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<document ID-CLB-Dataset="6049" ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.63775" ID-GBIF-Dataset="e6957b3b-f228-4d6a-b430-35d41cdb2caa" ID-PMC="PMC8222308" ID-Pensoft-Pub="1313-2970-1044-375" ID-Pensoft-UUID="EBD9E81E5EC55AA491952682787793A4" ID-PubMed="34183881" ID-ZooBank="D50CC77C2E6941FDA9BD395B025C43AE" ModsDocID="1313-2970-1044-375" checkinTime="1623875406767" checkinUser="pensoft" docAuthor="Will, Kipling &amp; Gueorguiev, Borislav" docDate="2021" docId="4EFF850B0A235B6EA046ADEC98DACFA3" docLanguage="en" docName="ZooKeys 1044: 375-425" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.63775" docTitle="Adelopomorpha tuberculata Will &amp; Guéorguiev 2021, sp. nov." docType="treatment" docUuid="9E94D72E-997A-4550-B6E7-0FED21757F42" docUuidSource="ZooBank" docVersion="6" id="EBD9E81E5EC55AA491952682787793A4" lastPageNumber="375" masterDocId="EBD9E81E5EC55AA491952682787793A4" masterDocTitle="Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini)" masterLastPageNumber="425" masterPageNumber="375" pageNumber="375" updateTime="1732931690188" updateUser="ExternalLinkService">
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<mods:title id="1B99E1485BBDE87C9907B7C364F7BEBD">Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini)</mods:title>
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<mods:namePart id="993D787CC5FC26E358D43AB818DA9B54">Will, Kipling</mods:namePart>
<mods:nameIdentifier id="578A243FC2FD4BF629362530AF359F68" type="ORCID">https://orcid.org/0000-0002-7056-9011</mods:nameIdentifier>
<mods:affiliation id="71A00AC2BE2243064D9C70B538462059">University of California Berkeley, Essig Museum of Entomology. Berkeley, California, USA</mods:affiliation>
<mods:nameIdentifier id="E48FDB1BCFB295FAE3D7E0C97F4B3168" type="email">kipwill@berkeley.edu</mods:nameIdentifier>
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<mods:namePart id="BE6DF94DFD9254428A8299C825A1B369">Gueorguiev, Borislav</mods:namePart>
<mods:affiliation id="7290B6156DDDBB2300FCD8E33A156F1F">National Museum of Natural History, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
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<mods:date id="DB816782BB6AC09A475F124374C8DA9D">2021</mods:date>
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<mods:classification id="6CB7CA4547BC5D83BD557B89A2DF5CCB">journal article</mods:classification>
<mods:identifier id="C666D70C3D106E343429E8D1A636FA40" type="DOI">http://dx.doi.org/10.3897/zookeys.1044.63775</mods:identifier>
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<treatment id="4EFF850B0A235B6EA046ADEC98DACFA3" ID-GBIF-Taxon="182754023" LSID="urn:lsid:zoobank.org:act:9E94D72E-997A-4550-B6E7-0FED21757F42" httpUri="http://treatment.plazi.org/id/4EFF850B0A235B6EA046ADEC98DACFA3" lastPageNumber="375" pageId="0" pageNumber="375" scope_family="Carabidae" scope_order="Coleoptera" scope_subTribe="Thryptocerina" scope_tribe="Oodini">
<subSubSection id="CE9046E9ACFD3DA92B33F280DE74DDA0" pageId="0" pageNumber="375" type="nomenclature">
<paragraph id="72B6754BCF6A45D9FAA058B713FAD066" pageId="0" pageNumber="375">
<taxonomicName id="FF3EFEDD428C9F814F5483B08BC5375F" ID-CoL="8MKN9" LSID="http://zoobank.org/9E94D72E-997A-4550-B6E7-0FED21757F42" authority="Will &amp; Guéorguiev, 2021" authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Adelopomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adelopomorpha tuberculata" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">Adelopomorpha tuberculata</taxonomicName>
<taxonomicNameLabel id="0DECADB2917A2C38555F76EA02A58326" pageId="0" pageNumber="375">sp. nov.</taxonomicNameLabel>
<figureCitation id="AC9DF8FE973BD0C30EA5F8E7074D28EE" captionStart="Figures 2231" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">Figs 28</figureCitation>
<figureCitation id="3E49F8F18BCD44505DB0781A3C4CFE01" captionStart="Figures 3240" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">, 38</figureCitation>
<figureCitation id="7EC44192D3695D4A468AFBF2825C49BD" captionStart="Figures 41, 42" captionStartId="F9" captionText="Figures 41, 42. Dorsal habitus of 41 Adelopomorpha tuberculata 42 holotype of A. glabra Heller, with labels. Images of A. glabra by O. Jaeger, reproduced with permission from MTKD." figureDoi="10.3897/zookeys.1044.63775.figures41-42" httpUri="https://binary.pensoft.net/fig/554056" pageId="0" pageNumber="375">, 41</figureCitation>
<figureCitation id="0F9A4BEF58D1C01CA5A25FC1A1A471FA" captionStart="Figures 52, 53" captionStartId="F15" captionText="Figures 52, 53. Maps of localities of New Caledonian Oodini." figureDoi="10.3897/zookeys.1044.63775.figures52-53" httpUri="https://binary.pensoft.net/fig/554062" pageId="0" pageNumber="375">, 53</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="53800F8E8E8D637D495AB9DE897009A6" pageId="0" pageNumber="375" type="materials_examined">
<paragraph id="76360252694E034B9375D1C7C7BF1E76" pageId="0" pageNumber="375">Material examined.</paragraph>
<paragraph id="F643B33D79EC7ED58DF185423231672C" pageId="0" pageNumber="375">
<materialsCitation id="F427A96F15084D7617658C6C8C407F80" collectingDate="2004-01-28" collectorName="G. B. Monteith" country="NEW CALEDONIA" elevation="950" location="New Caledonia" longitude="166.0" municipality="Mt. Do" specimenCount="♂" typeStatus="Holotype">
<emphasis id="7455A59A057D12B4715341B7B5DDDB50" bold="true" italics="true" pageId="0" pageNumber="375">
<typeStatus id="5DAE47237861C9BC596EF7F2C23D4F21">Holotype</typeStatus>
</emphasis>
:
<location id="E44DC68F9C93E0FA05024BFF605BA7AD" LSID="urn:lsid:plazi:treatment:4EFF850B0A235B6EA046ADEC98DACFA3:E44DC68F9C93E0FA05024BFF605BA7AD" country="NEW CALEDONIA" longitude="166.0" municipality="Mt. Do" name="New Caledonia">New Caledonia</location>
<specimenCount id="E9A7490D6E4A546D93D0992085B3B954"></specimenCount>
; &quot;
<collectingCountry id="CD3EC4689D0207022356D1799C4E53F4" name="New Caledonia">NEW CALEDONIA</collectingCountry>
<collectingMunicipality id="C555E528FA69DD92AAA41D1283A4E92B">Mt. Do</collectingMunicipality>
<quantity id="34FDD5EF0FD5538C6248E98F1C612213" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="km" value="0.5">0.5km</quantity>
below summit,
<collectingDate id="77D8C2BA3B29F8BA82380B18EF49358A" value="2004-01-28">28Jan2004</collectingDate>
<collectorName id="93036828AF35A4E1AA63B7171ED1C7EB">G.B.Monteith</collectorName>
&quot;// &quot; QM berleseate 1111 21°45'Sx
<geoCoordinate id="B942CF8645D1C1DB1C615129ED5648BA" degrees="166" direction="east" minutes="00" orientation="longitude" precision="925" value="166.0">166°00'E</geoCoordinate>
, rainforest,
<quantity id="5EF7BC2CF3E7DE3C77F3064A89807690" metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">
<elevation id="F9F32EE382D866F659B1357D40D71932" metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">950 m</elevation>
</quantity>
, sieved litter&quot;
</materialsCitation>
<typeStatus id="8359364D29F75EF84B8E3F5565353A44">Holotype</typeStatus>
pinned, with genitalia in a separate microvial. Source collection QM, deposited MNHN. Type locality as given on label.
</paragraph>
<paragraph id="AA363FB0E946437CDFC767D27EE3CF4C" pageId="0" pageNumber="375">
<materialsCitation id="1C7E7272366BC32555AA38C57260AFA7" collectingDate="1984-05-23" collectingDateMax="2003-11-22" collectingDateMin="1984-05-23" collectorName="G. Monteith, D. Cook" country="NEW CALEDONIA" county="Mt. Dzumac" elevation="1000" location="Sx" longitude="166.0" municipality="Mt. Do" specimenCount="♂" typeStatus="Paratypes">
<emphasis id="EB90883039991A838A3E47B02985A661" bold="true" italics="true" pageId="0" pageNumber="375">
<typeStatus id="B7165F85A869D479907788F992DD6530">Paratypes</typeStatus>
</emphasis>
: New Caledonia •
<specimenCount id="CB659A041A672BC3DA604994A1DBA910"></specimenCount>
; &quot;
<collectingCountry id="FDDCDFFEA79A75FD41425C84C96FA187" name="New Caledonia">NEW CALEDONIA</collectingCountry>
11420 21°45'
<location id="37AF6122D6E69B6E56C1FE865AAB4D16" LSID="urn:lsid:plazi:treatment:4EFF850B0A235B6EA046ADEC98DACFA3:37AF6122D6E69B6E56C1FE865AAB4D16" country="NEW CALEDONIA" county="Mt. Dzumac" longitude="166.0" municipality="Mt. Do" name="Sx">Sx</location>
<geoCoordinate id="7F610C47AE428D666A62CA02208456B9" degrees="166" direction="east" minutes="00" orientation="longitude" precision="925" value="166.0">166°00'E</geoCoordinate>
,
<collectingMunicipality id="0BF20D6EE50FABAEA750AC4878B83F47">Mt. Do</collectingMunicipality>
summit,
<quantity id="C1F38E860AF3B08924653A80A868040A" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
<elevation id="8FF0A93BD3DF005F4EB7605817AC6857" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000m</elevation>
</quantity>
,
<collectingDate id="56DDA5E87B0CBB6E4BA1E95162B269E6" value="2003-11-22">22 Nov 2003</collectingDate>
,
<collectorName id="972A78CB7CF6A4622045461397A12C01">G. Monteith</collectorName>
, Pyrethrum, trees &amp; logs.&quot; [QM] •
<specimenCount id="0DD81D1D89CEFE238D81D8211BE12B6E"></specimenCount>
; &quot;
<collectingCountry id="E885DE08E2FCE5EA775FBB7B34784613" name="New Caledonia">NEW CALEDONIA</collectingCountry>
<collectingCounty id="362F77C360E9D60D09297F1B8D987529">Mt. Dzumac</collectingCounty>
,
<quantity id="5255E9D4A26B0846818115D19D8A4E5E" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">
<elevation id="461F7335A21351339E692C2C3E38E7B2" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">800-1000m</elevation>
</quantity>
,
<collectingDate id="5869481C5EA013A8E387E8573B4A7C3F" value="1984-05-23">23 May 1984</collectingDate>
<collectorName id="33E27CBF8B7ECCF21A167D0301B42330">G. Monteith</collectorName>
&amp;
<collectorName id="5B7EFA983038A7DDA127D0EB0D096CD1">D. Cook</collectorName>
&quot; [QM]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AD9321C9C42177EDF34D59D2785F08C7" pageId="0" pageNumber="375" type="diagnosis">
<paragraph id="7DC0B64575F3B9EEA3849325A4DC472F" pageId="0" pageNumber="375">Diagnosis.</paragraph>
<paragraph id="71AA6377ED209A0A97902F54AAAD6F24" pageId="0" pageNumber="375">
This species is nearly indistinguishable externally from
<taxonomicName id="BD32A47170E76C9C77C1E9D340BFF470" lsidName="A. glabra" pageId="0" pageNumber="375" rank="species" species="glabra">
<emphasis id="185B4D8120E5973EDC6DFD2C74311DD2" italics="true" pageId="0" pageNumber="375">A. glabra</emphasis>
</taxonomicName>
except for the lack of the depressed lateral 1/3 of the pronotum present in
<taxonomicName id="62B9C8B8167D9505B9C95E0F2E44475C" lsidName="A. glabra" pageId="0" pageNumber="375" rank="species" species="glabra">
<emphasis id="05E8D7590614D44D61D1A556008FAA8E" italics="true" pageId="0" pageNumber="375">A. glabra</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="0C0A12906820E61E93DC19E6F78FC201" captionStart="Figures 43, 44" captionStartId="F10" captionText="Figures 43, 44. Forebody, left dorsolateral view of 43 Adelopomorpha glabra Heller 44 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures43-44" httpUri="https://binary.pensoft.net/fig/554057" pageId="0" pageNumber="375">43</figureCitation>
), and its smaller size. Also, it is smaller than
<taxonomicName id="3CCCF99D6351AA8F8A9E6378DBD33540" lsidName="A. tethys" pageId="0" pageNumber="375" rank="species" species="tethys">
<emphasis id="B898E228C17A7ECE49456DEEE93F8EA3" italics="true" pageId="0" pageNumber="375">A. tethys</emphasis>
</taxonomicName>
, whose male has setae on ventrite 6. See diagnosis above. The aedeagus is decisively different (Fig.
<figureCitation id="AA6572B64C4E46A93841961BEB8E6BAB" captionStart="Figures 2231" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">28</figureCitation>
), with a relatively long and moderately, narrowly rounded tip.
</paragraph>
</subSubSection>
<subSubSection id="A8C1A6CA9969BF92FA14C8A497D89500" pageId="0" pageNumber="375" type="description">
<paragraph id="7366CDB6BD72497B97DFA44663D98DEE" pageId="0" pageNumber="375">Description.</paragraph>
<paragraph id="5D36B2DD626E77C6DEB8F6BD71775AD2" pageId="0" pageNumber="375">
<emphasis id="A622EFD3A75A74065E5FD543F7D54072" bold="true" pageId="0" pageNumber="375">
<emphasis id="573BF8347C9DBE1B4EDE34EBE3467C16" italics="true" pageId="0" pageNumber="375">Habitus</emphasis>
.
</emphasis>
Very small sized, BL: 4.90 mm and BW: 2.45 mm (range, BL: 4.75-5.00 mm, BW: 2.45-2.60 mm), with rather oval and convex body (Fig.
<figureCitation id="046985AF367B38CED0DC34DFAC8B36D5" captionStart="Figures 41, 42" captionStartId="F9" captionText="Figures 41, 42. Dorsal habitus of 41 Adelopomorpha tuberculata 42 holotype of A. glabra Heller, with labels. Images of A. glabra by O. Jaeger, reproduced with permission from MTKD." figureDoi="10.3897/zookeys.1044.63775.figures41-42" httpUri="https://binary.pensoft.net/fig/554056" pageId="0" pageNumber="375">41</figureCitation>
).
<emphasis id="F6755CD68B1A7E1F3588C29D23FCF7E0" bold="true" pageId="0" pageNumber="375">
<emphasis id="16771C8F91FBF1B3F5364DC511A7C8DC" italics="true" pageId="0" pageNumber="375">Color and luster</emphasis>
.
</emphasis>
Head dark reddish brown; pronotum, elytra and ventral surface of body black, pronotum marginally diaphanous reddish black; legs deep reddish brown. Deep reddish black medially at the very apical edge of ventrite 6; antennae and palpi brown. Integument moderately dull, with slight, diffused spectral iridescence dorsally and ventrally.
<emphasis id="51E76B4C82E5AB03FE1DB629F787D685" bold="true" pageId="0" pageNumber="375">
<emphasis id="14C65E4FE2A31C430ECE7C12F7459737" italics="true" pageId="0" pageNumber="375">Microsculpture and punctation</emphasis>
.
</emphasis>
Dorsal surface of head and pronotum with prominent isodiametric meshes; elytra with irregular mesh of more or less transversely stretched sculpticells; ventral surface with scarcely-visible sculpticells on ventrite 6 and prosternum, otherwise sculpticells mostly not apparent. Head lacking macropunctation or wrinkles on vertex; head, pronotum and elytra without micropunctation. All abdominal ventrites very shallowly wrinkled to nearly smooth.
<emphasis id="E5B868A5841D4E15E2428007D93F6818" bold="true" pageId="0" pageNumber="375">
<emphasis id="25FF78AE44A1A38F0DB34C7976C120F5" italics="true" pageId="0" pageNumber="375">Head</emphasis>
.
</emphasis>
Slightly more than 1/4 as wide as pronotum (Suppl. material 2: Table S1). Eye moderate sized, somewhat protruded, EyW/HW: 1.31. Frontoclypeal sutures not evident. Mentum tooth without paramedial border.
<emphasis id="75982C45B2755D24928BF923559C5618" bold="true" pageId="0" pageNumber="375">
<emphasis id="42142CEDDCFC7C6C512335041797DEA7" italics="true" pageId="0" pageNumber="375">Thorax</emphasis>
.
</emphasis>
Pronotum 1 2/3 wider than long (PW/PL: 1.67 (range, 1.71-1.76)), with sides evenly, very shallowly rounded from anterior angles to posterior ones; maximal width in posterior 1/5; width at apex ~ 1/3 that of widest point (PW/PA: 2.88 (range 2.82-3.00)). Posterior angles ~ right angled, broadly rounded. Mesosternum with single, very prominent, medial tubercle. Metepisternum 1 1/2
<normalizedToken id="3DD4889392C0A736E7B8DD2CC6B50769" originalValue="×">x</normalizedToken>
wider than long, with lateral margin straight, coadunation with epipleuron along entire length.
<emphasis id="94C8D69265FA3DE6BB19E9427C6CBD4E" bold="true" pageId="0" pageNumber="375">
<emphasis id="4B7D44C28BDF192E5E7D345F1936B842" italics="true" pageId="0" pageNumber="375">Elytra</emphasis>
.
</emphasis>
Broad, as wide as long (EL/EW: 1.00 (range, 1.00-1.02)). Basal margin distinct, forming small, sharp tooth at shoulder, ended medially at level of parascutellar punctures, joined to parascutellar puncture fovea by a short, shallow striole. Apical sinuation not evident.
<emphasis id="84C9C3704D131B41251A32809494C4E9" bold="true" pageId="0" pageNumber="375">
<emphasis id="2F99699EF49F79CB6D9224C12AE4EAAD" italics="true" pageId="0" pageNumber="375">Female genitalia</emphasis>
.
</emphasis>
Female specimens unknown.
<emphasis id="5BB2558E58CD719476290ED67A5167C7" bold="true" pageId="0" pageNumber="375">
<emphasis id="1F2C9B48D480DD86EB57CF5F1AA1B208" italics="true" pageId="0" pageNumber="375">Male genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus in lateral view long, almost straight (Fig.
<figureCitation id="A31F77948C6676F1CE980CC450FC64F0" captionStart="Figures 3240" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">38</figureCitation>
); lobe in dorsal view with apical lamella moderately long, rounded, triangular and oriented to right (Fig.
<figureCitation id="300EC2CCE1495AF654010D7B440F4D9D" captionStart="Figures 2231" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">28</figureCitation>
); sclerotized portion of endophallus visible in repose, with two sclerites.
</paragraph>
</subSubSection>
<subSubSection id="E0D0F00F091D4447C799B80271558C4D" pageId="0" pageNumber="375" type="etymology">
<paragraph id="F8F41EBEA893E39905808401C85C3296" pageId="0" pageNumber="375">Etymology.</paragraph>
<paragraph id="B9E7E54BEF49E48892501B4BBD51DC89" pageId="0" pageNumber="375">
The specific epithet
<taxonomicName id="A1C82DE9EDAEAEB938C2883AC2692730" authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Adelopomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adelopomorpha tuberculata" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="F86085D028E477C4EB61C6BED8338255" italics="true" pageId="0" pageNumber="375">Adelopomorpha tuberculata</emphasis>
</taxonomicName>
draws attention to the relatively large, median mesosternal tubercle in these beetles. It is treated as an adjective in the nominative singular.
</paragraph>
</subSubSection>
</treatment>
</document>