307 lines
25 KiB
XML
307 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.49.8383" ID-PMC="PMC4432232" ID-Pensoft-Pub="1314-2003-49-33" ID-Pensoft-UUID="5D3B5F37691CEB6EEB4BFFC1FFADD024" ID-PubMed="25987884" ID-Zenodo-Dep="576299" ModsDocID="1314-2003-49-33" checkinTime="1451251539483" checkinUser="pensoft" docAuthor="Pedraza-Penalosa, Paola" docDate="2015" docId="4EC468444000EB69531B6F07D4AC8218" docLanguage="en" docName="PhytoKeys 49: 33-58" docOrigin="PhytoKeys 49" docPubDate="2015-04-22" docSource="http://dx.doi.org/10.3897/phytokeys.49.8383" docTitle="Satyria orquidiensis Pedraza 2015, sp. nov." docType="treatment" docVersion="6" id="5D3B5F37691CEB6EEB4BFFC1FFADD024" lastPageNumber="44" masterDocId="5D3B5F37691CEB6EEB4BFFC1FFADD024" masterDocTitle="New blueberry and mortino relatives (Ericaceae) from northwestern Colombia" masterLastPageNumber="58" masterPageNumber="33" pageNumber="41" updateTime="1668141352410" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New blueberry and mortino relatives (Ericaceae) from northwestern Colombia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pedraza-Penalosa, Paola</mods:namePart>
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<mods:affiliation>Institute of Systematic Botany, The New York Botanical Garden, 2900 Southern Blvd., Bronx, NY 10458</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-04-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>49</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>58</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.49.8383</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.49.8383</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-49-33</mods:identifier>
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<mods:identifier type="Pensoft-UUID">5D3B5F37691CEB6EEB4BFFC1FFADD024</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576299</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152025699" LSID="urn:lsid:plazi:treatment:4EC468444000EB69531B6F07D4AC8218" httpUri="http://treatment.plazi.org/id/4EC468444000EB69531B6F07D4AC8218" lastPageId="11" lastPageNumber="44" pageId="8" pageNumber="41">
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<subSubSection pageId="8" pageNumber="41" type="nomenclature">
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<paragraph pageId="8" pageNumber="41">
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<taxonomicName LSID="4EC46844-4000-EB69-531B-6F07D4AC8218" authority="Pedraza" authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria orquidiensis" order="Ericales" pageId="8" pageNumber="41" phylum="Tracheophyta" rank="species" species="orquidiensis" status="sp. nov.">
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<pageBreakToken pageId="8" pageNumber="41" start="start">Satyria</pageBreakToken>
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orquidiensis Pedraza
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</taxonomicName>
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<taxonomicNameLabel pageId="8" pageNumber="41">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Illustration of Satyria orquidiensis. A Branch with clustered leaves B Branch with inflorescences C Flower attached to the rachis with detail of a bracteole D Longitudinal section of a flower E Stamens in abaxial, adaxial, and lateral views, respectively F Fruit. [Drawn from the type, P. Pedraza-Penalosa et al. 2436 and 2447.]" pageId="8" pageNumber="41">Figures 9</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Satyria orquidiensis. A Branch with clustered leaves, adaxial side B Branch with clustered leaves, abaxial side C Cauliflorous inflorescences D Nearly mature (right) and immature fruits (left). [Photos by P. Pedraza-Penalosa.]" pageId="8" pageNumber="41">, 10</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Satyria orquidiensis. A Flowers at anthesis with detail of a longitudinal section B Close-up of flowers showing the ornamented calyces. [Photos by P. Pedraza-Penalosa.]" pageId="8" pageNumber="41">, 11</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="42" type="diagnosis">
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<paragraph pageId="9" pageNumber="42">
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<pageBreakToken pageId="9" pageNumber="42" start="start">Diagnosis</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="9" pageNumber="42">
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<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria orquidiensis" order="Ericales" pageId="9" pageNumber="42" phylum="Tracheophyta" rank="species" species="orquidiensis">
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<emphasis italics="true" pageId="9" pageNumber="42">Satyria orquidiensis</emphasis>
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</taxonomicName>
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has many distinctive characters and it differs from other species in the genus because its leaves, which are clustered (up to 8 leaves) and seemingly verticillate (the only one in the genus), oblanceolate, large, and basally subcordate. The abundant and large flowers of this new species also stand out because of the big ribs on the calyx tube and the sharp transversal constriction between the limb and tube, which gives the entire calyx a broad campanulate shape. The long obconic corolla (up to 4.4 cm long) is orange and green at the tip. Both calyx and corolla are covered with a fine glabrate indumentum.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="42" type="type">
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<paragraph pageId="9" pageNumber="42">Type.</paragraph>
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<paragraph pageId="9" pageNumber="42">
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COLOMBIA. Antioquia: Municipio Urrao. Corregimiento La
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<normalizedToken originalValue="Encarnación">Encarnacion</normalizedToken>
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, vereda Calles, Parque Nacional Natural Las
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<normalizedToken originalValue="Orquídeas">Orquideas</normalizedToken>
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, finca de Alfonso Pino, en la divisoria de aguas entre las quebradas La Virola y El Bosque, al noroccidente de la
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<normalizedToken originalValue="cabaña">cabana</normalizedToken>
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de Calles,
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<geoCoordinate degrees="6" direction="north" minutes="31" orientation="latitude" precision="15" seconds="35" value="6.5263886">6°31'35"N</geoCoordinate>
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;
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<geoCoordinate degrees="76" direction="west" minutes="15" orientation="longitude" precision="15" seconds="50" value="-76.263885">76°15'50"W</geoCoordinate>
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, 1450-1470 m, 27 Jan 2011 (fl, fr),
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<emphasis italics="true" pageId="9" pageNumber="42">
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Paola
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<normalizedToken originalValue="Pedraza-Peñalosa">Pedraza-Penalosa</normalizedToken>
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et al. 2016
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</emphasis>
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(holotype: COL!; isotypes, CAS!, CUVC, E!, HUA, MO!, NY! [NY02058404], PSO).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="43" pageId="9" pageNumber="42" type="description">
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<paragraph pageId="9" pageNumber="42">Description.</paragraph>
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<paragraph lastPageId="10" lastPageNumber="43" pageId="9" pageNumber="42">
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Epiphytic
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<emphasis italics="true" pageId="10" pageNumber="43">
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<pageBreakToken pageId="10" pageNumber="43" start="start">shrub</pageBreakToken>
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</emphasis>
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, lianoid; stems brown-grey, terete, glabrous, bark smooth; twigs terete, smooth, glabrous. Axillary buds compressed; prophylls 2, valvate, lanceolate, inconspicuous, 2.5-4.5 mm long, margin eciliate, apex acuminate, abaxially puberulous, the hairs inconspicuous, ca. <0.2 mm long, eglandular and unicellular (all indumentum composed of this type of hairs except when indicated).
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<emphasis italics="true" pageId="10" pageNumber="43">Leaves</emphasis>
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alternate but apparently pseudoverticillate, originating in clusters of (2-)5-8 leaves separated by leafless sections 11.5-17.5 cm long; petiole terete, pulvinate, 3-6 mm long, glabrate; lamina coriaceous, elliptic, sometimes oblanceolate, (14-)18.7-28
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<normalizedToken originalValue="×">x</normalizedToken>
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(3-)5-8 cm, base subcordate, margin entire and eciliate, apex acuminate, adaxially glabrous, abaxially glabrate (but appearing glabrous), the hairs inconspicuous, (<0.2 mm long), caducous but with persistent bases that are red and swollen, apparently multicellular and glandular; laminar glands inconspicuous, basal, rounded; venation plinerved (acrodromous), suprabasal, with 3-4 visible orders (consistently well-marked up to 3th order) in dry specimens, prominent lateral veins 2(-3) per side, subopposite or alternate, concentrated in the basal third, ascending, brochidodromous, midvein and secondaries adaxially impressed and abaxially raised.
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<emphasis italics="true" pageId="10" pageNumber="43">Inflorescence</emphasis>
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a 4-6-flowered raceme, more than one raceme arising from an axillary pad (pincushion like), often cauliflorous; inflorescence and floral bracts alike, persistent, chartaceous, ovate, 0.5-2.6
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<normalizedToken originalValue="×">x</normalizedToken>
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0.8-1.2 mm, margin entire and ciliolate, the hairs inconspicuous and eglandular, apex acute to acuminate, adaxially glabrous, abaxially glabrate, venation obscure; rachis orange, 4-8.5 mm long, glabrate, warts abundant and minute; pedicel dark or lightly orange, articulate with calyx, 1.9-2.3 cm long (3 cm when fruiting), basally (1-1.2 mm diam.) less than half the diameter of apex (3-4.5 mm) (
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<emphasis italics="true" pageId="10" pageNumber="43">in vivo</emphasis>
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2 mm basally, 5-7 mm apically), with the apex becoming discoid and almost as wide as the calyx tube, glabrate, minute warts present at very base both
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<emphasis italics="true" pageId="10" pageNumber="43">in vivo</emphasis>
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and when dry; bracteoles 2, basal, supopposite to opposite, chartaceous, ovate, 2-2.4
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<normalizedToken originalValue="×">x</normalizedToken>
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0.6-0.8 mm, margin entire and ciliolate, the hairs inconspicuous, caducous and eglandular, apex acuminate, glabrous on both sides, venation obscure.
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<emphasis italics="true" pageId="10" pageNumber="43">Flowers</emphasis>
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5-merous (some calyces 6-merous), actinomorphic, diplostemonous. Calyx dark or light orange, campanulate, with a marked transversal constriction between limb and tube, inconspicuously 5-angled, the angles alternating with lobes, conspicuously ribbed in the tube and with softer ribs in the limb, 4.1-5.8 mm long (6-7.2 mm
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<emphasis italics="true" pageId="10" pageNumber="43">in vivo</emphasis>
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), glabrate, the hairs inconspicuous, <0.1 mm long; tube oblate, 2-2.2 mm long (2.5-3 mm
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<emphasis italics="true" pageId="10" pageNumber="43">in vivo</emphasis>
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); limb slightly flaring, 2.1-3.6 mm long (3.5-4.7 mm
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<emphasis italics="true" pageId="10" pageNumber="43">in vivo</emphasis>
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); lobes deltate, 1-2
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<normalizedToken originalValue="×">x</normalizedToken>
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3.5-4.5 mm long (1.6-2
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<normalizedToken originalValue="×">x</normalizedToken>
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5 mm
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<emphasis italics="true" pageId="10" pageNumber="43">in vivo</emphasis>
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), margin entire, eglandular, and eciliate, apex acute; sinuses practically flat (broadly U-shaped
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<emphasis italics="true" pageId="10" pageNumber="43">in vivo</emphasis>
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); aestivation valvate. Corolla basal two thirds orange, apical third green, fleshy, not bistratose, obconic, 3.2-4 cm long (3.4-4.4 cm
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<emphasis italics="true" pageId="10" pageNumber="43">in vivo</emphasis>
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), 1-1.2 cm diam. (1-1.1 cm
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<emphasis italics="true" pageId="10" pageNumber="43">in vivo</emphasis>
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), 3 mm wide at throat (4.7-5.5 mm
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<emphasis italics="true" pageId="10" pageNumber="43">in vivo</emphasis>
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), inconspicuously 5-angled, glabrate without, the hairs inconspicuous, <0.1 mm long, glabrous within; lobes ovate, 1.5-2
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<normalizedToken originalValue="×">x</normalizedToken>
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1-2 mm, but sinuses often further tearing toward end of anthesis and the lobes then oblong and 4.8-7
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<normalizedToken originalValue="×">x</normalizedToken>
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1-2 mm, apex acute, not strongly reflexed at maturity; aestivation valvate.
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<emphasis italics="true" pageId="10" pageNumber="43">Stamens</emphasis>
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10, dimorphic, staminal cycles with different anther lengths and dehiscence orientation, included, not adherent to corolla. Long stamens 10-11.2 mm long; filaments connate into a tube, straight, 3.5-4 mm long, glabrous or distally glabrate on the abaxial side, the hairs a handful, inconspicuous, multicellular and apparently eglandular; anthers 9-9.5 mm long, slightly prognathous, narrowing at base and widening at apex, without a clear distinction between tubules and thecae; thecae 6-6.5 mm long, without basal appendage, minutely papillate at least basally; tubules 2, free, turned inwards like
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<normalizedToken originalValue="bull’s">bull's</normalizedToken>
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horns, 3 mm long, smooth, dehiscing by latrorse elliptical slits 3 mm long, distal margin ornamented with small and irregular lobes. Short stamens 9-10 mm long, same shape, indumentum and features as long stamens except when indicated; filaments 3.5-4 mm long; anthers 8-8.5 mm long; thecae 5.5 mm long; tubules pointing upwards and without space between them, 2.5-3 mm long, dehiscing by introrse elliptical slits 2.5-3 mm long. Nectary not pulvinate, slightly concave to flat, glabrous; style white, 2.6-3.7 cm long, included; stigma punctiform.
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<emphasis italics="true" pageId="10" pageNumber="43">Berry</emphasis>
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cream-colored and ribbed when immature, turning purple at maturity.
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</paragraph>
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<caption pageId="10" pageNumber="43" start="Figure 9" startId="F9">
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<paragraph pageId="10" pageNumber="43">
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<emphasis bold="true" pageId="10" pageNumber="43">Figure 9.</emphasis>
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Illustration of
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<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria orquidiensis" order="Ericales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="orquidiensis">
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<emphasis italics="true" pageId="10" pageNumber="43">Satyria orquidiensis</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="10" pageNumber="43">A</emphasis>
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Branch with clustered leaves
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<emphasis bold="true" pageId="10" pageNumber="43">B</emphasis>
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Branch with inflorescences
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<emphasis bold="true" pageId="10" pageNumber="43">C</emphasis>
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Flower attached to the rachis with detail of a bracteole
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<emphasis bold="true" pageId="10" pageNumber="43">D</emphasis>
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Longitudinal section of a flower
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<emphasis bold="true" pageId="10" pageNumber="43">E</emphasis>
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Stamens in abaxial, adaxial, and lateral views, respectively
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<emphasis bold="true" pageId="10" pageNumber="43">F</emphasis>
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Fruit. [Drawn from the type,
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<emphasis italics="true" pageId="10" pageNumber="43">
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P.
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<normalizedToken originalValue="Pedraza-Peñalosa">Pedraza-Penalosa</normalizedToken>
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et al. 2436
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</emphasis>
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and
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<emphasis italics="true" pageId="10" pageNumber="43">2447</emphasis>
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.]
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</paragraph>
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</caption>
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<caption pageId="10" pageNumber="43" start="Figure 10" startId="F10">
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<paragraph pageId="10" pageNumber="43">
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<emphasis bold="true" pageId="10" pageNumber="43">Figure 10.</emphasis>
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<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria orquidiensis" order="Ericales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="orquidiensis">
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<emphasis italics="true" pageId="10" pageNumber="43">Satyria orquidiensis</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="10" pageNumber="43">A</emphasis>
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Branch with clustered leaves, adaxial side
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<emphasis bold="true" pageId="10" pageNumber="43">B</emphasis>
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Branch with clustered leaves, abaxial side
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<emphasis bold="true" pageId="10" pageNumber="43">C</emphasis>
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Cauliflorous inflorescences
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<emphasis bold="true" pageId="10" pageNumber="43">D</emphasis>
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Nearly mature (right) and immature fruits (left). [Photos by P.
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<normalizedToken originalValue="Pedraza-Peñalosa">Pedraza-Penalosa</normalizedToken>
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.]
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</paragraph>
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</caption>
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<caption pageId="10" pageNumber="43" start="Figure 11" startId="F11">
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<paragraph pageId="10" pageNumber="43">
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<emphasis bold="true" pageId="10" pageNumber="43">Figure 11.</emphasis>
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<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria orquidiensis" order="Ericales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="orquidiensis">
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<emphasis italics="true" pageId="10" pageNumber="43">Satyria orquidiensis</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="10" pageNumber="43">A</emphasis>
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Flowers at anthesis with detail of a longitudinal section
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<emphasis bold="true" pageId="10" pageNumber="43">B</emphasis>
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Close-up of flowers showing the ornamented calyces. [Photos by P.
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<normalizedToken originalValue="Pedraza-Peñalosa">Pedraza-Penalosa</normalizedToken>
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.]
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="43" type="distribution and ecology">
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<paragraph pageId="10" pageNumber="43">Distribution and ecology.</paragraph>
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<paragraph pageId="10" pageNumber="43">
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<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria orquidiensis" order="Ericales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="orquidiensis">
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<emphasis italics="true" pageId="10" pageNumber="43">Satyria orquidiensis</emphasis>
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</taxonomicName>
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is endemic to Antioquia (Colombia) and it only known from collections from Las
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<normalizedToken originalValue="Orquídeas">Orquideas</normalizedToken>
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National Park. It is known to flower in January and fruit in January and July. This species grows in the canopy of humid premontane and montane forests at 880-1470 m, but it is possible that it could be found at lower altitudes as well.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="44" type="etymology">
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<paragraph pageId="11" pageNumber="44">
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<pageBreakToken pageId="11" pageNumber="44" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="11" pageNumber="44">
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Named after Las
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<normalizedToken originalValue="Orquídeas">Orquideas</normalizedToken>
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National Park (Colombia), where the species is endemic to.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="44" type="preliminary conservation status">
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<paragraph pageId="11" pageNumber="44">Preliminary conservation status.</paragraph>
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<paragraph pageId="11" pageNumber="44">
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<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria orquidiensis" order="Ericales" pageId="11" pageNumber="44" phylum="Tracheophyta" rank="species" species="orquidiensis">
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<emphasis italics="true" pageId="11" pageNumber="44">Satyria orquidiensis</emphasis>
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</taxonomicName>
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it is only known by specimens collected in Las
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<normalizedToken originalValue="Orquídeas">Orquideas</normalizedToken>
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National Park. It is commonly observed in conserved premontane and montane forests, but because it is a liana normally found high in the canopy, only a few collections exits. Because the observed abundance within the protected area I consider this species of least concern.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="44" type="discussion">
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<paragraph pageId="11" pageNumber="44">Discussion.</paragraph>
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<paragraph pageId="11" pageNumber="44">
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Morphologically,
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<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria orquidiensis" order="Ericales" pageId="11" pageNumber="44" phylum="Tracheophyta" rank="species" species="orquidiensis">
|
||
<emphasis italics="true" pageId="11" pageNumber="44">Satyria orquidiensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria pterocalyx" order="Ericales" pageId="11" pageNumber="44" phylum="Tracheophyta" rank="species" species="pterocalyx">
|
||
<emphasis italics="true" pageId="11" pageNumber="44">Satyria pterocalyx</emphasis>
|
||
</taxonomicName>
|
||
stand out within the genus and have more similarities among themselves than with other species; these are the only species in the genus with conspicuous wings and/or ribs on their calyces. Although their large corollas share similar colors and obconic shape, they can be easily differentiated because
|
||
<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria orquidiensis" order="Ericales" pageId="11" pageNumber="44" phylum="Tracheophyta" rank="species" species="orquidiensis">
|
||
<emphasis italics="true" pageId="11" pageNumber="44">Satyria orquidiensis</emphasis>
|
||
</taxonomicName>
|
||
has corollas inconspicuously 5-angled (vs. terete in
|
||
<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria pterocalyx" order="Ericales" pageId="11" pageNumber="44" phylum="Tracheophyta" rank="species" species="pterocalyx">
|
||
<emphasis italics="true" pageId="11" pageNumber="44">Satyria pterocalyx</emphasis>
|
||
</taxonomicName>
|
||
), orange with green lobes (vs. red-orange with the tube and lobes green-whitish) that gradually decrease in diameter toward the lobes (vs. dramatically constricted at the throat which is elongated into a tube ca. 8 mm long). Also, the dark or light orange (vs. light green) calyces of
|
||
<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria orquidiensis" order="Ericales" pageId="11" pageNumber="44" phylum="Tracheophyta" rank="species" species="orquidiensis">
|
||
<emphasis italics="true" pageId="11" pageNumber="44">Satyria orquidiensis</emphasis>
|
||
</taxonomicName>
|
||
are conspicuously ribbed on the tube and sharply constricted between the limb and tube (vs. calyces 5-winged, tube with two basal lobes in each of the facets demarked by the wings, not transversally constricted). Vegetetatively, these two new species are even more distinct as
|
||
<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria orquidiensis" order="Ericales" pageId="11" pageNumber="44" phylum="Tracheophyta" rank="species" species="orquidiensis">
|
||
<emphasis italics="true" pageId="11" pageNumber="44">Satyria orquidiensis</emphasis>
|
||
</taxonomicName>
|
||
has leaves that are clustered and seemingly verticillate (vs. not clustered in
|
||
<taxonomicName authorityName="Pedraza" authorityYear="2015" class="Magnoliopsida" family="Ericaceae" genus="Satyria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Satyria pterocalyx" order="Ericales" pageId="11" pageNumber="44" phylum="Tracheophyta" rank="species" species="pterocalyx">
|
||
<emphasis italics="true" pageId="11" pageNumber="44">Satyria pterocalyx</emphasis>
|
||
</taxonomicName>
|
||
), larger [(14-)18.7-28 cm long vs. 14-18 cm], basally subcordate (vs. obtuse or cuneate), apically acute to acuminate (long acuminate, acumen 1.8-2.8 cm long), and with inconspicuous basal laminar glands (vs. absent).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="11" pageNumber="44" type="specimens examined">
|
||
<paragraph pageId="11" pageNumber="44">Specimens examined.</paragraph>
|
||
<paragraph pageId="11" pageNumber="44">
|
||
<emphasis bold="true" pageId="11" pageNumber="44">COLOMBIA. Antioquia:</emphasis>
|
||
Municipio Frontino, Vereda Venados Abajo, Parque Nacional Natural Las
|
||
<normalizedToken originalValue="Orquídeas">Orquideas</normalizedToken>
|
||
, sector Venados, sitio La Miquera,
|
||
<geoCoordinate degrees="6" direction="north" minutes="32" orientation="latitude" precision="1" seconds="28.1" value="6.5411386">6°32'28.1"N</geoCoordinate>
|
||
;
|
||
<geoCoordinate degrees="76" direction="west" minutes="18" orientation="longitude" precision="1" seconds="05.3" value="-76.301476">76°18'05.3"W</geoCoordinate>
|
||
, 1000-1030 m, 27 Jul 2011 (fr),
|
||
<emphasis italics="true" pageId="11" pageNumber="44">
|
||
P.
|
||
<normalizedToken originalValue="Pedraza-Peñalosa">Pedraza-Penalosa</normalizedToken>
|
||
et al. 2436
|
||
</emphasis>
|
||
[COL!, NY! (2 sheets)]; Municipio Frontino, Vereda Venados Abajo, Parque Nacional Natural Las
|
||
<normalizedToken originalValue="Orquídeas">Orquideas</normalizedToken>
|
||
, sector Venados, sitio La Esperanza, cuenca de la quebrada Arenales,
|
||
<geoCoordinate degrees="6" direction="north" minutes="42" orientation="latitude" precision="1" seconds="06.8" value="6.7018886">6°42'06.8"N</geoCoordinate>
|
||
;
|
||
<geoCoordinate degrees="76" direction="west" minutes="18" orientation="longitude" precision="1" seconds="46.03" value="-76.31279">76°18'46.03"W</geoCoordinate>
|
||
, 880-920 m, 29 Jul 2011 (fl),
|
||
<emphasis italics="true" pageId="11" pageNumber="44">
|
||
P.
|
||
<normalizedToken originalValue="Pedraza-Peñalosa">Pedraza-Penalosa</normalizedToken>
|
||
et al. 2447
|
||
</emphasis>
|
||
(COL!, MO!, NY!).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |