plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/4C/30/A4/4C30A452FFFCFFEDB9B0FC6B3D79F8CE.xml
maintenance b79ca1a6b7 added 4C
2024-06-21 12:35:51 +02:00

310 lines
54 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="2A1044F24D6FCAD2A0DB720A9F859D32" ID-CLB-Dataset="31806" ID-DOI="10.5281/zenodo.893555" ID-GBIF-Dataset="4a2efe36-c48b-456b-9223-7b5793025a72" ID-ISSN="0035-418" ID-Zenodo-Dep="893555" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1508361162557" checkinUser="plazi" docAuthor="Peter J. Schwendinger" docDate="2017" docId="4C30A452FFFCFFEDB9B0FC6B3D79F8CE" docLanguage="en" docName="rsz_124_2_11_Schwendinger.pdf" docOrigin="Revue suisse de Zoologie 124 (2)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Liphistius negara Peter J. Schwendinger 2017, sp. nov." docType="treatment" docVersion="9" lastPageNumber="417" masterDocId="B009DC2AFFE4FFF7BA6FFFF13818FFE3" masterDocTitle="A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1" masterLastPageNumber="445" masterPageNumber="391" pageNumber="415" updateTime="1698469770161" updateUser="ExternalLinkService">
<mods:mods id="38869FF647973CAA91EA9A57E81D1794" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EB88F07968038E9BAB2F8F42F06C9212">
<mods:title id="556B7701E1A93F1EDB85CE96A7D37EC3">A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1</mods:title>
</mods:titleInfo>
<mods:name id="D54BD92F0369EA920ED1CC372EF922AE" type="personal">
<mods:role id="B9134799AE3357A1F62A8DF20F846DA5">
<mods:roleTerm id="9BC24C786D31BFCE180C3E1398A4ED16">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3A5A1DF081818F6D56AF62F7689D22C">Peter J. Schwendinger</mods:namePart>
</mods:name>
<mods:typeOfResource id="C86ABDA1A5C225ED18BB06CB7D31D918">text</mods:typeOfResource>
<mods:relatedItem id="9550B81B5BADFD8734442BCF891F1C62" type="host">
<mods:titleInfo id="6FD7D273551188AB0D3E70E74AF68494">
<mods:title id="C7AE2817929B314A119F043FCD0091A5">Revue suisse de Zoologie</mods:title>
</mods:titleInfo>
<mods:part id="B8C30682A2019D509EA9D9488411B0EA">
<mods:date id="14B9FD9B92FAD9F36E7AC86DC76B2F80">2017</mods:date>
<mods:detail id="7D1FFA2ADD63CCEE5B43CBDC9D761E74" type="pubDate">
<mods:number id="F437A332E27E1A8546EB14DAE5832B0E">2017-09-30</mods:number>
</mods:detail>
<mods:detail id="8E21C24631A1F0793E3BF84308E68318" type="volume">
<mods:number id="78B3B60FBED04F2DFED91C1D881C59F7">124</mods:number>
</mods:detail>
<mods:detail id="6462CD99C8007BB0DFAFE373EACE88F1" type="issue">
<mods:number id="74F6F2353C0B3E74BB85D0D9D3E71202">2</mods:number>
</mods:detail>
<mods:extent id="44F634E9149573D6E5EE7B463D040207" unit="page">
<mods:start id="3302ABCC3A5F3CC962CBBC2C27014C35">391</mods:start>
<mods:end id="248B8ACD39D6CF479101F0FE5F67BE1A">445</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="25978AAC43D07F5A392E858854ED1892">journal article</mods:classification>
<mods:identifier id="D18F8037B4CB1ADB79757FC2576CAF88" type="CLB-Dataset">31806</mods:identifier>
<mods:identifier id="3D367D16A31F1B4257A8710D09E6AF5D" type="DOI">10.5281/zenodo.893555</mods:identifier>
<mods:identifier id="2E55F2A5C2EF821EDB7004063F834C43" type="GBIF-Dataset">4a2efe36-c48b-456b-9223-7b5793025a72</mods:identifier>
<mods:identifier id="C9AD16B044DA6E0E723B594DF6A3AD01" type="ISSN">0035-418</mods:identifier>
<mods:identifier id="8443391632AA975E070488F4A76C57D7" type="Zenodo-Dep">893555</mods:identifier>
</mods:mods>
<treatment id="4C30A452FFFCFFEDB9B0FC6B3D79F8CE" ID-DOI="http://doi.org/10.5281/zenodo.6042366" ID-GBIF-Taxon="135450771" ID-Zenodo-Dep="6042366" LSID="urn:lsid:plazi:treatment:4C30A452FFFCFFEDB9B0FC6B3D79F8CE" httpUri="http://treatment.plazi.org/id/4C30A452FFFCFFEDB9B0FC6B3D79F8CE" lastPageId="26" lastPageNumber="417" pageId="24" pageNumber="415">
<subSubSection id="8C8346CFFFFCFFEFB9B0FC6B3CE5FC50" box="[991,1277,922,947]" pageId="24" pageNumber="415" type="nomenclature">
<paragraph id="C4261544FFFCFFEFB9B0FC6B3CE5FC50" blockId="24.[991,1277,922,979]" box="[991,1277,922,947]" pageId="24" pageNumber="415">
<heading id="9F6EA228FFFCFFEFB9B0FC6B3CE5FC50" bold="true" box="[991,1277,922,947]" centered="true" fontSize="10" level="3" pageId="24" pageNumber="415" reason="4">
<emphasis id="F6EDC956FFFCFFEFB9B0FC6B3CE5FC50" bold="true" box="[991,1277,922,947]" pageId="24" pageNumber="415">
<taxonomicName id="03996EC7FFFCFFEFB9B0FC6B3CB9FC51" ID-CoL="72J7G" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[991,1185,922,946]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="415" phylum="Arthropoda" rank="species" species="negara" status="sp. nov.">
<emphasis id="F6EDC956FFFCFFEFB9B0FC6B3CB9FC51" bold="true" box="[991,1185,922,946]" italics="true" pageId="24" pageNumber="415">Liphistius negara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFFCFFEFBEC8FC6A3CE5FC50" box="[1191,1277,923,947]" pageId="24" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFFCFFEFBE27FC4A3C8CFC30" box="[1096,1172,955,979]" pageId="24" pageNumber="415" type="materials_examined">
<paragraph id="C4261544FFFCFFEFBE27FC4A3C8CFC30" blockId="24.[991,1277,922,979]" box="[1096,1172,955,979]" pageId="24" pageNumber="415">
<figureCitation id="5CA209C1FFFCFFEFBE27FC4A3C8CFC30" box="[1096,1172,955,979]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="24" pageNumber="415">Fig. 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFFCFFEFB950FC1A3C85FB40" pageId="24" pageNumber="415" type="materials_examined">
<paragraph id="C4261544FFFCFFEFB950FC1A3C85FB40" blockId="24.[831,1437,1002,1189]" pageId="24" pageNumber="415">
<materialsCitation id="74F11F19FFFCFFEFB950FC1A3C85FB40" ID-GBIF-Occurrence="1675315712" collectingDate="2004-06-16" collectingDateMax="2006-07-09" collectingDateMin="2004-06-16" collectionCode="MHNG" collectorName="P. J. Schwendinger." country="Malaysia" county="Taman Negara" elevation="110" latitude="4.3969445" location="Abai Waterfall" longLatPrecision="21" longitude="102.43055" municipality="Nusa Camp" pageId="24" pageNumber="415" specimenCount="3" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
<emphasis id="F6EDC956FFFCFFEFB950FC1A3B94FBE0" bold="true" box="[831,908,1003,1027]" pageId="24" pageNumber="415">
<typeStatus id="1B22ABE6FFFCFFEFB950FC1A3B9EFBE0" box="[831,902,1003,1027]" pageId="24" pageNumber="415">Types</typeStatus>
:
</emphasis>
<collectionCode id="A2888D81FFFCFFEFB9F4FC1A3BE8FBE0" box="[923,1008,1003,1027]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="24" pageNumber="415">MHNG</collectionCode>
, sample MAL-04/14; male
<typeStatus id="1B22ABE6FFFCFFEFBF51FC1B3D85FBE2" box="[1342,1437,1002,1025]" pageId="24" pageNumber="415" type="holotype">holotype</typeStatus>
(matured
<collectingDate id="A063CA6CFFFCFFEFB9C1FBFA3C3FFBC1" box="[942,1063,1035,1058]" pageId="24" pageNumber="415" value="2006-07-09">
<date id="B0273384FFFCFFEFB9C1FBFA3C3FFBC1" box="[942,1063,1035,1058]" pageId="24" pageNumber="415" value="2006-07-09">9.VII.2006</date>
</collectingDate>
),
<specimenCount id="D29FDECDFFFCFFEFBE2EFBFA3C88FBC1" box="[1089,1168,1035,1059]" pageId="24" pageNumber="415" type="male">1 male</specimenCount>
<typeStatus id="1B22ABE6FFFCFFEFBEF2FBFD3CE2FBC7" box="[1181,1274,1036,1060]" pageId="24" pageNumber="415" type="paratype">paratype</typeStatus>
(matured end of
<collectingDate id="A063CA6CFFFCFFEFB930FBDA3BD7FBA1" box="[863,975,1067,1090]" pageId="24" pageNumber="415" value="2005-07">
<date id="B0273384FFFCFFEFB930FBDA3BD7FBA1" box="[863,975,1067,1090]" pageId="24" pageNumber="415" value="2005-07">July 2005</date>
</collectingDate>
), female
<typeStatus id="1B22ABE6FFFCFFEFBE59FBDA3C97FBA1" box="[1078,1167,1067,1090]" pageId="24" pageNumber="415" type="allotype">allotype</typeStatus>
,
<specimenCount id="D29FDECDFFFCFFEFBEF2FBDD3CC4FBA1" box="[1181,1244,1067,1092]" pageId="24" pageNumber="415" type="generic">2 juv.</specimenCount>
males;
<collectingCountry id="BC8E55D4FFFCFFEFBF5BFBDA3D80FBA1" box="[1332,1432,1067,1090]" name="Malaysia" pageId="24" pageNumber="415">Malaysia</collectingCountry>
,
<collectingRegion id="065DDBA6FFFCFFEFB950FBBA3B88FB81" box="[831,912,1099,1122]" country="Malaysia" name="Pahang" pageId="24" pageNumber="415">Pahang</collectingRegion>
,
<collectingCounty id="2D476DC8FFFCFFEFB9F1FBBC3C58FB86" box="[926,1088,1101,1125]" pageId="24" pageNumber="415">Taman Negara</collectingCounty>
, trail from
<collectingMunicipality id="24428F3EFFFCFFEFBEAEFBBC3D5AFB87" box="[1217,1346,1100,1124]" pageId="24" pageNumber="415">Nusa Camp</collectingMunicipality>
to
<location id="C146439FFFFCFFEFBF07FBBA3BBEFB60" LSID="urn:lsid:plazi:treatment:4C30A452FFFCFFEDB9B0FC6B3D79F8CE:C146439FFFFCFFEFBF07FBBA3BBEFB60" country="Malaysia" county="Taman Negara" latitude="4.3969445" longLatPrecision="21" longitude="102.43055" municipality="Nusa Camp" name="Abai Waterfall" pageId="24" pageNumber="415" stateProvince="Pahang">Abai Waterfall</location>
,
<geoCoordinate id="A1AD7383FFFCFFEFB9D5FB9C3C2BFB66" box="[954,1075,1133,1157]" direction="north" orientation="latitude" pageId="24" pageNumber="415" precision="15" value="4.3969445">4°23’49”N</geoCoordinate>
,
<geoCoordinate id="A1AD7383FFFCFFEFBE28FB9D3CC1FB60" box="[1095,1241,1132,1155]" direction="east" orientation="longitude" pageId="24" pageNumber="415" precision="15" value="102.43055">102°25’50”E</geoCoordinate>
,
<elevation id="4FB4F277FFFCFFEFBE82FB9C3D24FB67" box="[1261,1340,1133,1157]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="24" pageNumber="415" unit="m" value="110.0">
<quantity id="0361B8A1FFFCFFEFBE82FB9C3D24FB67" box="[1261,1340,1133,1157]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="24" pageNumber="415" unit="m" value="110.0">110 m</quantity>
</elevation>
;
<collectingDate id="A063CA6CFFFCFFEFBF3FFB9D3B83FB46" pageId="24" pageNumber="415" value="2004-06-16" valueMax="2004-06-17" valueMin="2004-06-16">
<date id="B0273384FFFCFFEFBF3FFB9D3B83FB46" pageId="24" pageNumber="415" value="2004-06-16" valueMax="2004-06-17" valueMin="2004-06-16">16.-17. VI.2004</date>
</collectingDate>
; leg.
<collectorName id="696C7092FFFCFFEFB9BBFB7F3C85FB40" box="[980,1181,1164,1189]" pageId="24" pageNumber="415">P.J. Schwendinger.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFFCFFEFB950FB4C3CA6FAF7" pageId="24" pageNumber="415" type="etymology">
<paragraph id="C4261544FFFCFFEFB950FB4C3CA6FAF7" blockId="24.[831,1437,1212,1302]" pageId="24" pageNumber="415">
<emphasis id="F6EDC956FFFCFFEFB950FB4C3BDCFB36" bold="true" box="[831,964,1213,1237]" pageId="24" pageNumber="415">Etymology:</emphasis>
The species epithet, a name in apposition, is taken from the Malay name of the
<typeStatus id="1B22ABE6FFFCFFEFBF68FB2F3D2DFB15" box="[1287,1333,1246,1270]" pageId="24" pageNumber="415">type</typeStatus>
locality: taman (= park),
<taxonomicName id="03996EC7FFFCFFEFB983FB0F3C2BFAF5" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[1004,1075,1278,1302]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="415" phylum="Arthropoda" rank="species" species="negara">negara</taxonomicName>
(= national).
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFFCFFEFB950FADD3CF6F84D" pageId="24" pageNumber="415" type="diagnosis">
<paragraph id="C4261544FFFCFFEFB950FADD3CF6F84D" blockId="24.[831,1437,1324,1966]" pageId="24" pageNumber="415">
<emphasis id="F6EDC956FFFCFFEFB950FADD3BAFFAA0" bold="true" box="[831,951,1324,1347]" pageId="24" pageNumber="415">Diagnosis:</emphasis>
Large, uniformly dark-coloured species in both sexes. Similar to
<taxonomicName id="03996EC7FFFCFFEFBE26FABF3CC8FA87" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" box="[1097,1232,1357,1381]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="415" phylum="Arthropoda" rank="species" species="panching">
<emphasis id="F6EDC956FFFCFFEFBE26FABF3CC8FA87" box="[1097,1232,1357,1381]" italics="true" pageId="24" pageNumber="415">L. panching</emphasis>
</taxonomicName>
, distinguished by slightly larger size (carapace length of males 10.28- 11.30, width 8.70-10.04 versus 9.8 and 8.4, respectively, in male of
<taxonomicName id="03996EC7FFFCFFEFB9DBFA413C2CFA26" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" box="[948,1076,1454,1479]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="415" phylum="Arthropoda" rank="species" species="panching">
<emphasis id="F6EDC956FFFCFFEFB9DBFA413C2CFA26" box="[948,1076,1454,1479]" italics="true" pageId="24" pageNumber="415">L. panching</emphasis>
</taxonomicName>
, see
<bibRefCitation id="A00868B5FFFCFFEFBE04FA5F3B6CFA05" author="Sedgwick W. C. &amp; Platnick N. I." journalOrPublisher="Malayan Nature Journal" pageId="24" pageNumber="415" pagination="203 - 206" part="39" refString="Sedgwick W. C., Platnick N. I. 1986. On the male of Liphistius panching (Araneae, Mesothelae). Malayan Nature Journal 39: 203 - 206." title="On the male of Liphistius panching (Araneae, Mesothelae)" type="journal article" year="1986">Sedgwick &amp; Platnick (1986: 205)</bibRefCitation>
. Males with tibial apophysis in ventral view narrower at base, more distinctly set back from anterior margin of tibia (
<figureCitation id="5CA209C1FFFCFFEFBE6BF9E13C72F9CB" box="[1028,1130,1552,1576]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="24" pageNumber="415">Fig. 14A</figureCitation>
cf.
<bibRefCitation id="A00868B5FFFCFFEFBECDF9E13B61F9A4" author="Sedgwick W. C. &amp; Platnick N. I." journalOrPublisher="Malayan Nature Journal" pageId="24" pageNumber="415" pagination="203 - 206" part="39" refString="Sedgwick W. C., Platnick N. I. 1986. On the male of Liphistius panching (Araneae, Mesothelae). Malayan Nature Journal 39: 203 - 206." title="On the male of Liphistius panching (Araneae, Mesothelae)" type="journal article" year="1986">Sedgwick &amp; Platnick, 1986</bibRefCitation>
: fig. 3); paracymbium deeper (
<figureCitation id="5CA209C1FFFCFFEFBF6DF9C13D76F9AA" box="[1282,1390,1584,1609]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="24" pageNumber="415">Fig. 14B</figureCitation>
cf.
<bibRefCitation id="A00868B5FFFCFFEFB950F9A03C6DF98B" author="Sedgwick W. C. &amp; Platnick N. I." box="[831,1141,1617,1641]" journalOrPublisher="Malayan Nature Journal" pageId="24" pageNumber="415" pagination="203 - 206" part="39" refString="Sedgwick W. C., Platnick N. I. 1986. On the male of Liphistius panching (Araneae, Mesothelae). Malayan Nature Journal 39: 203 - 206." title="On the male of Liphistius panching (Araneae, Mesothelae)" type="journal article" year="1986">Sedgwick &amp; Platnick, 1986</bibRefCitation>
: fig. 2); ventral process of contrategulum narrower and more pointed (
<figureCitation id="5CA209C1FFFCFFEFBF70F9803D75F969" box="[1311,1389,1649,1674]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="24" pageNumber="415">Fig. 14</figureCitation>
C-D cf.
<figureCitation id="5CA209C1FFFCFFEFB908F9603BD2F949" box="[871,970,1681,1706]" captionStart="Fig. 10" captionStartId="18.[831,866,1881,1902]" captionTargetBox="[838,1435,1514,1833]" captionTargetId="figure@18.[829,1439,1504,1842]" captionTargetPageId="18" captionText="Fig. 10. Liphistius johore, female holotype (A); Liphistius panching, male from type locality (B). (A) Dorsal view of vulval plate (redrawn from Platnick &amp; Sedgwick, 1984: fig. 79) (arrows indicating anterolateral invaginations in margin of poreplate). (B) Distal view of left palpal organ (dorsal side up) (redrawn from Sedgwick &amp; Platnick, 1986: fig. 5)." httpUri="https://zenodo.org/record/1021957/files/figure.png" pageId="24" pageNumber="415">Fig. 10B</figureCitation>
and
<bibRefCitation id="A00868B5FFFCFFEFBE68F9603D59F94B" author="Sedgwick W. C. &amp; Platnick N. I." box="[1031,1345,1681,1706]" journalOrPublisher="Malayan Nature Journal" pageId="24" pageNumber="415" pagination="203 - 206" part="39" refString="Sedgwick W. C., Platnick N. I. 1986. On the male of Liphistius panching (Araneae, Mesothelae). Malayan Nature Journal 39: 203 - 206." title="On the male of Liphistius panching (Araneae, Mesothelae)" type="journal article" year="1986">Sedgwick &amp; Platnick, 1986</bibRefCitation>
: fig. 5); distal edge of contrategulum with a large triangular tooth, dorsal apex rounded instead of pointed (
<figureCitation id="5CA209C1FFFCFFEFBF1EF9233B40F8EF" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="24" pageNumber="415">Fig. 14</figureCitation>
C-D cf.
<figureCitation id="5CA209C1FFFCFFEFB9D3F9023C07F8EF" box="[956,1055,1779,1804]" captionStart="Fig. 10" captionStartId="18.[831,866,1881,1902]" captionTargetBox="[838,1435,1514,1833]" captionTargetId="figure@18.[829,1439,1504,1842]" captionTargetPageId="18" captionText="Fig. 10. Liphistius johore, female holotype (A); Liphistius panching, male from type locality (B). (A) Dorsal view of vulval plate (redrawn from Platnick &amp; Sedgwick, 1984: fig. 79) (arrows indicating anterolateral invaginations in margin of poreplate). (B) Distal view of left palpal organ (dorsal side up) (redrawn from Sedgwick &amp; Platnick, 1986: fig. 5)." httpUri="https://zenodo.org/record/1021957/files/figure.png" pageId="24" pageNumber="415">Fig. 10B</figureCitation>
and
<bibRefCitation id="A00868B5FFFCFFEFBE31F9023D81F8E9" author="Sedgwick W. C. &amp; Platnick N. I." box="[1118,1433,1779,1804]" journalOrPublisher="Malayan Nature Journal" pageId="24" pageNumber="415" pagination="203 - 206" part="39" refString="Sedgwick W. C., Platnick N. I. 1986. On the male of Liphistius panching (Araneae, Mesothelae). Malayan Nature Journal 39: 203 - 206." title="On the male of Liphistius panching (Araneae, Mesothelae)" type="journal article" year="1986">Sedgwick &amp; Platnick, 1986</bibRefCitation>
: fig. 5). Female distinguished from those of
<taxonomicName id="03996EC7FFFCFFEFBF74F8E43D84F8C9" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" box="[1307,1436,1811,1836]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="415" phylum="Arthropoda" rank="species" species="panching">
<emphasis id="F6EDC956FFFCFFEFBF74F8E43D84F8C9" box="[1307,1436,1811,1836]" italics="true" pageId="24" pageNumber="415">L. panching</emphasis>
</taxonomicName>
by vulval plate wider than long; poreplate anteriorly distinctly narrower than posteriorly instead of anteriorly wider than posteriorly (
<figureCitation id="5CA209C1FFFCFFEFBE33F8843CABF86E" box="[1116,1203,1909,1933]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="24" pageNumber="415">Fig. 14</figureCitation>
I-J cf.
<figureCitation id="5CA209C1FFFCFFEFBF61F8843D7DF86E" box="[1294,1381,1909,1933]" captionStart="Fig. 13" captionStartId="23.[151,186,1966,1987]" captionTargetBox="[158,1411,716,1915]" captionTargetId="figure@23.[148,1435,692,1941]" captionTargetPageId="23" captionText="Fig. 13. Liphistius panching, vulval plate of four female specimens from the type locality: specimen moulted 16. X. 2002 (A-B), moulted VIII. 2005 (C), built egg case 16. VII. 2001 (D-F), moulted 30. XII. 2005 (G-H). (A, C-D, G) Entire structure, dorsal view. (B, E, H) Same, ventral view. (F) Receptacular cluster, ventral and slightly posterior view. Scale lines 1.0 mm: (A-C, F-H; D-E)." httpUri="https://zenodo.org/record/1021963/files/figure.png" pageId="24" pageNumber="415">Fig. 13</figureCitation>
and
<bibRefCitation id="A00868B5FFFCFFEFB950F8643C70F84D" author="Platnick N. &amp; Sedgwick W. C." box="[831,1128,1941,1966]" journalOrPublisher="American Museum Novitates" pageId="24" pageNumber="415" pagination="1 - 31" part="2781" refString="Platnick N. I., Sedgwick W. C. 1984. A revision of the spider genus Liphistius (Araneae, Mesothelae). American Museum Novitates 2781: 1 - 31." title="A revision of the spider genus Liphistius (Araneae, Mesothelae)" type="journal article" year="1984">Platnick &amp; Sedgwick, 1984</bibRefCitation>
: figs 77-78).
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFFCFFEDB950F8343C62FB0D" lastPageId="26" lastPageNumber="417" pageId="24" pageNumber="415" type="description">
<paragraph id="C4261544FFFCFFEDB950F83438E4FE1F" blockId="24.[831,1437,1989,2078]" lastBlockId="26.[151,757,258,1934]" lastPageId="26" lastPageNumber="417" pageId="24" pageNumber="415">
<emphasis id="F6EDC956FFFCFFEFB950F8343CA5F83F" bold="true" box="[831,1213,1989,2013]" pageId="24" pageNumber="415">
Description of male (
<typeStatus id="1B22ABE6FFFCFFEFBE28F8343CA9F83F" box="[1095,1201,1989,2012]" pageId="24" pageNumber="415" type="holotype">holotype</typeStatus>
):
</emphasis>
<emphasis id="F6EDC956FFFCFFEFBEA3F8343D84F83F" box="[1228,1436,1989,2013]" italics="true" pageId="24" pageNumber="415">Colour in alcohol</emphasis>
(much darker in life): Most sclerotised parts uniformly chestnut-brown, on ventral side slightly lighter than on dorsal side; carapace reddish chestnut-brown; proximal portion of chelicerae cream-coloured; membranes of opisthosoma greyish cream, other membranes white.
<emphasis id="F6EDC956FFFEFFEDBAF8FE923999FE98" box="[151,385,355,379]" italics="true" pageId="26" pageNumber="417">Bristles on carapace</emphasis>
: Stiff bristles along all margins (longest anteriorly); many weaker and shorter ones on coxal elevations and behind fovea, fewer of them on both sides of eye mound and in a W-shaped arrangement behind it.
</paragraph>
<caption id="90E645CCFFFDFFEEBAF8F8AE3CC9F80B" httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="25" pageNumber="416" targetBox="[154,1436,260,1826]" targetPageId="25">
<paragraph id="C4261544FFFDFFEEBAF8F8AE3CC9F80B" blockId="25.[151,1437,1887,2024]" pageId="25" pageNumber="416">
Fig. 14.
<taxonomicName id="03996EC7FFFDFFEEBA82F8AE398EF896" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[237,406,1887,1909]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="416" phylum="Arthropoda" rank="species" species="negara" status="sp. nov.">
<emphasis id="F6EDC956FFFDFFEEBA82F8AE398EF896" box="[237,406,1887,1909]" italics="true" pageId="25" pageNumber="416">Liphistius negara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFFDFFEEBBF3F89139F2F896" box="[412,490,1888,1909]" pageId="25" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
, details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J).
</paragraph>
</caption>
<paragraph id="C4261544FFFEFFEDBAF8FDF43906FDBE" blockId="26.[151,757,258,1934]" pageId="26" pageNumber="417">
<emphasis id="F6EDC956FFFEFFEDBAF8FDF43954FDFF" box="[151,332,517,540]" italics="true" pageId="26" pageNumber="417">Cheliceral teeth</emphasis>
: Eleven and twelve small ones on promargin of cheliceral groove on right and left chelicera, respectively.
</paragraph>
<paragraph id="C4261544FFFEFFEDBAF8FD9739BFFCDC" blockId="26.[151,757,258,1934]" pageId="26" pageNumber="417">
<emphasis id="F6EDC956FFFEFFEDBAF8FD9738F5FD9E" box="[151,237,614,637]" italics="true" pageId="26" pageNumber="417">Scopula</emphasis>
: Tarsus I with thin scopula confined to distal third of ventral side, divided for its entire length by narrow pale, glabrous longitudinal median stripe and by some bristles; tarsus II with thin scopula in distal half, only distally divided by median stripe; tarsi III-IV with dense scopula covering distal four-fifths, only distally divided by median stripe.
</paragraph>
<paragraph id="C4261544FFFEFFEDBAF8FCB83A9CFC43" blockId="26.[151,757,258,1934]" pageId="26" pageNumber="417">
<emphasis id="F6EDC956FFFEFFEDBAF8FCB838C2FC83" box="[151,218,841,864]" italics="true" pageId="26" pageNumber="417">Claws</emphasis>
: Paired tarsal claws on anterior legs with 4-5 denticles, on posterior legs with 4-6 denticles; unpaired claws in most cases carrying a single denticle.
</paragraph>
<paragraph id="C4261544FFFEFFEDBAF8FC5B3978F9A9" blockId="26.[151,757,258,1934]" pageId="26" pageNumber="417">
<emphasis id="F6EDC956FFFEFFEDBAF8FC5B38D1FC22" box="[151,201,938,961]" italics="true" pageId="26" pageNumber="417">Palp</emphasis>
: Tibial apophysis slightly set back from distal margin of tibia (
<figureCitation id="5CA209C1FFFEFFEDBB24FC3B39A8FC00" box="[331,432,970,995]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14A</figureCitation>
), carrying four medium-sized pointed megaspines (
<figureCitation id="5CA209C1FFFEFFEDBBE9FC1B3A0DFBE0" box="[390,533,1002,1027]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14B, G</figureCitation>
). Both apical lobes of cymbium indistinct and equally rounded (
<figureCitation id="5CA209C1FFFEFFEDB8E3FBFA38B6FBA7" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14E, H</figureCitation>
). Paracymbium basally deep (
<figureCitation id="5CA209C1FFFEFFEDBB93FBDA3A78FBA7" box="[508,608,1067,1092]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14B</figureCitation>
), its cumulus indistinctly elevated, carrying moderately long stiff bristles (
<figureCitation id="5CA209C1FFFEFFEDBA96FB9D3954FB66" box="[249,332,1132,1157]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14</figureCitation>
A-B). Subtegulum without apophysis. Tegulum short and wide, coarsely dentate along entire proximal margin (
<figureCitation id="5CA209C1FFFEFFEDBB37FB5C39AEFB25" box="[344,438,1197,1222]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14F</figureCitation>
). Contrategulum with distinct conical ventral process, its rounded apex directed proventrad (
<figureCitation id="5CA209C1FFFEFFEDBB73FB1C3999FAE5" box="[284,385,1261,1286]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14C</figureCitation>
); distal edge with several oblique ribs and with a large triangular tooth prodorsally, ending in spatulate asymmetrical dorsal apex (
<figureCitation id="5CA209C1FFFEFFEDB82CFADF3ABFFAA4" box="[579,679,1326,1351]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14C</figureCitation>
). Paraembolic plate short (
<figureCitation id="5CA209C1FFFEFFEDBB16FABE39C5FA84" box="[377,477,1359,1383]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14A</figureCitation>
, E-F); retrolateral side of sclerotised part of embolus proper without longitudinal keels or ribs and not carrying denticles, dorsal wall much wider than ventral wall and sharply bent prodorsad, thus opening of spermophor forming an L (
<figureCitation id="5CA209C1FFFEFFEDB803FA2138B2F9EA" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14A, C, E</figureCitation>
); membranous part of embolus proper distally wider than proximally, with only slightly pigmented proximal portion (
<figureCitation id="5CA209C1FFFEFFEDBA9CF9C0394BF9A9" box="[243,339,1585,1610]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14E</figureCitation>
).
</paragraph>
<paragraph id="C4261544FFFEFFEDBAF8F9A23A85F86F" blockId="26.[151,757,258,1934]" pageId="26" pageNumber="417">
<emphasis id="F6EDC956FFFEFFEDBAF8F9A2392AF989" box="[151,306,1619,1642]" italics="true" pageId="26" pageNumber="417">Measurements</emphasis>
: Total length 27.94; carapace 11.30 long, 10.04 wide; opisthosoma 11.55 long, 9.04 wide; eye mound 1.51 long, 1.61 wide, AME well-developed; palpal coxae 3.68 long, 2.26 wide; labium 1.00 long, 2.01 wide; sternum 5.35 long, 3.01 wide (1.34 on ventral surface); palp 18.76 long (5.69 + 3.18 + 6.44 + 3.45); leg I 32.80 long (9.20 + 4.69 + 7.20 + 8.28 + 3.43); leg II 35.06 long (9.45 + 4.69 + 7.70 + 9.54 + 3.68); leg III 38.66 long (9.45 + 4.69 + 8.20 + 12.05 + 4.27); leg IV 47.53 long (11.55 + 4.85 + 9.79 + 15.48 + 5.86).
</paragraph>
<paragraph id="C4261544FFFEFFEDBAF8F8553B80FE7A" blockId="26.[151,756,1956,2079]" lastBlockId="26.[831,1437,258,1086]" pageId="26" pageNumber="417">
<emphasis id="F6EDC956FFFEFFEDBAF8F8553A05F858" bold="true" box="[151,541,1956,1979]" pageId="26" pageNumber="417">
Description of female (
<typeStatus id="1B22ABE6FFFEFFEDBBC0F8553A09F858" box="[431,529,1956,1979]" pageId="26" pageNumber="417" type="allotype">allotype</typeStatus>
):
</emphasis>
Colour in alcohol (much darker in life) as in male. Carapace with stiff bristles as in male, plus a pair of bristles anterior of fovea. Eleven strong teeth on promargin of cheliceral groove on both sides. Paired tarsal claws on anterior legs with three denticles, on posterior claws with 3-4 denticles; unpaired claws in most cases with a single denticle (claws worn on all limbs). Tarsi without scopula.
</paragraph>
<paragraph id="C4261544FFFEFFEDB950FE523BF6FD18" blockId="26.[831,1437,258,1086]" pageId="26" pageNumber="417">
<emphasis id="F6EDC956FFFEFFEDB950FE523B63FE59" box="[831,891,419,442]" italics="true" pageId="26" pageNumber="417">Vulva</emphasis>
(
<figureCitation id="5CA209C1FFFEFFEDB9E0FE523BF8FE58" box="[911,992,419,443]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14</figureCitation>
I-J): Posterior margin of genital sternite slightly and widely invaginated. Vulval plate distinctly sclerotised and pigmented, wider than long. Genital atrium with several lateral and median hairs. Posterior stalk wide, with micropores, fused with poreplate. Poreplate entirely and distinctly pigmented, its anterior margin indistinctly invaginated and clearly narrower than posterior margin. Two central dorsal openings (CDO) separated by a longitudinal bridge; two longitudinal receptacular clusters on ventral side completely separated from each other.
</paragraph>
<paragraph id="C4261544FFFEFFEDB950FCF43BD1FBDF" blockId="26.[831,1437,258,1086]" pageId="26" pageNumber="417">
<emphasis id="F6EDC956FFFEFFEDB950FCF43BC2FCFF" box="[831,986,773,796]" italics="true" pageId="26" pageNumber="417">Measurements</emphasis>
: Total length 22.00; carapace 9.62 long, 8.28 wide; opisthosoma 10.29 long, 8.62 wide; eye mound 1.22 long, 1.42 wide; palpal coxae 3.18 long, 2.01 wide; labium 1.17 long, 2.18 wide; sternum 4.35 long, 2.59 wide (1.59 on ventral surface); palp 16.23 long (5.35 + 3.01 + 4.02 + 3.85); leg I 19.76 long (5.86 + 3.60 + 4.27 + 4.02 + 2.01); leg II 20.57 long (6.02 + 3.68 + 4.35 + 4.43 + 2.09); leg III 22.25 long (6.19 + 3.68 + 4.52 + 5.35 + 2.51); leg IV 30.37 long (8.03 + 3.85 + 6.11 + 8.70 + 3.68).
</paragraph>
<paragraph id="C4261544FFFEFFEDB950FBA53C62FB0D" blockId="26.[831,1436,1108,1262]" pageId="26" pageNumber="417">
<emphasis id="F6EDC956FFFEFFEDB950FBA53BAEFB88" bold="true" box="[831,950,1108,1131]" pageId="26" pageNumber="417">Variation:</emphasis>
Carapace lengths in males (n=2) 10.28- 11.30, widths 8.70-10.04. The male
<typeStatus id="1B22ABE6FFFEFFEDBE8DFB873D27FB6D" box="[1250,1343,1142,1166]" pageId="26" pageNumber="417" type="paratype">paratype</typeStatus>
has the same reduced scopula on anterior legs (especially on tarsus I) as the
<typeStatus id="1B22ABE6FFFEFFEDB980FB443C48FB2F" box="[1007,1104,1205,1228]" pageId="26" pageNumber="417" type="holotype">holotype</typeStatus>
. Variation in the shape of the palpal organ, see
<figureCitation id="5CA209C1FFFEFFEDB995FB243C5DFB0D" box="[1018,1093,1237,1262]" captionStart="Fig. 14" captionStartId="25.[151,186,1887,1908]" captionTargetBox="[154,1436,260,1826]" captionTargetId="figure@25.[148,1450,252,1850]" captionTargetPageId="25" captionText="Fig. 14. Liphistius negara sp. nov., details of left palp of male holotype (A-C, E-H) and of male paratype (D); vulval plate of female allotype (I-J). (A) Distal part of palp, ventral view. (B) Distal part of tibia and base of tarsus, retroventral view. (C-D) Palpal organ, distal view (dorsal side up). (E) Same, proventral view. (F) Same, retrodorsal view. (G) Tibial apophysis, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I) Vulval plate, dorsal view. (J) Same, ventral view. Abbreviation: t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A; B, G-H; C-D; E-F; I-J)." httpUri="https://zenodo.org/record/1021965/files/figure.png" pageId="26" pageNumber="417">Fig. 14</figureCitation>
C-D.
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFFEFFEDB950FAF53D5AFADE" pageId="26" pageNumber="417" type="distribution">
<paragraph id="C4261544FFFEFFEDB950FAF53D5AFADE" blockId="26.[831,1436,1284,1341]" pageId="26" pageNumber="417">
<emphasis id="F6EDC956FFFEFFEDB950FAF53BCEFAF8" bold="true" box="[831,982,1284,1307]" pageId="26" pageNumber="417">Distribution:</emphasis>
Known only from the
<typeStatus id="1B22ABE6FFFEFFEDBE82FAF43D03FAFE" box="[1261,1307,1285,1309]" pageId="26" pageNumber="417">type</typeStatus>
locality in central peninsular
<collectingCountry id="BC8E55D4FFFEFFEDBE6BFAD53C7EFAD8" box="[1028,1126,1316,1339]" name="Malaysia" pageId="26" pageNumber="417">Malaysia</collectingCountry>
(
<figureCitation id="5CA209C1FFFEFFEDBE1AFAD53CACFADE" box="[1141,1204,1316,1341]" captionStart="Fig. 1" captionStartId="1.[151,186,1392,1413]" captionTargetBox="[153,758,625,1349]" captionTargetId="figure@1.[149,499,895,1170]" captionTargetPageId="1" captionText="Fig. 1. Localities of Liphistius species of the malayanus - group, tioman - group, linang - group and batuenis - group in peninsular Malaysia and southern Thailand (coast of Sumatra omitted): 1 - Gunung Angsi (type locality of L. malayanus); 2 - Templer Park and Gua Anak Takun (L. malayanus, L. batuensis); 3 - Genting Highlands (L. malayanus); 4 - Fraser’s Hill (L. malayanus); 5 - Cameron Highlands (type locality of L. malayanus cameroni); 6 - Sungai Jasin in Endau Rompin National Park (type locality of L. endau); 7 - Gunung Belumut (L. endau); 8 - Gunung Muntahak (L. endau; type locality of L. gracilis sp. nov.); 9 - Sungai Rengit (type locality of L. johore); 10 - Gunung Kajang on Tioman Island (type locality of L. tioman); 11 - Gua Charas in Bukit Charas (type locality of L. panching); 12 - Nusa Camp in Taman Negara (type locality of L. negara sp. nov.); 13 - Jeram Linang Waterfall (type locality of L. linang sp. nov.); 14 - Sankalakhierie Mountains (type locality of L. indra); 15 - Batu Caves (type locality of L. batuensis); 16 - Gua Tempurung (type locality of L. tempurung); 17 - Gua Cicak (L. tempurung); 18 - Gua Keris (type locality of L. priceae sp. nov). Localities with conspecific populations are encircled. Colours distinguish species groups." httpUri="https://zenodo.org/record/1021937/files/figure.png" pageId="26" pageNumber="417">Fig. 1</figureCitation>
, locality 12).
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFFEFFEDB950FAA53D79F8CE" pageId="26" pageNumber="417" type="biology_ecology">
<paragraph id="C4261544FFFEFFEDB950FAA53D53F94F" blockId="26.[831,1437,1364,1838]" pageId="26" pageNumber="417">
<emphasis id="F6EDC956FFFEFFEDB950FAA53B87FA88" bold="true" box="[831,927,1364,1387]" pageId="26" pageNumber="417">Biology:</emphasis>
The
<typeStatus id="1B22ABE6FFFEFFEDB98DFAA43C03FA8F" box="[994,1051,1365,1388]" pageId="26" pageNumber="417">types</typeStatus>
were collected from sloping soil and from decaying wood in a rain forest. The largest burrow (of juvenile male, not yet penultimate) had a
<quantity id="0361B8A1FFFEFFEDB93AFA453BBAFA2F" box="[853,930,1460,1484]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="26" pageNumber="417" unit="cm" value="2.6">2.6 cm</quantity>
long ad
<quantity id="0361B8A1FFFEFFEDBE67FA443C4CFA2F" box="[1032,1108,1461,1485]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="26" pageNumber="417" unit="cm" value="4.1">4.1 cm</quantity>
wide trapdoor. The trapdoors of old females thus must be of similar size as those of
<taxonomicName id="03996EC7FFFEFFEDB950FA073BC0F9E8" authorityName="Abraham" authorityYear="1923" box="[831,984,1524,1549]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="417" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis id="F6EDC956FFFEFFEDB950FA073BC0F9E8" box="[831,984,1524,1549]" italics="true" pageId="26" pageNumber="417">L. malayanus</emphasis>
</taxonomicName>
and
<taxonomicName id="03996EC7FFFEFFEDBE75FA073C67F9E8" authorityName="Sedgwick &amp; Platnick" authorityYear="1987" box="[1050,1151,1524,1549]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="417" phylum="Arthropoda" rank="species" species="endau">
<emphasis id="F6EDC956FFFEFFEDBE75FA073C67F9E8" box="[1050,1151,1524,1549]" italics="true" pageId="26" pageNumber="417">L. endau</emphasis>
</taxonomicName>
. The female
<typeStatus id="1B22ABE6FFFEFFEDBF4EFA053D60F9E8" box="[1313,1400,1524,1547]" pageId="26" pageNumber="417" type="allotype">allotype</typeStatus>
of
<taxonomicName id="03996EC7FFFEFFEDB950F9E73BB3F9CE" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[831,939,1558,1581]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="417" phylum="Arthropoda" rank="species" species="negara" status="sp. nov.">
<emphasis id="F6EDC956FFFEFFEDB950F9E73BB3F9CE" box="[831,939,1558,1581]" italics="true" pageId="26" pageNumber="417">L. negara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFFEFFEDB9DBF9E73C12F9CE" box="[948,1034,1558,1581]" pageId="26" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
appears to be mature (judging from the degree of sclerotisation of its vulval plate), but not yet fully grown. Most burrow entrances had eight signal lines, one had nine. Both males became mature in July, after one and two years of captivity, respectively.
</paragraph>
<paragraph id="C4261544FFFEFFEDB950F9443D79F8CE" blockId="26.[831,1437,1364,1838]" pageId="26" pageNumber="417">
The male
<typeStatus id="1B22ABE6FFFEFFEDB9ACF9473C07F92D" box="[963,1055,1718,1742]" pageId="26" pageNumber="417" type="paratype">paratype</typeStatus>
has a pronounced pathological outgrowth on the right cheliceral fang, and bite marks of parasitic
<emphasis id="F6EDC956FFFEFFEDB9CCF9063C4EF8EF" box="[931,1110,1781,1806]" italics="true" pageId="26" pageNumber="417">
Ljungia
<taxonomicName id="03996EC7FFFEFFEDB990F9043C4EF8EF" box="[1023,1110,1781,1804]" class="Arachnida" family="Laelapidae" genus="Ljunghia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="417" phylum="Arthropoda" rank="species" species="bristowi">bristowi</taxonomicName>
</emphasis>
mites on its carapace (many) and opisthosomal tergites (few), but not elsewhere.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink