B28FF1D2C51FEF48CBB" bold="true" box="[199,314,1660,1687]" pageId="5" pageNumber="376">Aedeagus</emphasis>
asymmetrical, as in
<figureCitation id="77C302F8FF8D6B28FDFB2C51FD6C8CBB" box="[545,674,1660,1687]" captionStart="FIGURES 1923" captionStartId="7.[151,264,782,807]" captionTargetBox="[161,1431,182,687]" captionTargetId="figure@7.[153,1438,177,687]" captionTargetPageId="7" captionText="FIGURES 1923. Mesotibiae of Bergrothia barbakadzei sp. nov. (scale bar: 250 μm). 19, male; 20, female; aedeagus of Bergrothia barbakadzei sp. nov. 21, lateral view (scale bar: 200 μm); 22, dorsal view (scale bar: 200 μm); aedeagus of Bergrothia mingrelica (Reitter). 23, dorsal view (Reitter) (scale bar: 200 μm). Redrawn from Hlaváč, 1999." httpUri="https://zenodo.org/record/3055424/files/figure.png" pageId="5" pageNumber="376">Figs 2122</figureCitation>
.
</paragraph>
<paragraph id="EF471E7DFF8D6B28FF1D2C8DFA918CF3" blockId="5.[151,1437,1552,2011]" pageId="5" pageNumber="376">
<emphasis id="DD8CC26FFF8D6B28FF1D2C8DFE788C97" bold="true" box="[199,438,1696,1723]" pageId="5" pageNumber="376">Sexual dimorphism.</emphasis>
Mesotrochanters of male with thick and strong tooth (
<figureCitation id="77C302F8FF8D6B28FBCB2C8DFBAA8C97" box="[1041,1124,1696,1723]" captionStart="FIGURES 1518" captionStartId="6.[151,264,1953,1978]" captionTargetBox="[153,1435,177,1887]" captionTargetId="figure@6.[153,1438,177,1887]" captionTargetPageId="6" captionText="FIGURES 1518. Bergrothia barbakadzei sp. nov. 15, elytra (female); 16, median leg (male); 17, abdominal tergites IIVII, dorsal view (female); 18, abdominal tergites IIVII, ventral view (male)." httpUri="https://zenodo.org/record/3055410/files/figure.png" pageId="5" pageNumber="376">Fig. 16</figureCitation>
); in females simple, lacking tooth. Mesotibia with small spur on apical third in males (
<figureCitation id="77C302F8FF8D6B28FCFA2CE9FCBD8CF3" box="[800,883,1732,1759]" captionStart="FIGURES 1923" captionStartId="7.[151,264,782,807]" captionTargetBox="[161,1431,182,687]" captionTargetId="figure@7.[153,1438,177,687]" captionTargetPageId="7" captionText="FIGURES 1923. Mesotibiae of Bergrothia barbakadzei sp. nov. (scale bar: 250 μm). 19, male; 20, female; aedeagus of Bergrothia barbakadzei sp. nov. 21, lateral view (scale bar: 200 μm); 22, dorsal view (scale bar: 200 μm); aedeagus of Bergrothia mingrelica (Reitter). 23, dorsal view (Reitter) (scale bar: 200 μm). Redrawn from Hlaváč, 1999." httpUri="https://zenodo.org/record/3055424/files/figure.png" pageId="5" pageNumber="376">Fig. 19</figureCitation>
); in females simple, lacking tooth (
<figureCitation id="77C302F8FF8D6B28FB242CE9FA9C8CF3" box="[1278,1362,1732,1759]" captionStart="FIGURES 1923" captionStartId="7.[151,264,782,807]" captionTargetBox="[161,1431,182,687]" captionTargetId="figure@7.[153,1438,177,687]" captionTargetPageId="7" captionText="FIGURES 1923. Mesotibiae of Bergrothia barbakadzei sp. nov. (scale bar: 250 μm). 19, male; 20, female; aedeagus of Bergrothia barbakadzei sp. nov. 21, lateral view (scale bar: 200 μm); 22, dorsal view (scale bar: 200 μm); aedeagus of Bergrothia mingrelica (Reitter). 23, dorsal view (Reitter) (scale bar: 200 μm). Redrawn from Hlaváč, 1999." httpUri="https://zenodo.org/record/3055424/files/figure.png" pageId="5" pageNumber="376">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A7E24DF6FF8D6B28FF1D2CC5FD508DBF" pageId="5" pageNumber="376" type="diagnosis">
<paragraph id="EF471E7DFF8D6B28FF1D2CC5FD508DBF" blockId="5.[151,1437,1552,2011]" pageId="5" pageNumber="376">
<emphasis id="DD8CC26FFF8D6B28FF1D2CC5FE018D2F" bold="true" box="[199,463,1768,1795]" pageId="5" pageNumber="376">Differential diagnosis.</emphasis>
The new species strongly resembles
<taxonomicName id="28F865FEFF8D6B28FCAC2CC4FBC08D2F" baseAuthorityName="Reitter" baseAuthorityYear="1885" box="[886,1038,1769,1795]" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="mingrelica">
<emphasis id="DD8CC26FFF8D6B28FCAC2CC4FBC08D2F" box="[886,1038,1769,1795]" italics="true" pageId="5" pageNumber="376">B. mingrelica</emphasis>
</taxonomicName>
by head roughness between lateral carinae, first visible abdominal tergite (IV) shorter than elytra, aedeagus with a dorsoapical brush of setae, and by a similar body size. The new species differs from
<taxonomicName id="28F865FEFF8D6B28FD762D1CFC8D8D67" baseAuthorityName="Reitter" baseAuthorityYear="1885" box="[684,835,1841,1867]" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="mingrelica">
<emphasis id="DD8CC26FFF8D6B28FD762D1CFC8D8D67" box="[684,835,1841,1867]" italics="true" pageId="5" pageNumber="376">B. mingrelica</emphasis>
</taxonomicName>
in the lack of a dorsobasal brush of setae on aedeagus (
<figureCitation id="77C302F8FF8D6B28FF452D79FEED8D43" box="[159,291,1876,1903]" captionStart="FIGURES 1923" captionStartId="7.[151,264,782,807]" captionTargetBox="[161,1431,182,687]" captionTargetId="figure@7.[153,1438,177,687]" captionTargetPageId="7" captionText="FIGURES 1923. Mesotibiae of Bergrothia barbakadzei sp. nov. (scale bar: 250 μm). 19, male; 20, female; aedeagus of Bergrothia barbakadzei sp. nov. 21, lateral view (scale bar: 200 μm); 22, dorsal view (scale bar: 200 μm); aedeagus of Bergrothia mingrelica (Reitter). 23, dorsal view (Reitter) (scale bar: 200 μm). Redrawn from Hlaváč, 1999." httpUri="https://zenodo.org/record/3055424/files/figure.png" pageId="5" pageNumber="376">Figs 2122</figureCitation>
), which is present in
<taxonomicName id="28F865FEFF8D6B28FDCD2D78FD7E8D43" baseAuthorityName="Reitter" baseAuthorityYear="1885" box="[535,688,1877,1903]" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="mingrelica">
<emphasis id="DD8CC26FFF8D6B28FDCD2D78FD7E8D43" box="[535,688,1877,1903]" italics="true" pageId="5" pageNumber="376">B. mingrelica</emphasis>
</taxonomicName>
(
<figureCitation id="77C302F8FF8D6B28FD1B2D79FCDD8D43" box="[705,787,1876,1903]" captionStart="FIGURES 1923" captionStartId="7.[151,264,782,807]" captionTargetBox="[161,1431,182,687]" captionTargetId="figure@7.[153,1438,177,687]" captionTargetPageId="7" captionText="FIGURES 1923. Mesotibiae of Bergrothia barbakadzei sp. nov. (scale bar: 250 μm). 19, male; 20, female; aedeagus of Bergrothia barbakadzei sp. nov. 21, lateral view (scale bar: 200 μm); 22, dorsal view (scale bar: 200 μm); aedeagus of Bergrothia mingrelica (Reitter). 23, dorsal view (Reitter) (scale bar: 200 μm). Redrawn from Hlaváč, 1999." httpUri="https://zenodo.org/record/3055424/files/figure.png" pageId="5" pageNumber="376">Fig. 23</figureCitation>
) and the presence of pale and small ommatidia, while
<taxonomicName id="28F865FEFF8D6B28FA5E2D78FEC18DBF" baseAuthorityName="Reitter" baseAuthorityYear="1885" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="mingrelica">
<emphasis id="DD8CC26FFF8D6B28FA5E2D78FEC18DBF" italics="true" pageId="5" pageNumber="376">B. mingrelica</emphasis>
</taxonomicName>
has dark and welldeveloped ones.
</paragraph>
</subSubSection>
<subSubSection id="A7E24DF6FF8D6B28FF1D2DB0FD8A8DF7" pageId="5" pageNumber="376" type="etymology">
<paragraph id="EF471E7DFF8D6B28FF1D2DB0FD8A8DF7" blockId="5.[151,1437,1552,2011]" pageId="5" pageNumber="376">
<emphasis id="DD8CC26FFF8D6B28FF1D2DB0FE808D9B" bold="true" box="[199,334,1949,1975]" pageId="5" pageNumber="376">Etymology.</emphasis>
Patronymic, dedicated to Mr. Valeri Barbakadze, Georgian speleologist and cave diver, who helped us during the expeditions in the caves.
</paragraph>
</subSubSection>
<caption id="BB874EF5FF8E6B2BFF4D2D8CFCB78DF6" httpUri="https://zenodo.org/record/3055410/files/figure.png" pageId="6" pageNumber="377" startId="6.[151,264,1953,1978]" targetBox="[153,1435,177,1887]" targetPageId="6">
<paragraph id="EF471E7DFF8E6B2BFF4D2D8CFCB78DF6" blockId="6.[151,1436,1953,2010]" pageId="6" pageNumber="377">
<emphasis id="DD8CC26FFF8E6B2BFF4D2D8CFE818D96" bold="true" box="[151,335,1953,1978]" pageId="6" pageNumber="377">FIGURES 1518</emphasis>
.
<taxonomicName id="28F865FEFF8E6B2BFE862D8FFD808D96" authority="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč, 2019" authorityName="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč" authorityYear="2019" box="[348,590,1954,1978]" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="barbakadzei" status="sp. nov.">
<emphasis id="DD8CC26FFF8E6B2BFE862D8FFD808D96" box="[348,590,1954,1978]" italics="true" pageId="6" pageNumber="377">Bergrothia barbakadzei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C6BF7F14FF8E6B2BFD8F2D8FFD698D96" box="[597,679,1954,1978]" pageId="6" pageNumber="377" rank="species">
<emphasis id="DD8CC26FFF8E6B2BFD8F2D8FFD698D96" bold="true" box="[597,679,1954,1978]" pageId="6" pageNumber="377">sp. nov.</emphasis>
</taxonomicNameLabel>
15, elytra (female); 16, median leg (male); 17, abdominal tergites IIVII, dorsal view (female); 18, abdominal tergites IIVII, ventral view (male).
</paragraph>
</caption>
<caption id="BB874EF5FF8F6B2AFF4D2923FBAA894A" httpUri="https://zenodo.org/record/3055424/files/figure.png" pageId="7" pageNumber="378" startId="7.[151,264,782,807]" targetBox="[161,1431,182,687]" targetPageId="7">
<paragraph id="EF471E7DFF8F6B2AFF4D2923FBAA894A" blockId="7.[151,1437,782,871]" pageId="7" pageNumber="378">
<emphasis id="DD8CC26FFF8F6B2AFF4D2923FE9B890B" bold="true" box="[151,341,782,807]" pageId="7" pageNumber="378">FIGURES 1923.</emphasis>
Mesotibiae of
<taxonomicName id="28F865FEFF8F6B2AFE342923FD11890A" authority="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč, 2019" authorityName="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč" authorityYear="2019" box="[494,735,782,806]" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="barbakadzei" status="sp. nov.">
<emphasis id="DD8CC26FFF8F6B2AFE342923FD11890A" box="[494,735,782,806]" italics="true" pageId="7" pageNumber="378">Bergrothia barbakadzei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C6BF7F14FF8F6B2AFD3C2923FCF9890A" box="[742,823,782,806]" pageId="7" pageNumber="378" rank="species">
<emphasis id="DD8CC26FFF8F6B2AFD3C2923FCF9890A" bold="true" box="[742,823,782,806]" pageId="7" pageNumber="378">sp. nov.</emphasis>
</taxonomicNameLabel>
(scale bar: 250 μm). 19, male; 20, female; aedeagus of
<taxonomicName id="28F865FEFF8F6B2AFAAA2922FEAA896A" authority="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč, 2019" authorityName="Maghradze &amp; Faille &amp; Barjadze &amp; Hlaváč" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="barbakadzei" status="sp. nov.">
<emphasis id="DD8CC26FFF8F6B2AFAAA2922FEAA896A" italics="true" pageId="7" pageNumber="378">Bergrothia barbakadzei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C6BF7F14FF8F6B2AFEB12903FE72896A" box="[363,444,814,838]" pageId="7" pageNumber="378" rank="species">
<emphasis id="DD8CC26FFF8F6B2AFEB12903FE72896A" bold="true" box="[363,444,814,838]" pageId="7" pageNumber="378">sp. nov.</emphasis>
</taxonomicNameLabel>
21, lateral view (scale bar: 200 μm); 22, dorsal view (scale bar: 200 μm); aedeagus of
<taxonomicName id="28F865FEFF8F6B2AFAF52903FEAD894B" authority="(Reitter)" authorityName="Reitter" baseAuthorityName="Reitter" baseAuthorityYear="1885" class="Insecta" family="Staphylinidae" genus="Bergrothia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="mingrelica">
<emphasis id="DD8CC26FFF8F6B2AFAF52903FECA894A" italics="true" pageId="7" pageNumber="378">Bergrothia mingrelica</emphasis>
(Reitter)
</taxonomicName>
. 23, dorsal view (Reitter) (scale bar: 200 μm). Redrawn from Hlaváč, 1999.
</paragraph>
</caption>
<subSubSection id="A7E24DF6FF8F6B2AFF1D2985FAA489C9" pageId="7" pageNumber="378" type="biology_ecology">
<paragraph id="EF471E7DFF8F6B2AFF1D2985FAA489C9" blockId="7.[151,1436,935,1069]" pageId="7" pageNumber="378">
<emphasis id="DD8CC26FFF8F6B2AFF1D2985FEE289EE" bold="true" box="[199,300,936,962]" pageId="7" pageNumber="378">Ecology.</emphasis>
The new species was collected on guano, under plant debris and stones as well as on wet walls in dark zones of Prometheus, Datvis and Melouri caves. The presence of eyes suggests that it is a troglophilic species.
</paragraph>
</subSubSection>
<subSubSection id="A7E24DF6FF8F6B2AFF1D29C2FE908E01" pageId="7" pageNumber="378" type="distribution">
<paragraph id="EF471E7DFF8F6B2AFF1D29C2FE908E01" blockId="7.[151,1436,935,1069]" pageId="7" pageNumber="378">
<emphasis id="DD8CC26FFF8F6B2AFF1D29C2FEAD8E26" bold="true" box="[199,355,1007,1034]" pageId="7" pageNumber="378">Distribution:</emphasis>
The species is apparently endemic to
<collectingCountry id="97EF5EEDFF8F6B2AFCD129C2FCA68E25" box="[779,872,1007,1033]" name="Georgia" pageId="7" pageNumber="378">Georgia</collectingCountry>
,
<collectingRegion id="2D3CD09FFF8F6B2AFCA829C2FBDA8E25" box="[882,1044,1007,1033]" country="Georgia" name="Imereti" pageId="7" pageNumber="378">Imereti region</collectingRegion>
, Tskaltubo district, SatapliaTskal-tubo karst massif.
</paragraph>
</subSubSection>
</treatment>
</document>