treatments-xml/data/54/41/A0/5441A02CB133FFA83EACB5A1FACF535C.xml
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<document id="7B5AACCF1BF1C6625FC56D0E09591A25" ID-DOI="10.11646/zootaxa.4999.5.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5119429" ID-ZooBank="C5BC0813-D7ED-4192-A726-7560C1BC28DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626875728399" checkinUser="plazi" docAuthor="Opresko, Dennis M. &amp; Molodtsova, Tina N." docDate="2021" docId="5441A02CB133FFA83EACB5A1FACF535C" docLanguage="en" docName="zootaxa.4999.5.1.pdf" docOrigin="Zootaxa 4999 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alternatipathes mirabilis Opresko &amp; Molodtsova 2021, n. sp." docType="treatment" docUuid="C810F528-71BD-4604-A08F-5180236EDCB8" docUuidSource="ZooBank" docVersion="3" lastPageNumber="405" masterDocId="A878D854B131FFAC3E3BB064FF8C5651" masterDocTitle="New species of deep-sea Antipatharians from the North Pacific (Cnidaria: Anthozoa: Antipatharia), Part 2" masterLastPageNumber="422" masterPageNumber="401" pageNumber="403" updateTime="1699012000698" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E0220FE88DE73EAE35FF45DB9642474A">New species of deep-sea Antipatharians from the North Pacific (Cnidaria: Anthozoa: Antipatharia), Part 2</mods:title>
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<mods:name id="AF5479787BE6178842E415B7A8CAFE65" type="personal">
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<mods:roleTerm id="2C7880270768C754A13F87C23042DE40">Author</mods:roleTerm>
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<mods:namePart id="F0AE1BACAEDDF53575310D90E2AA2890">Opresko, Dennis M.</mods:namePart>
<mods:nameIdentifier id="E2B98CAA4FCAA7E35D0C1F4AB141264C" type="ORCID">0000-0001-9946-1533</mods:nameIdentifier>
<mods:affiliation id="E383D6952FF8AFC3E9C2F6A91BF64703">Department of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, USA. dmopresko @ hotmail. com; https: // orcid. org / 0000 - 0001 - 9946 - 1533</mods:affiliation>
<mods:nameIdentifier id="858BF29628AC97A451A41B06176B1E12" type="email">dmopresko@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="D3DBCC2BAE424CF7024D6A26DF341BC8">Molodtsova, Tina N.</mods:namePart>
<mods:affiliation id="3B0A8B85C9CC1B8AAF6E77BB5A15BFAE">P. P. Shirshov Institute of Oceanology, RAS, 36 Nakhimovsky prospect, Moscow 117997, Russia</mods:affiliation>
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<mods:title id="D4D97FC7589C28D070FA8E6956A7C2CA">Zootaxa</mods:title>
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<mods:date id="CE1CD02175476A347C2087BF3E18C096">2021</mods:date>
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<mods:number id="5824903765441D42D5047988534CD121">4999</mods:number>
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<treatment id="5441A02CB133FFA83EACB5A1FACF535C" LSID="urn:lsid:zoobank.org:act:C810F528-71BD-4604-A08F-5180236EDCB8" httpUri="http://treatment.plazi.org/id/5441A02CB133FFA83EACB5A1FACF535C" lastPageId="4" lastPageNumber="405" pageId="2" pageNumber="403">
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<paragraph id="DC57113AB133FFAE3EACB5A1FD82538E" blockId="2.[151,961,1477,1576]" box="[151,526,1477,1504]" pageId="2" pageNumber="403">
<heading id="871FA656B133FFAE3EACB5A1FD82538E" bold="true" box="[151,526,1477,1504]" fontSize="11" level="1" pageId="2" pageNumber="403" reason="1">
<emphasis id="EE9CCD28B133FFAE3EACB5A1FD82538E" bold="true" box="[151,526,1477,1504]" pageId="2" pageNumber="403">
<taxonomicName id="1BE86AB9B133FFAE3EACB5A1FE4853B1" ID-CoL="8M2TJ" authority="Opresko &amp; Molodtsova, 2021" authorityName="Opresko &amp; Molodtsova" authorityYear="2021" box="[151,452,1477,1504]" class="Anthozoa" family="Schizopathidae" genus="Alternatipathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="2" pageNumber="403" phylum="Cnidaria" rank="species" species="mirabilis" status="sp. nov.">
<emphasis id="EE9CCD28B133FFAE3EACB5A1FE4853B1" bold="true" box="[151,452,1477,1504]" italics="true" pageId="2" pageNumber="403">Alternatipathes mirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F5AF7053B133FFAE3FF0B5A1FD82538E" box="[459,526,1477,1503]" pageId="2" pageNumber="403" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="94F242B1B133FFAE3EACB58DFC4D5079" pageId="2" pageNumber="403" type="description">
<paragraph id="DC57113AB133FFAE3EACB58DFF675052" blockId="2.[151,961,1477,1576]" box="[151,235,1513,1539]" pageId="2" pageNumber="403">
(
<figureCitation id="44D30DBFB133FFAE3E9BB58DFF6C5052" box="[160,224,1513,1539]" captionStart="FIGURE 1" captionStartId="3.[151,250,1980,2004]" captionTargetBox="[151,1436,489,1957]" captionTargetId="figure-202@3.[151,1436,489,1957]" captionTargetPageId="3" captionText="FIGURE 1. Alternatipathes mirabilis n. sp., holotype, USNM 1070972: A. corallum; B. enlarged view of spines; C. polyps; D. sections of pinnules showing spines, scale bars 0.2 mm (B and D from SEM stub 488)." figureDoi="http://doi.org/10.5281/zenodo.5119431" httpUri="https://zenodo.org/record/5119431/files/figure.png" pageId="2" pageNumber="403">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="DC57113AB133FFAE3EACB669FC4D5079" blockId="2.[151,961,1477,1576]" box="[151,961,1549,1576]" pageId="2" pageNumber="403">
<uri id="A8791D38B133FFAE3EACB669FC4D5079" box="[151,961,1549,1576]" pageId="2" pageNumber="403">
urn:lsid:zoobank.org:act:
<uuid id="A84E2BEFB133FFAE3F9CB669FC4D5079" box="[423,961,1549,1576]" pageId="2" pageNumber="403">C810F528-71BD-4604-A08F-5180236EDCB8</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="94F242B1B133FFAE3EACB631FE0950E6" pageId="2" pageNumber="403" type="materials_examined">
<paragraph id="DC57113AB133FFAE3EACB631FE0950E6" blockId="2.[151,1437,1621,2008]" pageId="2" pageNumber="403">
<emphasis id="EE9CCD28B133FFAE3EACB631FE0F503E" bold="true" box="[151,387,1621,1647]" pageId="2" pageNumber="403">Material examined.</emphasis>
<materialsCitation id="6C801B67B133FFAE3FB0B631FE0950E6" collectingDate="2004-07-20" collectionCode="SEM, ROV" elevation="4685" latitude="53.0419" location="North Pacific" longLatPrecision="6" longitude="-161.183" municipality="Derickson Seamount" pageId="2" pageNumber="403" specimenCount="1" typeStatus="holotype">
<emphasis id="EE9CCD28B133FFAE3FB0B631FE71503E" bold="true" box="[395,509,1621,1648]" pageId="2" pageNumber="403">
<emphasis id="EE9CCD28B133FFAE3FB0B631FE7F5021" bold="true" box="[395,499,1621,1648]" italics="true" pageId="2" pageNumber="403">
<typeStatus id="0353AF98B133FFAE3FB0B631FE7F5021" box="[395,499,1621,1648]" pageId="2" pageNumber="403" type="holotype">Holotype</typeStatus>
</emphasis>
:
</emphasis>
USNM 1070972 (
<collectionCode id="BAF989FFB133FFAE3CE8B631FC82503E" box="[723,782,1621,1647]" pageId="2" pageNumber="403">SEM</collectionCode>
stubs 451 and 488),
<location id="D93747E1B133FFAE3A3AB631FB10503E" LSID="urn:lsid:plazi:treatment:5441A02CB133FFA83EACB5A1FACF535C:D93747E1B133FFAE3A3AB631FB10503E" box="[1025,1180,1621,1647]" latitude="53.0419" longLatPrecision="6" longitude="-161.183" municipality="Derickson Seamount" name="North Pacific" pageId="2" pageNumber="403">North Pacific</location>
,
<collectingMunicipality id="3C338B40B133FFAE3A92B631FA14503E" box="[1193,1432,1621,1647]" pageId="2" pageNumber="403">Derickson Seamount</collectingMunicipality>
,
<geoCoordinate id="B9DC77FDB133FFAE3EACB61DFE9850C5" box="[151,276,1657,1684]" degrees="53.0419" direction="north" orientation="latitude" pageId="2" pageNumber="403" precision="5" value="53.0419">53.0419°N</geoCoordinate>
,
<geoCoordinate id="B9DC77FDB133FFAE3F19B61DFE2250C5" box="[290,430,1657,1684]" degrees="161.1830" direction="west" orientation="longitude" pageId="2" pageNumber="403" precision="5" value="-161.183">161.1830°W</geoCoordinate>
,
<collectionCode id="BAF989FFB133FFAE3F85B61DFE7550C2" box="[446,505,1657,1683]" pageId="2" pageNumber="403">ROV</collectionCode>
<taxonomicName id="1BE86AB9B133FFAE3C3AB61EFD2850C2" authority="II, Dive" authorityName="II, Dive" box="[513,676,1657,1684]" class="Gastropoda" family="Facelinidae" genus="Jason" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="403" phylum="Mollusca" rank="genus">
<emphasis id="EE9CCD28B133FFAE3C3AB61EFDCE50C2" box="[513,578,1658,1683]" italics="true" pageId="2" pageNumber="403">Jason</emphasis>
II, Dive
</taxonomicName>
93 (Field Identification Number: JD-093),
<quantity id="1B10BCDFB133FFAE3AADB61DFB6350C5" box="[1174,1263,1657,1684]" metricMagnitude="3" metricUnit="m" metricValue="4.685" pageId="2" pageNumber="403" unit="m" value="4685.0">
<elevation id="57C5F609B133FFAE3AADB61DFB6350C5" box="[1174,1263,1657,1684]" metricMagnitude="3" metricUnit="m" metricValue="4.685" pageId="2" pageNumber="403" unit="m" value="4685.0">4685 m</elevation>
</quantity>
, coll. A. Baco- Taylor,
<date id="A85637FAB133FFAE3ED6B6F9FE0D50E6" box="[237,385,1693,1719]" pageId="2" pageNumber="403" value="2004-07-20">
<collectingDate id="B812CE12B133FFAE3ED6B6F9FE0D50E6" box="[237,385,1693,1719]" pageId="2" pageNumber="403" value="2004-07-20">20 July 2004</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="94F242B1B133FFAE3EFCB6A5FD9C51C1" pageId="2" pageNumber="403" type="diagnosis">
<paragraph id="DC57113AB133FFAE3EFCB6A5FD9C51C1" blockId="2.[151,1437,1621,2008]" pageId="2" pageNumber="403">
<emphasis id="EE9CCD28B133FFAE3EFCB6A5FEB6508A" bold="true" box="[199,314,1729,1755]" pageId="2" pageNumber="403">Diagnosis</emphasis>
. Colony attached, monopodial, unbranched, and pinnulate. Pinnules simple, arranged alternately in two lateral rows along upper part of stem. Lower unpinnulated section of the stem longer than upper pinnulated section. Pinnules generally decreasing in length proximally to distally. Pinnular density 1011 per
<quantity id="1B10BCDFB133FFAE3AD1B76DFAAD5175" box="[1258,1313,1801,1828]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="403" unit="cm" value="3.0">3 cm</quantity>
(including pinnules in both rows). Spines on pinnules smooth, triangular in profile, moderately acute, laterally compressed, and mostly
<quantity id="1B10BCDFB133FFAE3F27B735FE4C513D" box="[284,448,1873,1900]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="2" pageNumber="403" unit="mm" value="0.05" valueMax="0.06" valueMin="0.04">0.040.06 mm</quantity>
tall on polypar side of axis. Polyps on pinnules mostly
<quantity id="1B10BCDFB133FFAE3A03B735FB1A513D" box="[1080,1174,1873,1900]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="403" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
in transverse diameter, with four to five polyps per
<quantity id="1B10BCDFB133FFAE3FEAB711FD8051C1" box="[465,524,1909,1936]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="403" unit="cm" value="3.0">3 cm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="94F242B1B133FFA83EFCB7FDFCCF5294" lastPageId="4" lastPageNumber="405" pageId="2" pageNumber="403" type="description">
<paragraph id="DC57113AB133FFAF3EFCB7FDFDE25780" blockId="2.[151,1437,1621,2008]" lastBlockId="3.[151,1437,150,465]" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403">
<emphasis id="EE9CCD28B133FFAE3EFCB7FDFD8251E2" bold="true" box="[199,526,1945,1971]" pageId="2" pageNumber="403">
Description of the
<typeStatus id="0353AF98B133FFAE3F9CB7FDFD8251E2" box="[423,526,1945,1971]" pageId="2" pageNumber="403" type="holotype">holotype</typeStatus>
</emphasis>
. The
<typeStatus id="0353AF98B133FFAE3C74B7FDFD3D51E2" box="[591,689,1945,1971]" pageId="2" pageNumber="403" type="holotype">holotype</typeStatus>
(USNM 1070972,
<figureCitation id="44D30DBFB133FFAE3DB6B7FDFC6F51E2" box="[909,995,1945,1971]" captionStart="FIGURE 1" captionStartId="3.[151,250,1980,2004]" captionTargetBox="[151,1436,489,1957]" captionTargetId="figure-202@3.[151,1436,489,1957]" captionTargetPageId="3" captionText="FIGURE 1. Alternatipathes mirabilis n. sp., holotype, USNM 1070972: A. corallum; B. enlarged view of spines; C. polyps; D. sections of pinnules showing spines, scale bars 0.2 mm (B and D from SEM stub 488)." figureDoi="http://doi.org/10.5281/zenodo.5119431" httpUri="https://zenodo.org/record/5119431/files/figure.png" pageId="2" pageNumber="403">Fig. 1A</figureCitation>
) is an almost complete colony with an intact basal holdfast; the very tip of the stem, however, is broken off. The stem is about
<quantity id="1B10BCDFB133FFAE3A42B7D9FB585189" box="[1145,1236,1981,2008]" metricMagnitude="-1" metricUnit="m" metricValue="4.65" pageId="2" pageNumber="403" unit="cm" value="46.5">46.5 cm</quantity>
long with a basal diameter of
<quantity id="1B10BCDFB132FFAF3F24B0F3FEF556E0" box="[287,377,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="404" unit="mm" value="2.4">2.4 mm</quantity>
. The lower unpinnulated section of the stem is
<quantity id="1B10BCDFB132FFAF3DB7B0F2FC6656E0" box="[908,1002,150,177]" metricMagnitude="-1" metricUnit="m" metricValue="3.25" pageId="3" pageNumber="404" unit="cm" value="32.5">32.5 cm</quantity>
, and the upper pinnulated section covers a distance of
<quantity id="1B10BCDFB132FFAF3F6DB0DFFE2C5684" box="[342,416,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="404" unit="cm" value="14.0">14 cm</quantity>
. The pinnules are simple, bilateral and alternately arranged; generally decreasing in length from the lower part of pinnulated stalk to the distal end. Most of the pinnules are broken off at the end; the longest remaining ones are about
<quantity id="1B10BCDFB132FFAF3F87B166FD8E574C" box="[444,514,258,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="404" unit="cm" value="15.0">15 cm</quantity>
in length with a basal diameter of about
<quantity id="1B10BCDFB132FFAF3DEEB167FBA3574C" box="[981,1071,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="404" unit="mm" value="0.8">0.8 mm</quantity>
. Forty-six pinnules occur on the corallum, 23 on each side of the stem. Within each row, the pinnules are spaced
<quantity id="1B10BCDFB132FFAF3A1DB142FB085710" box="[1062,1156,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="404" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
apart. The resulting pinnular density is 10 (total for both rows) per
<quantity id="1B10BCDFB132FFAF3CBFB12FFD375734" box="[644,699,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="404" unit="cm" value="3.0">3 cm</quantity>
near the basal end of the stem to about 11 per
<quantity id="1B10BCDFB132FFAF3AF2B12FFA8C5734" box="[1225,1280,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="404" unit="cm" value="3.0">3 cm</quantity>
in the middle and towards the distal end. In the preserved specimen the lower rows of pinnules form an interior angle of about 60°, which increases towards the top of the corallum to 180°. The distal angle that the pinnules form with the stem is 6080°, even at the top of the corallum.
</paragraph>
<caption id="889741B2B132FFAF3EACB7D8FBAA51A9" ID-DOI="http://doi.org/10.5281/zenodo.5119431" ID-Zenodo-Dep="5119431" httpUri="https://zenodo.org/record/5119431/files/figure.png" pageId="3" pageNumber="404" startId="3.[151,250,1980,2004]" targetBox="[151,1436,489,1957]" targetPageId="3">
<paragraph id="DC57113AB132FFAF3EACB7D8FBAA51A9" blockId="3.[151,1436,1980,2040]" pageId="3" pageNumber="404">
<emphasis id="EE9CCD28B132FFAF3EACB7D8FE985185" bold="true" box="[151,276,1980,2004]" pageId="3" pageNumber="404">FIGURE 1.</emphasis>
<taxonomicName id="1BE86AB9B132FFAF3F27B7D8FD975185" authority="Opresko &amp; Molodtsova, 2021" authorityName="Opresko &amp; Molodtsova" authorityYear="2021" box="[284,539,1980,2004]" class="Anthozoa" family="Schizopathidae" genus="Alternatipathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="3" pageNumber="404" phylum="Cnidaria" rank="species" species="mirabilis" status="sp. nov.">
<emphasis id="EE9CCD28B132FFAF3F27B7D8FD975185" box="[284,539,1980,2004]" italics="true" pageId="3" pageNumber="404">Alternatipathes mirabilis</emphasis>
</taxonomicName>
<emphasis id="EE9CCD28B132FFAF3C19B7D8FDD15185" bold="true" box="[546,605,1980,2004]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="F5AF7053B132FFAF3C19B7D8FDD15185" box="[546,605,1980,2004]" pageId="3" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, USNM 1070972:
<emphasis id="EE9CCD28B132FFAF3DB2B7D8FC175185" bold="true" box="[905,923,1980,2004]" pageId="3" pageNumber="404">A</emphasis>
. corallum;
<emphasis id="EE9CCD28B132FFAF3A2AB7D8FBAE5185" bold="true" box="[1041,1058,1980,2004]" pageId="3" pageNumber="404">B</emphasis>
. enlarged view of spines;
<emphasis id="EE9CCD28B132FFAF3B09B7D8FAC85185" bold="true" box="[1330,1348,1980,2004]" pageId="3" pageNumber="404">C</emphasis>
. polyps;
<emphasis id="EE9CCD28B132FFAF3EACB784FF2551A9" bold="true" box="[151,169,2016,2040]" pageId="3" pageNumber="404">D</emphasis>
. sections of pinnules showing spines, scale bars 0.2 mm (B and D from SEM stub 488).
</paragraph>
</caption>
<paragraph id="DC57113AB135FFA83EFCB0F3FDDD57FC" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="405">
The spines on the pinnules are very small, triangular in profile, moderately acute, smooth, and laterally compressed (
<figureCitation id="44D30DBFB135FFA83EC1B0DFFE0E5684" box="[250,386,187,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1980,2004]" captionTargetBox="[151,1436,489,1957]" captionTargetId="figure-202@3.[151,1436,489,1957]" captionTargetPageId="3" captionText="FIGURE 1. Alternatipathes mirabilis n. sp., holotype, USNM 1070972: A. corallum; B. enlarged view of spines; C. polyps; D. sections of pinnules showing spines, scale bars 0.2 mm (B and D from SEM stub 488)." figureDoi="http://doi.org/10.5281/zenodo.5119431" httpUri="https://zenodo.org/record/5119431/files/figure.png" pageId="4" pageNumber="405">Fig. 1B, 1D</figureCitation>
). There is only a slight difference in the size of the polypar and abpolypar spines. On sections of pinnules about
<quantity id="1B10BCDFB135FFA83F59B0BBFE3556A8" box="[354,441,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="405" unit="mm" value="0.4">0.4 mm</quantity>
in diameter the polypar spines are mostly
<quantity id="1B10BCDFB135FFA83DA3B0BBFBC656A8" box="[920,1098,223,249]" metricMagnitude="-5" metricUnit="m" metricValue="4.35" metricValueMax="4.699999999999999" metricValueMin="4.0" pageId="4" pageNumber="405" unit="mm" value="0.0435" valueMax="0.047" valueMin="0.04">0.040.047 mm</quantity>
tall (maximum
<quantity id="1B10BCDFB135FFA83AC4B0BBFAE856A8" box="[1279,1380,223,249]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="405" unit="mm" value="0.06">0.06 mm</quantity>
) and the abpolypar spines are
<quantity id="1B10BCDFB135FFA83F97B167FDC2574C" box="[428,590,259,285]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="405" unit="mm" value="0.035" valueMax="0.04" valueMin="0.03">0.030.04 mm</quantity>
tall. Double spines are present, but are not common. The central axial canal is
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in diameter on a pinnule
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in diameter. Five or six axial rows are visible in lateral view, and a few individual spines occur randomly between the rows. In each row the spines are a variable distance apart,
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; however, on average, there are 2.53.5 spines per mm in each axial row. The spines on the stem are similar to those on the pinnules in size and shape.
</paragraph>
<paragraph id="DC57113AB135FFA83EFCB1D3FDE35448" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="405">
The polyps (
<figureCitation id="44D30DBFB135FFA83F6FB1D3FE255780" box="[340,425,439,465]" captionStart="FIGURE 1" captionStartId="3.[151,250,1980,2004]" captionTargetBox="[151,1436,489,1957]" captionTargetId="figure-202@3.[151,1436,489,1957]" captionTargetPageId="3" captionText="FIGURE 1. Alternatipathes mirabilis n. sp., holotype, USNM 1070972: A. corallum; B. enlarged view of spines; C. polyps; D. sections of pinnules showing spines, scale bars 0.2 mm (B and D from SEM stub 488)." figureDoi="http://doi.org/10.5281/zenodo.5119431" httpUri="https://zenodo.org/record/5119431/files/figure.png" pageId="4" pageNumber="405">Fig. 1C</figureCitation>
) occur in a single series on one side of the pinnules; they are
<quantity id="1B10BCDFB135FFA83A5AB1D2FB325780" box="[1121,1214,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="4" pageNumber="405" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
in transverse diameter. Near the distal end of the pinnules there are four polyps per
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, and in the mid to basal sections of the pinnules there are up to five polyps per
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.
</paragraph>
<paragraph id="DC57113AB135FFA83EFCB246FBCA546C" blockId="4.[151,1437,151,1293]" box="[199,1094,546,573]" pageId="4" pageNumber="405">
<emphasis id="EE9CCD28B135FFA83EFCB246FEF6546D" bold="true" box="[199,378,546,572]" pageId="4" pageNumber="405">Other material</emphasis>
. No other specimens can be assigned to the species at this time.
</paragraph>
<paragraph id="DC57113AB135FFA83EFCB222FC0554D4" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="405">
<emphasis id="EE9CCD28B135FFA83EFCB222FEED5430" bold="true" box="[199,353,582,609]" pageId="4" pageNumber="405">Genetic data</emphasis>
.
<materialsCitation id="6C801B67B135FFA83F4AB223FC0954D4" collectionCode="USNM" pageId="4" pageNumber="405" specimenCode="USNM 1070972" specimenCount="1" typeStatus="holotype">
DNA analysis (
<emphasis id="EE9CCD28B135FFA83C1AB223FD175430" box="[545,667,583,609]" italics="true" pageId="4" pageNumber="405">nad5-nad1</emphasis>
) of the
<typeStatus id="0353AF98B135FFA83CC3B223FCD65430" box="[760,858,583,609]" pageId="4" pageNumber="405" type="holotype">holotype</typeStatus>
(
<specimenCode id="8C4EB941B135FFA83D51B223FBA65430" box="[874,1066,583,609]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="405" type="Museum">USNM 1070972</specimenCode>
) suggests a close relationship to
<taxonomicName id="1BE86AB9B135FFA83EACB20FFD8554D4" authority="(Chery et al. 2018)" baseAuthorityName="Chery" baseAuthorityYear="2018" box="[151,521,619,645]" class="Anthozoa" family="Schizopathidae" genus="Schizopathes" kingdom="Animalia" order="Antipatharia" pageId="4" pageNumber="405" phylum="Cnidaria" rank="genus">
<emphasis id="EE9CCD28B135FFA83EACB20FFEA554D4" box="[151,297,619,645]" italics="true" pageId="4" pageNumber="405">Schizopathes</emphasis>
(
<bibRefCitation id="B8796CCBB135FFA83F03B20FFD8D54D4" author="Chery, N. &amp; Parra, K. &amp; Evankow, A. &amp; Stein, D. &amp; Distel, D. &amp; Appiah-Madson, H. &amp; Ross, R. &amp; Sanon, E. &amp; Alomari, N. &amp; Johnson, R. &amp; Vasovic, A. &amp; Horowitz, A. &amp; Popa, H. &amp; Short, B. &amp; Kourehjan, D. &amp; Vasquez, D. M. &amp; Rodriguez, E. &amp; Opresko, D. M. &amp; Brugler, M. R." box="[312,513,619,645]" pageId="4" pageNumber="405" refId="ref14506" refString="Chery, N., Parra, K., Evankow, A., Stein, D., Distel, D., Appiah-Madson, H., Ross, R., Sanon, E., Alomari, N., Johnson, R., Vasovic, A., Horowitz, A., Popa, H., Short, B., Kourehjan, D., Vasquez, D. M., Rodriguez, E., Opresko, D. M. &amp; Brugler, M. R. (2018) Partnering with the Ocean Genome Legacy to advance our understanding of black corals (Order Antipatharia). 15 th Deep-Sea Biology Symposium, Monterey, California, 9 - 14 September 2018. poster presentation. [preprint available from M. Brugler at https: // tidalmarshtaskforce. wixsite. com / uscb / publications]" type="url" year="2018">
Chery
<emphasis id="EE9CCD28B135FFA83FBEB20FFE3154D4" box="[389,445,619,645]" italics="true" pageId="4" pageNumber="405">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
; however, further study is needed
</materialsCitation>
.
</paragraph>
<paragraph id="DC57113AB135FFA83EFCB2EAFA0D54A0" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="405">
<emphasis id="EE9CCD28B135FFA83EFCB2EAFEEA54F9" bold="true" box="[199,358,654,680]" pageId="4" pageNumber="405">Comparisons</emphasis>
.
<taxonomicName id="1BE86AB9B135FFA83F48B2EBFD0754F8" authority="Opresko &amp; Molodtsova, 2021" authorityName="Opresko &amp; Molodtsova" authorityYear="2021" box="[371,651,655,681]" class="Anthozoa" family="Schizopathidae" genus="Alternatipathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="4" pageNumber="405" phylum="Cnidaria" rank="species" species="mirabilis" status="sp. nov.">
<emphasis id="EE9CCD28B135FFA83F48B2EBFD0754F8" box="[371,651,655,681]" italics="true" pageId="4" pageNumber="405">Alternatipathes mirabilis</emphasis>
</taxonomicName>
<emphasis id="EE9CCD28B135FFA83CAAB2EBFD4354F8" bold="true" box="[657,719,655,681]" pageId="4" pageNumber="405">
<taxonomicNameLabel id="F5AF7053B135FFA83CAAB2EBFD4354F8" box="[657,719,655,681]" pageId="4" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from the
<typeStatus id="0353AF98B135FFA83DB2B2EBFC3554F8" box="[905,953,655,681]" pageId="4" pageNumber="405">type</typeStatus>
species of the genus,
<emphasis id="EE9CCD28B135FFA83A91B2EBFAB654F8" box="[1194,1338,655,681]" italics="true" pageId="4" pageNumber="405">
<taxonomicName id="1BE86AB9B135FFA83A91B2EBFABA54F8" authorityName="Opresko" authorityYear="2005" box="[1194,1334,655,681]" class="Anthozoa" family="Schizopathidae" genus="Umbellapathes" kingdom="Animalia" order="Antipatharia" pageId="4" pageNumber="405" phylum="Cnidaria" rank="species" species="bipinnata">A. bipinnata</taxonomicName>
,
</emphasis>
in that it does not form branches; it has longer pinnules (
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vs.
<quantity id="1B10BCDFB135FFA83D2CB2D6FCE4549C" box="[791,872,690,717]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="405" unit="cm" value="4.5">4.5 cm</quantity>
); much smaller spines (
<quantity id="1B10BCDFB135FFA83A50B2D7FA83549C" box="[1131,1295,691,717]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="4" pageNumber="405" unit="mm" value="0.05" valueMax="0.06" valueMin="0.04">0.040.06 mm</quantity>
vs. up to
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); much larger polyps (
<quantity id="1B10BCDFB135FFA83FF9B2B2FDAC54A0" box="[450,544,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="4" pageNumber="405" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
vs.
<quantity id="1B10BCDFB135FFA83C76B2B3FD2254A0" box="[589,686,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="4" pageNumber="405" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
); and a lower polyp density (45 per
<quantity id="1B10BCDFB135FFA83A74B2B3FB0A54A0" box="[1103,1158,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="405" unit="cm" value="3.0">3 cm</quantity>
vs. about 9 per
<quantity id="1B10BCDFB135FFA83B02B2B3FAF954A0" box="[1337,1397,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="405" unit="cm" value="3.0">3 cm</quantity>
).
</paragraph>
<paragraph id="DC57113AB135FFA83EFCB29FFA035264" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="405">
<taxonomicName id="1BE86AB9B135FFA83EFCB29FFE6C5544" authority="Opresko &amp; Molodtsova, 2021" authorityName="Opresko &amp; Molodtsova" authorityYear="2021" box="[199,480,763,789]" class="Anthozoa" family="Schizopathidae" genus="Alternatipathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="4" pageNumber="405" phylum="Cnidaria" rank="species" species="mirabilis" status="sp. nov.">
<emphasis id="EE9CCD28B135FFA83EFCB29FFE6C5544" box="[199,480,763,789]" italics="true" pageId="4" pageNumber="405">Alternatipathes mirabilis</emphasis>
</taxonomicName>
<emphasis id="EE9CCD28B135FFA83FDCB29FFDAA5544" bold="true" box="[487,550,763,789]" pageId="4" pageNumber="405">
<taxonomicNameLabel id="F5AF7053B135FFA83FDCB29FFDAA5544" box="[487,550,763,789]" pageId="4" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from specimens assigned to
<taxonomicName id="1BE86AB9B135FFA83D81B29FFBCD5544" authorityName="Brook" authorityYear="1889" box="[954,1089,763,789]" class="Anthozoa" family="Schizopathidae" genus="Bathypathes" kingdom="Animalia" order="Antipatharia" pageId="4" pageNumber="405" phylum="Cnidaria" rank="species" species="alternata">
<emphasis id="EE9CCD28B135FFA83D81B29FFBCD5544" box="[954,1089,763,789]" italics="true" pageId="4" pageNumber="405">A. alternata</emphasis>
</taxonomicName>
in having: 1) relatively thicker pinnules (
<quantity id="1B10BCDFB135FFA83F3FB37BFED75568" box="[260,347,799,825]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="405" unit="mm" value="0.8">0.8 mm</quantity>
vs.
<quantity id="1B10BCDFB135FFA83FBDB37AFE505568" box="[390,476,798,825]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="405" unit="mm" value="0.5">0.5 mm</quantity>
basal diameter for pinnules
<quantity id="1B10BCDFB135FFA83D2EB37AFC0F5568" box="[789,899,798,825]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="4" pageNumber="405" unit="cm" value="14.0" valueMax="15.0" valueMin="13.0">1315 cm</quantity>
long); 2) a wider central axial canal (
<quantity id="1B10BCDFB135FFA83B1BB37BFAFB5568" box="[1312,1399,799,825]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="405" unit="mm" value="0.3">0.3 mm</quantity>
vs.
<quantity id="1B10BCDFB135FFA83EACB327FEB6550C" box="[151,314,835,861]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="1.7999999999999998" metricValueMin="1.2" pageId="4" pageNumber="405" unit="mm" value="0.15" valueMax="0.18" valueMin="0.12">0.120.18 mm</quantity>
on pinnules
<quantity id="1B10BCDFB135FFA83FF3B326FDE7550C" box="[456,619,834,861]" metricMagnitude="-4" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.5" metricValueMin="2.2" pageId="4" pageNumber="405" unit="mm" value="0.235" valueMax="0.25" valueMin="0.22">0.220.25 mm</quantity>
in diameter, excluding spines); 3) less of a difference in the relative sizes of the polypar and abpolypar spines (about one-third larger in
<taxonomicName id="1BE86AB9B135FFA83D6FB303FC5455D0" authorityName="Opresko &amp; Molodtsova" authorityYear="2021" box="[852,984,871,897]" class="Anthozoa" family="Schizopathidae" genus="Alternatipathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="4" pageNumber="405" phylum="Cnidaria" rank="species" species="mirabilis">
<emphasis id="EE9CCD28B135FFA83D6FB303FCE955D0" box="[852,869,871,897]" italics="true" pageId="4" pageNumber="405">A</emphasis>
.
<emphasis id="EE9CCD28B135FFA83D48B303FC5455D0" box="[883,984,871,897]" italics="true" pageId="4" pageNumber="405">mirabilis</emphasis>
</taxonomicName>
vs. more than twice as large in
<taxonomicName id="1BE86AB9B135FFA83B04B303FF4555F5" authorityName="Brook" authorityYear="1889" class="Anthozoa" family="Schizopathidae" genus="Bathypathes" kingdom="Animalia" order="Antipatharia" pageId="4" pageNumber="405" phylum="Cnidaria" rank="species" species="alternata">
<emphasis id="EE9CCD28B135FFA83B04B303FADC55D0" box="[1343,1360,871,897]" italics="true" pageId="4" pageNumber="405">A</emphasis>
.
<emphasis id="EE9CCD28B135FFA83B64B303FF4555F5" italics="true" pageId="4" pageNumber="405">alternata</emphasis>
</taxonomicName>
); 4) relatively longer pinnules (
<quantity id="1B10BCDFB135FFA83C0AB3EEFDFB55F4" box="[561,631,906,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="405" unit="cm" value="15.0">15 cm</quantity>
or more vs.
<quantity id="1B10BCDFB135FFA83D3CB3EEFCE855F4" box="[775,868,906,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="405" unit="cm" value="10.5">10.5 cm</quantity>
) on colonies having pinnulated sections of similar length (
<quantity id="1B10BCDFB135FFA83ED6B3CBFEBF5598" box="[237,307,943,969]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="405" unit="cm" value="13.0">13 cm</quantity>
vs.
<quantity id="1B10BCDFB135FFA83F5AB3CBFE265598" box="[353,426,943,969]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="405" unit="cm" value="11.0">11 cm</quantity>
); 5), a wider distal angle of the pinnules at the top of the corallum (6080° vs. 45° or less) and 6) larger polyps (
<quantity id="1B10BCDFB135FFA83FBDB3B6FE6855BC" box="[390,484,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="4" pageNumber="405" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
vs.
<quantity id="1B10BCDFB135FFA83C2AB3B7FDE155BC" box="[529,621,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="405" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
in transverse diameter), resulting in a smaller polyp density (45 vs. 68 per
<quantity id="1B10BCDFB135FFA83EF9B393FF735240" box="[194,255,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="405" unit="cm" value="3.0">3 cm</quantity>
). However, it is important to note that these comparisons are based on a single specimen that is currently assigned to
<taxonomicName id="1BE86AB9B135FFA83F20B47FFDB85264" authorityName="Opresko &amp; Molodtsova" authorityYear="2021" box="[283,564,1051,1077]" class="Anthozoa" family="Schizopathidae" genus="Alternatipathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="4" pageNumber="405" phylum="Cnidaria" rank="species" species="mirabilis">
<emphasis id="EE9CCD28B135FFA83F20B47FFDB85264" box="[283,564,1051,1077]" italics="true" pageId="4" pageNumber="405">Alternatipathes mirabilis</emphasis>
</taxonomicName>
; thus, the ranges reported for this species should not be considered absolute.
</paragraph>
<paragraph id="DC57113AB135FFA83EFCB45BFCCF5294" blockId="4.[151,1437,151,1293]" pageId="4" pageNumber="405">
<taxonomicName id="1BE86AB9B135FFA83EFCB45BFE6C5208" authority="Opresko &amp; Molodtsova, 2021" authorityName="Opresko &amp; Molodtsova" authorityYear="2021" box="[199,480,1087,1113]" class="Anthozoa" family="Schizopathidae" genus="Alternatipathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="4" pageNumber="405" phylum="Cnidaria" rank="species" species="mirabilis" status="sp. nov.">
<emphasis id="EE9CCD28B135FFA83EFCB45BFE6C5208" box="[199,480,1087,1113]" italics="true" pageId="4" pageNumber="405">Alternatipathes mirabilis</emphasis>
</taxonomicName>
<emphasis id="EE9CCD28B135FFA83FDDB45BFDA95208" bold="true" box="[486,549,1087,1113]" pageId="4" pageNumber="405">
<taxonomicNameLabel id="F5AF7053B135FFA83FDDB45BFDA95208" box="[486,549,1087,1113]" pageId="4" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from specimens assigned to
<taxonomicName id="1BE86AB9B135FFA83D83B45BFBA15209" authorityName="Opresko &amp; Wagner" authorityYear="2020" box="[952,1069,1087,1113]" class="Anthozoa" family="Schizopathidae" genus="Alternatipathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="4" pageNumber="405" phylum="Cnidaria" rank="species" species="venusta">
<emphasis id="EE9CCD28B135FFA83D83B45BFBA15209" box="[952,1069,1087,1113]" italics="true" pageId="4" pageNumber="405">A. venusta</emphasis>
</taxonomicName>
in that it has a wider distal angle of the pinnules at the top of the corallum (6080° vs. 30°), no difference between polypar and abpolypar spines, much shorter pinnular spines (
<quantity id="1B10BCDFB135FFA83FDEB4E3FD0552F0" box="[485,649,1159,1185]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="4" pageNumber="405" unit="mm" value="0.05" valueMax="0.06" valueMin="0.04">0.040.06 mm</quantity>
vs.
<quantity id="1B10BCDFB135FFA83C8FB4E3FCD552F0" box="[692,857,1159,1185]" metricMagnitude="-4" metricUnit="m" metricValue="1.65" metricValueMax="2.2" metricValueMin="1.1" pageId="4" pageNumber="405" unit="mm" value="0.165" valueMax="0.22" valueMin="0.11">0.110.22 mm</quantity>
); slightly larger polyps (
<quantity id="1B10BCDFB135FFA83A5EB4E2FB4D52F0" box="[1125,1217,1158,1185]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="4" pageNumber="405" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
vs.
<quantity id="1B10BCDFB135FFA83AD6B4E2FAED52F0" box="[1261,1377,1158,1185]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="4" pageNumber="405" unit="mm" value="4.25" valueMax="5.0" valueMin="3.5">3.55 mm</quantity>
); and a lower polyp density (45 per
<quantity id="1B10BCDFB135FFA83FCEB4CFFDA05294" box="[501,556,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="405" unit="cm" value="3.0">3 cm</quantity>
vs. about 68 per
<quantity id="1B10BCDFB135FFA83CC0B4CFFCBB5294" box="[763,823,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="405" unit="cm" value="3.0">3 cm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="94F242B1B135FFA83EFCB4ABFAA052B8" box="[199,1324,1231,1257]" pageId="4" pageNumber="405" type="etymology">
<paragraph id="DC57113AB135FFA83EFCB4ABFAA052B8" blockId="4.[151,1437,151,1293]" box="[199,1324,1231,1257]" pageId="4" pageNumber="405">
<emphasis id="EE9CCD28B135FFA83EFCB4ABFEC452B8" bold="true" box="[199,328,1231,1257]" pageId="4" pageNumber="405">Etymology</emphasis>
. The species name is derived from the Latin,
<taxonomicName id="1BE86AB9B135FFA83D73B4ABFC2152B8" authorityName="Opresko &amp; Molodtsova" authorityYear="2021" box="[840,941,1231,1257]" class="Anthozoa" family="Schizopathidae" genus="Alternatipathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="4" pageNumber="405" phylum="Cnidaria" rank="species" species="mirabilis">
<emphasis id="EE9CCD28B135FFA83D73B4ABFC2152B8" box="[840,941,1231,1257]" italics="true" pageId="4" pageNumber="405">mirabilis</emphasis>
</taxonomicName>
, meaning “wonderful or strange”.
</paragraph>
</subSubSection>
<subSubSection id="94F242B1B135FFA83EFCB496FACF535C" box="[199,1347,1266,1293]" pageId="4" pageNumber="405" type="distribution">
<paragraph id="DC57113AB135FFA83EFCB496FACF535C" blockId="4.[151,1437,151,1293]" box="[199,1347,1266,1293]" pageId="4" pageNumber="405">
<emphasis id="EE9CCD28B135FFA83EFCB496FED5535D" bold="true" box="[199,345,1266,1292]" pageId="4" pageNumber="405">Distribution</emphasis>
. Known only from a single specimen collected in the North Pacific at a depth of
<quantity id="1B10BCDFB135FFA83ADCB496FAB3535C" box="[1255,1343,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="4.685" pageId="4" pageNumber="405" unit="m" value="4685.0">4685 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>