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<document ID-CLB-Dataset="122795" ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-GBIF-Dataset="5f938945-980c-433c-942a-f05d39529a25" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende &amp; Pinto-da-Rocha, Ricardo" docDate="2022" docId="600BA0C567F5516BBC1AAB18D4488B84" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Ayacucho glauberrochai Benedetti &amp; Pinto-da-Rocha 2022, sp. nov." docType="treatment" docUuid="9347FBBA-3028-4F71-9ACE-668A0FEB159B" docUuidSource="ZooBank" docVersion="5" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1732651490011" updateUser="ExternalLinkService">
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<mods:title id="8CD89F0084CF16430856CB4B2C28DF32">Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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<mods:name id="A57DAB5F543FA72BDD978D89BA2FC322" type="personal">
<mods:role id="8BF8F033B5BE73060A7F6F0B6C702E95">
<mods:roleTerm id="C7B078FC32998055F8AD86B7127F14A2">Author</mods:roleTerm>
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<mods:namePart id="9CDEEEB83BDA4EAC8C685B6D0DE5A84D">Benedetti, Alipio Rezende</mods:namePart>
<mods:affiliation id="BD1070DA8B41F758C18DB071FFB38612">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil &amp; Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
<mods:nameIdentifier id="CDEA6A3601D7C648629A9AEE3CE86E64" type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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<mods:namePart id="EF0B722E4DF40F58E05D7DCDB09CA599">Pinto-da-Rocha, Ricardo</mods:namePart>
<mods:nameIdentifier id="ED91E9B134128C2379F6918F6FB28C08" type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
<mods:affiliation id="C12B7F18F9E36AC757BE7B6A723BCA84">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:title id="A291F26E8BE8492EF7C5E1BF2CE1B922">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:part id="F8863456822858EFCE88B4D308D48861">
<mods:date id="1B58835DA17FBFE03B6924B85F5B7BE2">2022</mods:date>
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<mods:number id="7A69C85C91DDA2D316BD71432425F175">2022-08-16</mods:number>
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<treatment id="600BA0C567F5516BBC1AAB18D4488B84" ID-GBIF-Taxon="199468694" LSID="urn:lsid:zoobank.org:act:9347FBBA-3028-4F71-9ACE-668A0FEB159B" httpUri="http://treatment.plazi.org/id/600BA0C567F5516BBC1AAB18D4488B84" lastPageNumber="309" pageId="0" pageNumber="309" scope_family="Metasarcidae" scope_order="Opiliones" scope_subOrder="Laniatores">
<subSubSection id="D4BF487085067DBA9691C001E6ED3CBC" pageId="0" pageNumber="309" type="nomenclature">
<paragraph id="1703D3A5A1C1291E9E004567E8FDA57D" pageId="0" pageNumber="309">
3.12.
<taxonomicName id="07F621BE3CA3CDC6EA96392359BCF097" ID-CoL="9X665" LSID="http://zoobank.org/9347FBBA-3028-4F71-9ACE-668A0FEB159B" authority="Benedetti &amp; Pinto-da-Rocha, 2022" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho glauberrochai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="glauberrochai" status="sp. nov.">Ayacucho glauberrochai</taxonomicName>
<taxonomicNameLabel id="1C3FF3F0AE21F1A4C7EAB9C09E46BFC3" pageId="0" pageNumber="309">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="5A039EDB78C39C460E11FB04ACBD8A32" pageId="0" pageNumber="309" type="description">
<paragraph id="179F8212499FD2269CEBAD3A49BEC372" pageId="0" pageNumber="309">
<figureCitation id="DFC8286AC77D175E7EBD101B3074A41C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal of Ayacucho. A A. bambamarca (Roewer, 1957) comb. nov., male; B A. glauberrochai sp. nov., male; C A. inermis (Roewer, 1957) comb. nov., male; D A. insignitus (Roewer, 1956) comb. nov., female; E A. pasolinii sp. nov., male; F A. pomacocha sp. nov., male; G A. querococha sp. nov., male; H A. roeweri nom. nov., female; I A. silvae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure2" httpUri="https://binary.pensoft.net/fig/729876" pageId="0" pageNumber="309">Figs 2B</figureCitation>
<figureCitation id="E2FE07B96361F048CD5BB677AD34D7FE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">, 7C, D</figureCitation>
<figureCitation id="2165A62E9F4EE6CF1CA53A15C83D4A07" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Penis of Ayacucho, dorsal, lateral and ventral views, respectively. A - C A. spiniger (Roewer, 1957) comb. nov.; D - F A. glauberrochai sp. nov.; G - I A. tapacocha nom. nov.; Legend bars = 0.1 mm" figureDoi="10.3897/asp.80.e73829.figure16" httpUri="https://binary.pensoft.net/fig/729890" pageId="0" pageNumber="309">, 16D-F</figureCitation>
<figureCitation id="47D217339C1A2440C093F7B398E2ABC8" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Live specimens of Ayacucho. A: A. glauberrochai sp. nov., male. B: female. C: A. pomacocha sp. nov., male. D, female. E: A. silvae sp. nov., male. F: female. G: A. pasolinii sp. nov., male. H: A. spielbergi sp. nov., male." figureDoi="10.3897/asp.80.e73829.figure22" httpUri="https://binary.pensoft.net/fig/729896" pageId="0" pageNumber="309">, 22A, B</figureCitation>
<figureCitation id="5E8A498697524CE9C9BE7F204EAD1F35" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">, 28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FC4246337C29DB54D300D8D7F5398C75" pageId="0" pageNumber="309" type="description">
<paragraph id="4A2DC1907987FF002736E65A1D635C45" pageId="0" pageNumber="309">Description.</paragraph>
<paragraph id="14AFBAC113680A565A6B90FBDA8A3EF9" pageId="0" pageNumber="309">
<emphasis id="3D7D26A500E8561BDEE97F0AD1F5E650" bold="true" pageId="0" pageNumber="309">
MALE:
<emphasis id="E564AB302BAF62ABF3DA539511A25EA2" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis id="8CF2408C50A6E9AD888B112D0308022D" italics="true" pageId="0" pageNumber="309">n</emphasis>
=9) DSW: 3.3-3.4 (3.3); DSL: 4.6-5.2 (4.6); CL: 1.5-2.0 (1.5). FIVL: 5.0-5.3 (5.0). ChL: 1.7-3.0 (2.1).
<emphasis id="3CAD2591151B851A546B1466159CD780" bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
: (Fig.
<figureCitation id="94C2D4666054BD763B3B58E56EEC13F8" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Live specimens of Ayacucho. A: A. glauberrochai sp. nov., male. B: female. C: A. pomacocha sp. nov., male. D, female. E: A. silvae sp. nov., male. F: female. G: A. pasolinii sp. nov., male. H: A. spielbergi sp. nov., male." figureDoi="10.3897/asp.80.e73829.figure22" httpUri="https://binary.pensoft.net/fig/729896" pageId="0" pageNumber="309">22A</figureCitation>
): Predominantly orange, with two longitudinal black spots in the lateral portion of areas I-IV. Lateral margins of the dorsal scutum yellowish, with small black spots. Posterior margin of the dorsal scutum, free tergites and legs predominantly black. Chelicerae and pedipalpus orange and reddish.
<emphasis id="DF429A73974D804313BA608965109825" bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
: (Fig.
<figureCitation id="719B1F743CB9264A4A8AA23ABA50D28A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal of Ayacucho. A A. bambamarca (Roewer, 1957) comb. nov., male; B A. glauberrochai sp. nov., male; C A. inermis (Roewer, 1957) comb. nov., male; D A. insignitus (Roewer, 1956) comb. nov., female; E A. pasolinii sp. nov., male; F A. pomacocha sp. nov., male; G A. querococha sp. nov., male; H A. roeweri nom. nov., female; I A. silvae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure2" httpUri="https://binary.pensoft.net/fig/729876" pageId="0" pageNumber="309">2B</figureCitation>
) Alpha-type DSS, with the constrictions weakly marked (when compared to most
<taxonomicName id="B7804E004B1450038F362F783AC3D145" authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="A62E18B9E8A562E905092A86008DB845" italics="true" pageId="0" pageNumber="309">Ayacucho</emphasis>
</taxonomicName>
spp.). Anterior margin of carapace covered with granules in all its width. Ocularium unarmed, smooth or with a few granules in some individuals. Carapace with scattered granules distributed throughout its surface. Areas I-IV with one pair of small median tubercles; densely granulate. Posterior margin of dorsal scutum with a median row of 6-8 (6) acuminate tubercles (larger than tubercles of areas I-IV). Free tergites I-III with a row of tubercles similar to those on the posterior margin of dorsal scutum.
<emphasis id="CF33957DE2690E89CD7872401064479E" bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
: (Fig.
<figureCitation id="637C1CE97DAABCEE9B0EB0C43894E662" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal of Ayacucho. A A. bambamarca (Roewer, 1957) comb. nov., male; B A. glauberrochai sp. nov., male; C A. inermis (Roewer, 1957) comb. nov., male; D A. insignitus (Roewer, 1956) comb. nov., female; E A. pasolinii sp. nov., male; F A. pomacocha sp. nov., male; G A. querococha sp. nov., male; H A. roeweri nom. nov., female; I A. silvae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure2" httpUri="https://binary.pensoft.net/fig/729876" pageId="0" pageNumber="309">2B</figureCitation>
) Swollen in some males (not swollen in the holotype). Segment I covered with granules throughout their length. Segment II predominantly smooth; finger with three teeth. Segment III with two teeth.
<taxonomicName id="EED251198FF647822D6FCF1B3DC07A5F" genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis id="922FF4794FF779636D2A41F706215963" bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
</taxonomicName>
: With granules sparsely distributed on the dorsal surface of the femur and patella. Trochanter with a distal ventral setiferous tubercle. Femur with a ventrobasal setiferous tubercle and a row of 5-6 (5) small ventromedial setiferous tubercles. Tibia: prolateral IIII, retrolateral IIII. Tarsus: prolateral IIi, retrolateral Iii.
<emphasis id="F55B13134AE2226B7ECC42622BE22DAD" bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
: Coxa I with scattered small tubercles; coxae II-III, predominantly smooth, with sparse granules; Coxa IV with denser granulation than coxae II-III. Genital area smooth. Free sternites I-IV and anal operculum with granules sparsely distributed.
<emphasis id="921963B6833A07E6F89D524232E99B72" bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
: (Figs
<figureCitation id="5FE0D50A75366F024EF830F2CCE7F68C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal of Ayacucho. A A. bambamarca (Roewer, 1957) comb. nov., male; B A. glauberrochai sp. nov., male; C A. inermis (Roewer, 1957) comb. nov., male; D A. insignitus (Roewer, 1956) comb. nov., female; E A. pasolinii sp. nov., male; F A. pomacocha sp. nov., male; G A. querococha sp. nov., male; H A. roeweri nom. nov., female; I A. silvae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure2" httpUri="https://binary.pensoft.net/fig/729876" pageId="0" pageNumber="309">2B</figureCitation>
,
<figureCitation id="6A2D3D4CDD3F86CC0ED50689D4FB320D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7C, D</figureCitation>
) Coxa I with a prolateral apophysis; coxa II with a retrolateral and a prolateral apophysis. Coxa IV densely granulate. Trochanters I-IV with few granules, unarmed. Femora I-IV, with granules distributed throughout its length. Tarsal formula: (
<emphasis id="833FEA348F8DD3B13A5FB40685B0908A" italics="true" pageId="0" pageNumber="309">n</emphasis>
= 9) 7-8 (7), 11, 7, 8.
<emphasis id="6DE4EAB75956C8081002F7A7C653B2D9" bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
: (Fig.
<figureCitation id="5986EB6A6A077D6438387B5278F5AD43" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Penis of Ayacucho, dorsal, lateral and ventral views, respectively. A - C A. spiniger (Roewer, 1957) comb. nov.; D - F A. glauberrochai sp. nov.; G - I A. tapacocha nom. nov.; Legend bars = 0.1 mm" figureDoi="10.3897/asp.80.e73829.figure16" httpUri="https://binary.pensoft.net/fig/729890" pageId="0" pageNumber="309">16D-F</figureCitation>
) VP subrectangular; distal margin with a V-notch, and with conspicuous laterodistal projections; slightly curved in lateral view. MS C1-C7 subdistal long and curved; MS A1 long and straight; medially placed; MS B1 sub basal long and straight; MS D1 subdistal very short, dorsally placed. Lateral sacs long and apically acuminate; with long T3-like microsetae. Stylus apically robust and with projections. Dorsal process absent. Promontory convex. -
<emphasis id="CD9EFF2C1120EFB9543F78894E72ECA9" bold="true" pageId="0" pageNumber="309">
FEMALE:
<emphasis id="B663D0E6E30E94F82FB90FB65F1947D3" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis id="90CB3FC174C4FC225E3B975CDA2F4B0A" italics="true" pageId="0" pageNumber="309">n</emphasis>
=10) DSW: 3.5-4.0; DSL: 4.5-5.2; CL: 1.6-1.8. FIVL: 4.7-5.6. ChL: 1.7-2.1. (Fig.
<figureCitation id="1456C4B00ABFAAFF9F8EED8414D2F610" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Live specimens of Ayacucho. A: A. glauberrochai sp. nov., male. B: female. C: A. pomacocha sp. nov., male. D, female. E: A. silvae sp. nov., male. F: female. G: A. pasolinii sp. nov., male. H: A. spielbergi sp. nov., male." figureDoi="10.3897/asp.80.e73829.figure22" httpUri="https://binary.pensoft.net/fig/729896" pageId="0" pageNumber="309">22B</figureCitation>
) Chelicerae not swollen. Strongly similar to small males. Tarsal segmentation: (
<emphasis id="5D1EC9509AD31BA9493D4921EEC324BB" italics="true" pageId="0" pageNumber="309">n</emphasis>
=10) 6, 10-11, 7, 8.
</paragraph>
<caption id="15D7363B7EB72BAA9112C300FDB8B232" doi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309" start="Figure 5" startId="F5">
<paragraph id="0C0C12C5A07322D557F900B958654DFB" pageId="0" pageNumber="309">
<emphasis id="B2AE6CC5A673B051296DE26B095D7030" bold="true" pageId="0" pageNumber="309">Figure 5.</emphasis>
Habitus, dorsal of
<taxonomicName id="7BD59EF0B45132863D2A1C974340F3A3" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="819244BE5A1A58A2DCC7C0CE89F0DE59" italics="true" pageId="0" pageNumber="309">Metasarcus</emphasis>
</taxonomicName>
.
<emphasis id="C6D5AFDFD6BA00E8FA5D40F8E90F8607" bold="true" pageId="0" pageNumber="309">A</emphasis>
<taxonomicName id="945BA3FF41B80BD7C4E39BDA3665B1DE" class="Arachnida" kingdom="Animalia" lsidName="M. beni" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="beni">
<emphasis id="27101D52B7EAA8DBB6B659A88A90C788" italics="true" pageId="0" pageNumber="309">M. beni</emphasis>
</taxonomicName>
<emphasis id="434C19F8B6A248450C43FB56B508777E" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, male;
<emphasis id="64755413DD22685F1424C85AE80AC38B" bold="true" pageId="0" pageNumber="309">B</emphasis>
<taxonomicName id="AA52164225269F96AEA71D3B4CE8A7A8" class="Arachnida" kingdom="Animalia" lsidName="M. bergmani" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bergmani">
<emphasis id="610869118C54EC3D7D97B6C49714476F" italics="true" pageId="0" pageNumber="309">M. bergmani</emphasis>
</taxonomicName>
<emphasis id="EA8F5A02FE0D67689069816B303B4B08" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, male;
<emphasis id="CF2885728C37B16C9D57E1D4DCF99D03" bold="true" pageId="0" pageNumber="309">C</emphasis>
<taxonomicName id="B289FC04F9067AB132A9EE3152D27EAA" class="Arachnida" kingdom="Animalia" lsidName="M. bolivianus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bolivianus">
<emphasis id="E9FC9D69FDCC27E15AEFD92DA46E9A12" italics="true" pageId="0" pageNumber="309">M. bolivianus</emphasis>
</taxonomicName>
Roewer, 1913, female;
<emphasis id="4E14ED3DED459C48B83AD86010182DDD" bold="true" pageId="0" pageNumber="309">D</emphasis>
<taxonomicName id="7793C70219CE876AB15A29D7F5C4E689" class="Arachnida" kingdom="Animalia" lsidName="M. clavifemur" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="clavifemur">
<emphasis id="68C8F808EA0D7C8E0F07E00FF3E6D3D4" italics="true" pageId="0" pageNumber="309">M. clavifemur</emphasis>
</taxonomicName>
(Roewer, 1929), male;
<emphasis id="1757058237879E8DEC920C9D91EEE0E4" bold="true" pageId="0" pageNumber="309">E</emphasis>
<taxonomicName id="52F2F5464CC54834107AD2CCD5A9B3E1" class="Arachnida" kingdom="Animalia" lsidName="M. fellinii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="fellinii">
<emphasis id="4A0231E78BE34A01670A870FDF246EF4" italics="true" pageId="0" pageNumber="309">M. fellinii</emphasis>
</taxonomicName>
<emphasis id="908110666B0C64DD47D41C11D47E0D0B" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, male;
<emphasis id="42400C0C8B93E20B17D61AD2D3F4E75C" bold="true" pageId="0" pageNumber="309">F</emphasis>
<taxonomicName id="B66724943172462362CD0BAAA5E96B25" class="Arachnida" kingdom="Animalia" lsidName="M. kurosawai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="kurosawai">
<emphasis id="6FB823A570DE277FC921DC4703C33BE6" italics="true" pageId="0" pageNumber="309">M. kurosawai</emphasis>
</taxonomicName>
<emphasis id="2C5377278ED8437B59EC6B68F225060A" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, male;
<emphasis id="7F1D03864E62A77072F00B082D3768A0" bold="true" pageId="0" pageNumber="309">G</emphasis>
<taxonomicName id="68952800BE4C9BA15A0B15588DB4344D" class="Arachnida" kingdom="Animalia" lsidName="M. limachii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="limachii">
<emphasis id="8CA02D563CD94B51403012BA71F967CD" italics="true" pageId="0" pageNumber="309">M. limachii</emphasis>
</taxonomicName>
<emphasis id="F16898C4FB5D445FEA34785DDEAAD6E3" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, male;
<emphasis id="D143A873B94B7BB92298AD917FCFC096" bold="true" pageId="0" pageNumber="309">H</emphasis>
<taxonomicName id="8E5DC0192739775F72B8002379A26F5D" class="Arachnida" kingdom="Animalia" lsidName="M. trispinosus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="trispinosus">
<emphasis id="0BB6D180D034A8C0A1C191CA8249F3B8" italics="true" pageId="0" pageNumber="309">M. trispinosus</emphasis>
</taxonomicName>
<emphasis id="3CBE65664793CAFA483DDF47A243E902" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, male;
<emphasis id="4033DB768887A47994D7C118413C7603" bold="true" pageId="0" pageNumber="309">I</emphasis>
<taxonomicName id="1B0C69E36BAEA834D2CC26EFF2930355" class="Arachnida" kingdom="Animalia" lsidName="M. vacafloresae" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="vacafloresae">
<emphasis id="30BEBC65D403492CB6940E31F4A05396" italics="true" pageId="0" pageNumber="309">M. vacafloresae</emphasis>
</taxonomicName>
<emphasis id="D8FDC3ED9F180973160A5B21FB72CAF4" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, male; Legend bars = 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D70EC2671A476B6411F8542A8FDFC687" pageId="0" pageNumber="309" type="diagnosis">
<paragraph id="6C3712BE6CCDCCAD29F78D9E6658AAA9" pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph id="4A85A9FD24DA9F89C52CCF8F37FFA3D5" pageId="0" pageNumber="309">
Similar to
<taxonomicName id="6D631801FF97782884E802A8988BA905" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho pomacocha" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="pomacocha">
<emphasis id="C37C212D9D66C0EDF28A4C1673E29165" italics="true" pageId="0" pageNumber="309">Ayacucho pomacocha</emphasis>
</taxonomicName>
<emphasis id="46F710152E26FCF5580B9024C24D7A80" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
,
<taxonomicName id="689710A34FDFCF75ABF8939E1B368677" lsidName="A. silvae" pageId="0" pageNumber="309" rank="species" species="silvae">
<emphasis id="759E77DF5B2D0A997E88498302DCD5D2" italics="true" pageId="0" pageNumber="309">A. silvae</emphasis>
</taxonomicName>
<emphasis id="1B27C0E1E1C3CFC5A555BC984360DEB6" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
,
<taxonomicName id="DFB97C449A346D70BABE6E19980FCE40" lsidName="A. titschacki" pageId="0" pageNumber="309" rank="species" species="titschacki">
<emphasis id="A48247C8A6147D7AAA04E29C72087F6E" italics="true" pageId="0" pageNumber="309">A. titschacki</emphasis>
</taxonomicName>
and
<taxonomicName id="6E7F6DBE30109FDA8080D9ED1E83F613" lsidName="A. vargasllosai" pageId="0" pageNumber="309" rank="species" species="vargasllosai">
<emphasis id="D0FCEBA5311A6CC8E4B696AFDBFD4259" italics="true" pageId="0" pageNumber="309">A. vargasllosai</emphasis>
</taxonomicName>
<emphasis id="FEFA0C39292F0876DF44E8ED5DD6C790" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
in the following combination of characteristics: dorsal scutum densely granulate; ocularium and areas I-IV of DS unarmed or armed with tiny tubercles, slightly greater than granules; posterior margin of DS and free tergites I-III with median rows of acuminate tubercles (Fig.
<figureCitation id="BAF41E2F6AC3E0C6034BFC689476166C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal of Ayacucho. A A. bambamarca (Roewer, 1957) comb. nov., male; B A. glauberrochai sp. nov., male; C A. inermis (Roewer, 1957) comb. nov., male; D A. insignitus (Roewer, 1956) comb. nov., female; E A. pasolinii sp. nov., male; F A. pomacocha sp. nov., male; G A. querococha sp. nov., male; H A. roeweri nom. nov., female; I A. silvae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure2" httpUri="https://binary.pensoft.net/fig/729876" pageId="0" pageNumber="309">2B</figureCitation>
); femur IV of males without strong armature (except in ventral surface of femur IV in
<taxonomicName id="D0437053A77C9DC77883D2973E574D00" lsidName="A. silvae" pageId="0" pageNumber="309" rank="species" species="silvae">
<emphasis id="4C5FE8705813693A4C002EC2B1C1821F" italics="true" pageId="0" pageNumber="309">A. silvae</emphasis>
</taxonomicName>
<emphasis id="06AA45A5E542AFB28750A7A9F0BE08BF" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, Fig.
<figureCitation id="2247DCEF8F34382A8FC53690A7661449" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7C, D</figureCitation>
). It differs from the four previously mentioned species in the following combination of characteristics: DSS with weakly marked constrictions (Fig.
<figureCitation id="A464BAF2E4B74D82068EBC8F765C271B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal of Ayacucho. A A. bambamarca (Roewer, 1957) comb. nov., male; B A. glauberrochai sp. nov., male; C A. inermis (Roewer, 1957) comb. nov., male; D A. insignitus (Roewer, 1956) comb. nov., female; E A. pasolinii sp. nov., male; F A. pomacocha sp. nov., male; G A. querococha sp. nov., male; H A. roeweri nom. nov., female; I A. silvae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure2" httpUri="https://binary.pensoft.net/fig/729876" pageId="0" pageNumber="309">2B</figureCitation>
); ocularium unarmed and smooth or with a few granules (Fig.
<figureCitation id="A84DC380D0C9C7C7804BC988699BFAD1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal of Ayacucho. A A. bambamarca (Roewer, 1957) comb. nov., male; B A. glauberrochai sp. nov., male; C A. inermis (Roewer, 1957) comb. nov., male; D A. insignitus (Roewer, 1956) comb. nov., female; E A. pasolinii sp. nov., male; F A. pomacocha sp. nov., male; G A. querococha sp. nov., male; H A. roeweri nom. nov., female; I A. silvae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure2" httpUri="https://binary.pensoft.net/fig/729876" pageId="0" pageNumber="309">2B</figureCitation>
); male femur IV unarmed (unlike
<taxonomicName id="B2AB6CE71B7C5774B3A7C189EE13E8EB" lsidName="A. silvae" pageId="0" pageNumber="309" rank="species" species="silvae">
<emphasis id="0A003F051590784E4AE7A01E65F3362F" italics="true" pageId="0" pageNumber="309">A. silvae</emphasis>
</taxonomicName>
<emphasis id="E99DEDCD993298223BAFE6B5421E6749" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, Fig.
<figureCitation id="28F8A4BD2F11F78A7555AD1CA8CAB4F9" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7C, D</figureCitation>
); and penis VP subrectangular with a V-notch on distal margin (Fig.
<figureCitation id="2097B13D575BB8106364E28FAAB777F2" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Penis of Ayacucho, dorsal, lateral and ventral views, respectively. A - C A. spiniger (Roewer, 1957) comb. nov.; D - F A. glauberrochai sp. nov.; G - I A. tapacocha nom. nov.; Legend bars = 0.1 mm" figureDoi="10.3897/asp.80.e73829.figure16" httpUri="https://binary.pensoft.net/fig/729890" pageId="0" pageNumber="309">16D, F</figureCitation>
).
</paragraph>
<caption id="DCC80189674D4A77FE54C402AEC57B47" doi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309" start="Figure 6" startId="F6">
<paragraph id="D7CF2F9193E1399FA9E420580782CC00" pageId="0" pageNumber="309">
<emphasis id="86418D255585A6F364A74C08F44D97D8" bold="true" pageId="0" pageNumber="309">Figure 6.</emphasis>
Habitus, dorsal of males of
<taxonomicName id="DDF7CE2C6A2D45134ACFC5604F980C23" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Lumieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lumieria" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="4ECFAB7B3D9A902C5E3A93611E5E1627" italics="true" pageId="0" pageNumber="309">Lumieria</emphasis>
</taxonomicName>
<emphasis id="4C3EB510D0E314404ED3DB6171F5A7C5" bold="true" pageId="0" pageNumber="309">gen. nov.</emphasis>
and
<taxonomicName id="7F640629B18D6ABAF2EEE266470261B1" authorityName="Roewer" authorityYear="1957" class="Arachnida" family="Metasarcidae" genus="Tschaidicancha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tschaidicancha" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="F765D10171855BC82F93265CF4345D93" italics="true" pageId="0" pageNumber="309">Tschaidicancha</emphasis>
</taxonomicName>
.
<emphasis id="3969A4C5C462F46B622ED4C533069C53" bold="true" pageId="0" pageNumber="309">A</emphasis>
<taxonomicName id="750735EEC5D837F72D19916DD84110A8" lsidName="L. antonionii" pageId="0" pageNumber="309" rank="species" species="antonionii">
<emphasis id="644ADE53ED9906151AE6DDE31DEE8D40" italics="true" pageId="0" pageNumber="309">L. antonionii</emphasis>
</taxonomicName>
<emphasis id="5FD7BB07E29A58D0CA76380A378DAD5C" bold="true" pageId="0" pageNumber="309">gen. et sp. nov.</emphasis>
;
<emphasis id="BE170DB3126F5909A03AAE2AB30C4B02" bold="true" pageId="0" pageNumber="309">B</emphasis>
<taxonomicName id="2E79A4BAD151D696BCAB8EEFF2A6965E" lsidName="L. woodyalleni" pageId="0" pageNumber="309" rank="species" species="woodyalleni">
<emphasis id="401608737D811316AF2F26A73E2DDD47" italics="true" pageId="0" pageNumber="309">L. woodyalleni</emphasis>
</taxonomicName>
<emphasis id="719C33145F1497A95817B633F4273268" bold="true" pageId="0" pageNumber="309">gen. et sp. nov.</emphasis>
;
<emphasis id="A5F3DDCBACE95A090D8C63577ED5EE47" bold="true" pageId="0" pageNumber="309">C</emphasis>
<taxonomicName id="BF56B1867F8EB83FDFCD454991E83EB9" class="Arachnida" family="Metasarcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. chaplini" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="chaplini">
<emphasis id="3F714939A63C800528368A90662F16C9" italics="true" pageId="0" pageNumber="309">T. chaplini</emphasis>
</taxonomicName>
<emphasis id="0D60461F25EFB565C8B67A7FD0832BF1" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
;
<emphasis id="23BC162B11EC821E117F6ED63A8BF7FE" bold="true" pageId="0" pageNumber="309">D</emphasis>
<taxonomicName id="9C402B96BA50F5467F7F5E6B18F80D7C" class="Arachnida" family="Metasarcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. joseochoai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="joseochoai">
<emphasis id="120BC83F1A67711E3AAA6B6D12D173CE" italics="true" pageId="0" pageNumber="309">T. joseochoai</emphasis>
</taxonomicName>
<emphasis id="FEC12143010E2E26D48990B740C57E54" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
;
<emphasis id="9EDF60FE2E1FC64E96997C2E6DC1FFF7" bold="true" pageId="0" pageNumber="309">E</emphasis>
<taxonomicName id="1A4CF4301CF79FD95660CFE58C393435" class="Arachnida" family="Metasarcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. scorsesei" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="scorsesei">
<emphasis id="DE4494351BCE438CE6164C606A8D414E" italics="true" pageId="0" pageNumber="309">T. scorsesei</emphasis>
</taxonomicName>
<emphasis id="060774E0A0509D95407081FD11E4E89A" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
;
<emphasis id="88918164840D6E6FA761B18861654825" bold="true" pageId="0" pageNumber="309">F</emphasis>
<taxonomicName id="6FD8F58963FB886A3902904B9D4FC519" class="Arachnida" family="Metasarcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. weyrauchi" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="weyrauchi">
<emphasis id="FB40C6EC7AF4D58288942C07887E665B" italics="true" pageId="0" pageNumber="309">T. weyrauchi</emphasis>
</taxonomicName>
Roewer, 1957; Legend bars = 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="A7AA8CBF175714CAE6A8CC85DE709EF0" pageId="0" pageNumber="309" type="etymology">
<paragraph id="A90CA978A8D5CA6E52B59DB5D45195D5" pageId="0" pageNumber="309">Derivatio nominis.</paragraph>
<paragraph id="75A8B00AC26CEF8F347E2B885C7A5CCD" pageId="0" pageNumber="309">The specific epithet of masculine gender, in the genitive form, dedicated to the Brazilian filmmaker, actor and writer Glauber de Andrade Rocha (1939-1981).</paragraph>
</subSubSection>
<subSubSection id="A28CD0E32EAC87DA6AB7D2115A8A03E3" pageId="0" pageNumber="309" type="distribution">
<paragraph id="49D52F154506875A0230677F1B499B88" pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph id="547E875FCE6280DA8CCFC467286D6B6F" pageId="0" pageNumber="309">
(Fig.
<figureCitation id="309390D9011483820B6CEA40325D68AF" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">28</figureCitation>
) PERU. Huancavellica. Huancavellica, Quebrada Potreros.
</paragraph>
</subSubSection>
<subSubSection id="0413E982CD249F4DE6842D92AD67FAE7" pageId="0" pageNumber="309" type="materials_examined">
<paragraph id="5E29995A250B1A3D370D45832742966D" pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph id="419136C28A59FA71CDD0E6B768E1AFBE" pageId="0" pageNumber="309">
<emphasis id="243A7F44E8E82B77FD0F9BB72FDC12C3" bold="true" pageId="0" pageNumber="309">
<emphasis id="2994C812B6822F8B257AE143AA07B696" italics="true" pageId="0" pageNumber="309">Type material</emphasis>
:
<materialsCitation id="657F561F9A359B522E9C21602AAA66E4" ID-GBIF-Occurrence="3889641358" collectingDate="2011-04-28" collectorName="R. Pinto-da-Rocha, A. Benedetti, J. Ochoa, D. Silva" country="PERU" county="Quebrada Potreros" latitude="-12.769639" location="Huancavellica" longLatPrecision="1" longitude="-75.017365" municipality="Huancavellica" specimenCount="♂" typeStatus="Holotype">
<typeStatus id="05DD20DD9F3FF4147F660E050B82BFF3">Holotype</typeStatus>
<specimenCount id="125D95EFD7043DE886412B7E064CCAEA">♂</specimenCount>
, '
<collectingCountry id="C77E86E7A1AD44005DCA5367294D0981" name="Peru">PERU</collectingCountry>
,
<location id="1C807DB039CD93F21B6D0BBF0F336617" LSID="urn:lsid:plazi:treatment:600BA0C567F5516BBC1AAB18D4488B84:1C807DB039CD93F21B6D0BBF0F336617" country="PERU" county="Quebrada Potreros" latitude="-12.769639" longLatPrecision="1" longitude="-75.017365" municipality="Huancavellica" name="Huancavellica">Huancavellica</location>
,
<collectingMunicipality id="92F9DA34A82F9C1C46756F8875D60F52">Huancavellica</collectingMunicipality>
,
<collectingCounty id="196C1F984599A670620055F363D2F5C5">Quebrada Potreros</collectingCounty>
|
<geoCoordinate id="5099DDC7C321E109212CFC8644F950FB" degrees="12" direction="south" minutes="46" orientation="latitude" precision="1" seconds="10.7" value="-12.769639">12°46′10.7″S</geoCoordinate>
<geoCoordinate id="D486E3C0ABEC251D988A54B33001AB95" degrees="75" direction="west" minutes="01" orientation="longitude" precision="1" seconds="02.5" value="-75.017365">75°01′02.5″W</geoCoordinate>
|
<collectingDate id="56938467C261F8E65EF2E453284C626E" value="2011-04-28">28/IV/2011</collectingDate>
,
<collectorName id="F535BCFD85668DB268B0B3F965F89FC7">R. Pinto-da-Rocha</collectorName>
,
<collectorName id="47245EB3F395EC903ACF5F1DB6218194">A. Benedetti</collectorName>
,
<collectorName id="EDBC7383659CF6571EC1A691C7B7C221">J. Ochoa</collectorName>
&amp;
<collectorName id="4AB95456731B184CC16D6409592C3889">D. Silva</collectorName>
leg. (MUBI)
</materialsCitation>
</emphasis>
-
<emphasis id="B39D2B4C499376F4D00A4BDC12476A0B" bold="true" pageId="0" pageNumber="309">
<typeStatus id="C18D915D9FF8F3A7BFEE28725B6BFB32">Paratypes</typeStatus>
</emphasis>
<specimenCount id="F006B81044FF57AAC4AA64C810C8E825" type="male">1 ♂</specimenCount>
,
<specimenCount id="55B4DA6C03AE92C9F692EBB75A298196" type="female">5 ♀</specimenCount>
,
<normalizedToken id="E8959CEFC9B6556BCFACFE3290600018" originalValue="‘ditto’">'ditto'</normalizedToken>
(MUBI);
<emphasis id="209EF7CB05E76AEFFEED17E0D9C389AE" bold="true" pageId="0" pageNumber="309">
<typeStatus id="025CC63066C3B409423F14762097433F">Paratypes</typeStatus>
</emphasis>
<specimenCount id="00122AD3D89E9611C230B447D4C19B17" type="male">5 ♂</specimenCount>
,
<specimenCount id="16ECE0BFF368C461B2FAB12E48CEDD38" type="female">11 ♀</specimenCount>
,
<normalizedToken id="0C5B6CCF2085D6279CC3342511C77AD0" originalValue="‘ditto’">'ditto'</normalizedToken>
(MZSP 36974);
<emphasis id="0AFD1C4DFDBD568CFE4C2D6CEF80E12D" bold="true" pageId="0" pageNumber="309">
<typeStatus id="3718E10C923C9EF55042F18C518F6B91">Paratypes</typeStatus>
</emphasis>
<specimenCount id="E82F6744D88BE66F03DFE95CA34E7882" type="male">2 ♂</specimenCount>
,
<specimenCount id="EA64F545A5C68C7DFDBAACBB7DA7BA9A" type="female">5 ♀</specimenCount>
,
<normalizedToken id="E2FA4B05B5A47F89112944EDB0734716" originalValue="‘ditto’">'ditto'</normalizedToken>
(MUSM).
</paragraph>
</subSubSection>
</treatment>
</document>
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