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<mods:title id="CEF7E49D2237DF3894E164ADA67DFCC4">Phylogenetic reassessment and biogeography of the Ectateus generic group (Coleoptera: Tenebrionidae: Platynotina)</mods:title>
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<subSubSection id="8CB51DC61521BB4DFC74FC83FA8D9A58" box="[905,1322,852,874]" pageId="20" pageNumber="93" type="nomenclature">
<paragraph id="C4104E4D1521BB4DFC74FC83FA8D9A58" blockId="20.[905,1322,852,874]" box="[905,1322,852,874]" pageId="20" pageNumber="93">
<heading id="9F58F9211521BB4DFC74FC83FA8D9A58" box="[905,1322,852,874]" centered="true" fontSize="8" level="2" pageId="20" pageNumber="93" reason="8">
SUBGENUS
<taxonomicName id="03AF35CE1521BB4DFBF1FC82FA8D9A58" authority="KOCH, 1956" authorityName="Koch" authorityYear="1956" box="[1036,1322,852,874]" class="Insecta" family="Tenebrionidae" genus="Upembarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1521BB4DFBF1FC82FB309A5A" box="[1036,1175,853,874]" italics="true" pageId="20" pageNumber="93">UPEMBARUS</emphasis>
KOCH, 1956
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8CB51DC61521BB4DFCDBFCABFCDC9E5D" pageId="20" pageNumber="93" type="materials_examined">
<paragraph id="C4104E4D1521BB4DFCDBFCABFC479A82" blockId="20.[806,1421,891,944]" pageId="20" pageNumber="93">
<emphasis id="F6DB925F1521BB4DFCDBFCABFC119AA3" box="[806,950,892,913]" italics="true" pageId="20" pageNumber="93">
<typeStatus id="1B14F0EF1521BB4DFCDBFCABFCFD9AA3" box="[806,858,892,913]" pageId="20" pageNumber="93">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="03AF35CE1521BB4DFC43FCACFA859AA3" authority="Koch, 1956" authorityName="Koch" authorityYear="1956" box="[958,1314,891,913]" class="Insecta" family="Tenebrionidae" genus="Upembarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="93" phylum="Arthropoda" rank="species" species="saegeri">
<emphasis id="F6DB925F1521BB4DFC43FCACFB3F9AA3" box="[958,1176,891,913]" italics="true" pageId="20" pageNumber="93">Upembarus saegeri</emphasis>
Koch, 1956
</taxonomicName>
; by original designation.
</paragraph>
<paragraph id="C4104E4D1521BB4DFCDBFC0EFBBD9D3E" blockId="20.[806,1421,984,1036]" pageId="20" pageNumber="93">
<emphasis id="F6DB925F1521BB4DFCDBFC0EFBB09ADF" box="[806,1047,984,1006]" italics="true" pageId="20" pageNumber="93">Diagnostic characters:</emphasis>
<collectingRegion id="066B80AF1521BB4DFBE3FC0FFBE29ADF" box="[1054,1093,984,1005]" country="South Sudan" name="Lakes" pageId="20" pageNumber="93">See</collectingRegion>
diagnostic characters for subgenus
<taxonomicName id="03AF35CE1521BB4DFC8EFC20FBB49D3E" authorityName="Kamiński" authorityYear="2015" box="[883,1043,1015,1036]" class="Insecta" family="Tenebrionidae" genus="Pseudoselinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1521BB4DFC8EFC20FBB49D3E" box="[883,1043,1015,1036]" italics="true" pageId="20" pageNumber="93">Pseudoselinus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C4104E4D1521BB4DFCDBFBE2FC239D58" blockId="20.[806,1421,1077,1130]" pageId="20" pageNumber="93">
<emphasis id="F6DB925F1521BB4DFCDBFBE2FC1F9D78" box="[806,952,1077,1098]" italics="true" pageId="20" pageNumber="93">Distribution:</emphasis>
<collectingCountry id="BCB80EDD1521BB4DFC39FBE2FAF09D78" box="[964,1367,1077,1099]" name="Democratic Republic of the Congo" pageId="20" pageNumber="93">Democratic Republic of the Congo</collectingCountry>
and
<collectingCountry id="BCB80EDD1521BB4DFCDBFB83FCD89D58" box="[806,895,1108,1130]" name="Zambia" pageId="20" pageNumber="93">Zambia</collectingCountry>
.
</paragraph>
<paragraph id="C4104E4D1521BB4DFCDBFB44FC099C36" blockId="20.[806,1421,1170,1284]" pageId="20" pageNumber="93">
<emphasis id="F6DB925F1521BB4DFCDBFB44FB0D9D9A" box="[806,1194,1170,1192]" italics="true" pageId="20" pageNumber="93">Species composition (four species):</emphasis>
<taxonomicName id="03AF35CE1521BB4DFB4FFB45FC129DF5" authority="Koch, 1956" authorityName="Koch" authorityYear="1956" class="Insecta" family="Tenebrionidae" genus="Upembarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="93" phylum="Arthropoda" rank="species" species="saegeri">
<emphasis id="F6DB925F1521BB4DFB4FFB45FA2A9D9A" box="[1202,1421,1170,1192]" italics="true" pageId="20" pageNumber="93">Upembarus saegeri</emphasis>
Koch, 1956
</taxonomicName>
,
<taxonomicName id="03AF35CE1521BB4DFC35FB66FA2F9DF5" authority="Koch, 1956" authorityName="Koch" authorityYear="1956" box="[968,1416,1201,1223]" class="Insecta" family="Tenebrionidae" genus="Upembarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="93" phylum="Arthropoda" rank="species" species="sympatrius">
<emphasis id="F6DB925F1521BB4DFC35FB66FB4C9DF5" box="[968,1259,1201,1223]" italics="true" pageId="20" pageNumber="93">Upembarus sympatrius</emphasis>
Koch, 1956
</taxonomicName>
,
<taxonomicName id="03AF35CE1521BB4DFCDBFB18FB1A9DD7" authority="Koch, 1956" authorityName="Koch" authorityYear="1956" box="[806,1213,1231,1253]" class="Insecta" family="Tenebrionidae" genus="Upembarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="93" phylum="Arthropoda" rank="species" species="upembaensis">
<emphasis id="F6DB925F1521BB4DFCDBFB18FB9D9DD6" box="[806,1082,1231,1252]" italics="true" pageId="20" pageNumber="93">Upembarus upembaensis</emphasis>
Koch, 1956
</taxonomicName>
,
<taxonomicName id="03AF35CE1521BB4DFB34FB18FC0D9C36" authority="Koch, 1956" authorityName="Koch" authorityYear="1956" class="Insecta" family="Tenebrionidae" genus="Upembarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="93" phylum="Arthropoda" rank="species" species="wittei">
<emphasis id="F6DB925F1521BB4DFB34FB18FA2A9DD7" box="[1225,1421,1231,1253]" italics="true" pageId="20" pageNumber="93">Upembarus wittei</emphasis>
Koch, 1956
</taxonomicName>
.
</paragraph>
<paragraph id="C4104E4D1521BB4DFC3EFA97FAAE9C6A" blockId="20.[963,1289,1344,1368]" box="[963,1289,1344,1368]" pageId="20" pageNumber="93">ACKNOWLEDGEMENTS</paragraph>
<paragraph id="C4104E4D1521BB4DFCDBFAB8FCDC9E5D" blockId="20.[806,1422,1391,1903]" pageId="20" pageNumber="93">
The author is grateful to Dariusz Iwan (
<collectionCode id="A2BED6881521BB4DFB08FAB8FA969CB7" box="[1269,1329,1391,1413]" pageId="20" pageNumber="93">MIIZ</collectionCode>
) for the supervision during the PhD project. Additionally, for the valuable comments on the manuscript, the author thanks Joanna Ma˛ kol (Wrocław University of Environmental and Life Sciences), Grzegorz Pas´nik (Institue of Systematics and Evolution of Animals,
<collectionCode id="A2BED6881521BB4DFAEBF9DFFAEE9F2C" box="[1302,1353,1544,1566]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15187" name="Java Sugar Experimental Station" pageId="20" pageNumber="93" type="Herbarium">PAS</collectionCode>
), Rolf Aalbu (
<collectingRegion id="066B80AF1521BB4DFC8AF9F0FC419F0F" box="[887,998,1575,1597]" country="United States of America" name="California" pageId="20" pageNumber="93">California</collectingRegion>
Academy of Sciences), and the anonymous reviewers assigned by the Zoological Journal of the Linnean Society. The author would also like to thank the curators of the institutions cited above for the loan of specimens in their care. This research received support from the SYNTHESYS Project (ES-TAF- 3891; http://www.synthesys.info/), which is financed by European Community Research Infrastructure Action under the FP7 Capacities Programme at the Museo Museo Nacional de Ciencias Naturales (
<collectionCode id="A2BED6881521BB4DFCD3F88EFCC89E5D" box="[814,879,1881,1903]" pageId="20" pageNumber="93">CSIC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="8CB51DC61520BB4CFEA9FF0FFA28981F" pageId="21" pageNumber="94" type="key">
<paragraph id="C4104E4D1520BB4CFEA9FF0FFB5699DE" blockId="21.[181,1425,215,1838]" box="[340,1265,215,237]" pageId="21" pageNumber="94">
KEY
<collectingRegion id="066B80AF1520BB4CFE70FF0CFE0D99DE" box="[397,426,219,236]" country="South Sudan" name="Lakes" pageId="21" pageNumber="94">TO</collectingRegion>
THE AFROTROPICAL PLATYNOTOID
<taxonomicName id="03AF35CE1520BB4CFCC8FF0FFC7799DE" baseAuthorityName="Iwan" baseAuthorityYear="2002" box="[821,976,216,237]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="subTribe" subTribe="Platynotina">PLATYNOTINA</taxonomicName>
GENERA AND SUBGENERA
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FF29FA28981F" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">1. Pronotal disc without basal indentations. Ratio of the pronotal discs total width to its height &lt;5.0. Ratio of pronotal disc height to prothorax height&gt; 0.4 ................................................................................................. (2)</paragraph>
</subSubSection>
<subSubSection id="8CB51DC61520BB4CFF48FEE3FA379E1C" pageId="21" pageNumber="94" type="description">
<paragraph id="C4104E4D1520BB4CFF48FEE3FA289857" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
Pronotal disc with a pair of basal indentations. Ratio of the pronotal discs total width to its height&gt; 6.0. Ratio of pronotal disc height to prothorax height &lt;0.3 ..................................................
<emphasis id="F6DB925F1520BB4CFB85FE84FB769857" box="[1144,1233,339,357]" italics="true" pageId="21" pageNumber="94">Ectateus</emphasis>
generic group (6)
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FEB9FA3798B3" blockId="21.[181,1425,215,1838]" box="[181,1424,366,385]" pageId="21" pageNumber="94">
2. Anaepisternum and metaventrite long (
<figureCitation id="5C9452C81520BB4CFD76FEB9FD7198B3" box="[651,726,366,385]" captionStart="Figures 2226" captionStartId="25.[164,254,1238,1257]" captionTargetBox="[431,1174,199,1206]" captionTargetId="figure-0@25.[428,1177,196,1209]" captionTargetPageId="25" captionText="Figures 2226. Apex of elytra (22), epipleuron (23, 26), abdominal ventrites (24), and anaepisternum (25). Phymatoplata halsteadi (22); Anchophthalmus (Kochogaster) impressicollis (23); Anchophthalmops gridellii (24); Zidalus latipes (25); Selinus planus (26). Arrows indicate particular states of the characters used in the phylogenetic analysis and autapomorphies (ap)." figureDoi="http://doi.org/10.5281/zenodo.10831742" httpUri="https://zenodo.org/record/10831742/files/figure.png" pageId="21" pageNumber="94">Fig. 25</figureCitation>
). Wings present (reduced in
<taxonomicName id="03AF35CE1520BB4CFBF0FEB8FBD898B3" baseAuthorityName="Koch" baseAuthorityYear="1956" box="[1037,1151,366,385]" class="Insecta" family="Tenebrionidae" genus="Zidalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="exalatus">
<emphasis id="F6DB925F1520BB4CFBF0FEB8FBD898B3" box="[1037,1151,366,385]" italics="true" pageId="21" pageNumber="94">Z. exalatus</emphasis>
</taxonomicName>
and Z.
<taxonomicName id="03AF35CE1520BB4CFB2EFEB9FB5C98B2" baseAuthorityName="Ardoin" baseAuthorityYear="1965" box="[1235,1275,366,384]" class="Insecta" family="Tenebrionidae" genus="Zidalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="species" species="royi">
<emphasis id="F6DB925F1520BB4CFB2EFEB9FB5C98B2" box="[1235,1275,366,384]" italics="true" pageId="21" pageNumber="94">royi</emphasis>
</taxonomicName>
) .......
<taxonomicName id="03AF35CE1520BB4CFAC2FEB9FA3798B3" authorityName="Mulsant &amp; Rey" authorityYear="1853" box="[1343,1424,366,385]" class="Insecta" family="Tenebrionidae" genus="Zidalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFAC2FEB9FA3798B3" box="[1343,1424,366,385]" italics="true" pageId="21" pageNumber="94">Zidalus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FE5FFA3798AF" blockId="21.[181,1425,215,1838]" box="[181,1424,392,413]" pageId="21" pageNumber="94">
Anaepisternum and metaventrite reduced. Wings absent...................................
<taxonomicName id="03AF35CE1520BB4CFBA5FE5DFB7698AF" authorityName="Koch" authorityYear="1956" box="[1112,1233,394,413]" class="Insecta" family="Tenebrionidae" genus="Upembarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFBA5FE5DFB7698AF" box="[1112,1233,394,413]" italics="true" pageId="21" pageNumber="94">Upembarus</emphasis>
</taxonomicName>
generic group (3)
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FE71FA28988B" blockId="21.[181,1425,215,1838]" box="[181,1423,422,441]" pageId="21" pageNumber="94">
3. Ventral margin of eyes with a groove below (
<figureCitation id="5C9452C81520BB4CFD3AFE71FCB7988B" box="[711,784,422,441]" captionStart="Figures 1115" captionStartId="23.[164,254,1326,1345]" captionTargetBox="[390,1218,197,1294]" captionTargetId="figure-0@23.[387,1220,195,1296]" captionTargetPageId="23" captionText="Figures 1115. Head dorsal view (11), antenna (12), and pronotum (1315). Anchophthalmops gridellii (12A), Ectateus ghesquierei (15); Glyptopteryx interioris (13); Lechius abacoides (11); Phymatoplata halsteadi (12B); Selinus planus (14). Arrows indicate particular states of the characters used in the phylogenetic analysis and autapomorphies (ap)." figureDoi="http://doi.org/10.5281/zenodo.10831738" httpUri="https://zenodo.org/record/10831738/files/figure.png" pageId="21" pageNumber="94">Fig. 11</figureCitation>
). Distribution:
<collectingCountry id="BCB80EDD1520BB4CFC49FE71FB94988B" box="[948,1075,422,441]" name="Madagascar" pageId="21" pageNumber="94">Madagascar</collectingCountry>
.................................
<taxonomicName id="03AF35CE1520BB4CFAC2FE71FA28988B" authorityName="Iwan" authorityYear="1995" box="[1343,1423,422,441]" class="Insecta" family="Tenebrionidae" genus="Lechius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFAC2FE71FA28988B" box="[1343,1423,422,441]" italics="true" pageId="21" pageNumber="94">Lechius</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FE17FA2898E7" blockId="21.[181,1425,215,1838]" box="[181,1423,448,469]" pageId="21" pageNumber="94">Ventral margin of eyes without a groove below. Distribution: continental Africa........................................(4)</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FE09FA289B3F" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
4. Middle of mentum not projecting anteriad. Anterior tentorial pits invisible. Intercoxal process of prosternum narrow.............................................................................................................................
<taxonomicName id="03AF35CE1520BB4CFAEEFE2DFA289B3F" authorityName="KAMINSKI" authorityYear="2013" box="[1299,1423,506,525]" class="Insecta" family="Tenebrionidae" genus="Paraselinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFAEEFE2DFA289B3F" box="[1299,1423,506,525]" italics="true" pageId="21" pageNumber="94">Paraselinus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FDC3FA379B76" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
Middle part of mentum projecting anteriad (
<figureCitation id="5C9452C81520BB4CFD48FDC2FD559B1B" box="[693,754,533,553]" captionStart="Figures 410" captionStartId="22.[144,236,1361,1380]" captionTargetBox="[369,1198,199,1328]" captionTargetId="figure-0@22.[367,1200,196,1331]" captionTargetPageId="22" captionText="Figures 410. Head ventral view (4, 6), mentum (5, 7, 8), and anterior tentorial pits (9, 10). Anchophthalmus (Anchophthalmus) algoensis (4); Anchophthalmus (Kochogaster) impressicollis (5); Angolositus lidiae (8); Nesopatrum josephi (6); Selinus planus (10); Upembarus (Pseudoselinus) punctatostriatus (7, 9). Arrows indicate particular states of the characters used in the phylogenetic analysis, autapomorphies (ap), and diagnostic characters (d)." figureDoi="http://doi.org/10.5281/zenodo.10831736" httpUri="https://zenodo.org/record/10831736/files/figure.png" pageId="21" pageNumber="94">Fig. 7</figureCitation>
). Anterior tentorial pits longitudinal (
<figureCitation id="5C9452C81520BB4CFB7EFDC2FB189B1B" box="[1155,1215,533,553]" captionStart="Figures 410" captionStartId="22.[144,236,1361,1380]" captionTargetBox="[369,1198,199,1328]" captionTargetId="figure-0@22.[367,1200,196,1331]" captionTargetPageId="22" captionText="Figures 410. Head ventral view (4, 6), mentum (5, 7, 8), and anterior tentorial pits (9, 10). Anchophthalmus (Anchophthalmus) algoensis (4); Anchophthalmus (Kochogaster) impressicollis (5); Angolositus lidiae (8); Nesopatrum josephi (6); Selinus planus (10); Upembarus (Pseudoselinus) punctatostriatus (7, 9). Arrows indicate particular states of the characters used in the phylogenetic analysis, autapomorphies (ap), and diagnostic characters (d)." figureDoi="http://doi.org/10.5281/zenodo.10831736" httpUri="https://zenodo.org/record/10831736/files/figure.png" pageId="21" pageNumber="94">Fig. 9</figureCitation>
). Intercoxal process of prosternum wide (
<figureCitation id="5C9452C81520BB4CFE3FFDE6FDAD9B76" box="[450,522,561,580]" captionStart="Figures 1621" captionStartId="24.[144,235,1318,1337]" captionTargetBox="[369,1198,197,1287]" captionTargetId="figure-0@24.[367,1200,196,1289]" captionTargetPageId="24" captionText="Figures 1621. Intercoxal process of prosternum (17), hypomeron (16), scutellum (18), and base of elytron (1921). Anchophthalmus (Kochogaster) impressicollis (16); Upembarus (Pseudoselinus) punctatostriatus (17); Anchophthalmops platessa (18); Ectateus modestus (19); Glyptopteryx lesnei (20); Eleoselinus villiersi (21). Arrows indicate particular states of the characters used in the phylogenetic analysis." figureDoi="http://doi.org/10.5281/zenodo.10831740" httpUri="https://zenodo.org/record/10831740/files/figure.png" pageId="21" pageNumber="94">Fig. 17</figureCitation>
)............................................................................................
<taxonomicName id="03AF35CE1520BB4CFB0FFDE6FACC9B76" authorityName="Koch" authorityYear="1956" box="[1266,1387,561,580]" class="Insecta" family="Tenebrionidae" genus="Upembarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFB0FFDE6FACC9B76" box="[1266,1387,561,580]" italics="true" pageId="21" pageNumber="94">Upembarus</emphasis>
</taxonomicName>
(5)
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FD9AFA379B4E" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
5. The upper edge of the elytral base disappearing before humerus (cf. Kamin´ ski, 2013c). Fifth abdominal ventrite without bordering. Bursa copulatrix without sclerites..................................................subgenus
<taxonomicName id="03AF35CE1520BB4CFAEAFDBEFA379B4E" authorityName="Koch" authorityYear="1956" box="[1303,1424,617,636]" class="Insecta" family="Tenebrionidae" genus="Upembarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFAEAFDBEFA379B4E" box="[1303,1424,617,636]" italics="true" pageId="21" pageNumber="94">Upembarus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FD54FA379B86" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
The upper edge of the elytral base reaching the humerus. Fifth abdominal ventrite bordered. Bursa copulatrix with longitudinal sclerites...................................................................................subgenus
<taxonomicName id="03AF35CE1520BB4CFB00FD76FA379B86" authorityName="Kamiński" authorityYear="2015" box="[1277,1424,673,692]" class="Insecta" family="Tenebrionidae" genus="Pseudoselinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFB00FD76FA379B86" box="[1277,1424,673,692]" italics="true" pageId="21" pageNumber="94">Pseudoselinus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FD6AFA379BE2" blockId="21.[181,1425,215,1838]" box="[181,1424,701,720]" pageId="21" pageNumber="94">
6. Anaepisternum and metaventrite long. Wings reduced (but present) (
<figureCitation id="5C9452C81520BB4CFC37FD6AFBB49BE2" box="[970,1043,701,720]" captionStart="Figure 30" captionStartId="25.[826,905,1882,1901]" captionTargetBox="[831,1027,1404,1847]" captionTargetId="figure-88@25.[826,1032,1399,1852]" captionTargetPageId="25" captionText="Figure 30. Reduced wing of Pteroselinus insularis." figureDoi="http://doi.org/10.5281/zenodo.10831746" httpUri="https://zenodo.org/record/10831746/files/figure.png" pageId="21" pageNumber="94">Fig. 30</figureCitation>
) ..............
<emphasis id="F6DB925F1520BB4CFB73FD6AFA379BE2" bold="true" box="[1166,1424,701,720]" pageId="21" pageNumber="94">
<taxonomicName id="03AF35CE1520BB4CFB73FD6AFAB99BE2" authority="Kamiński, 2015" authorityName="Kamiński" authorityYear="2015" box="[1166,1310,701,720]" class="Insecta" family="Lucanidae" genus="Pteroselinus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F6DB925F1520BB4CFB73FD6AFAB99BE2" bold="true" box="[1166,1310,701,720]" italics="true" pageId="21" pageNumber="94">Pteroselinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDE82F241520BB4CFADAFD6AFA379BE2" box="[1319,1424,701,720]" pageId="21" pageNumber="94" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FD00FA289BDE" blockId="21.[181,1425,215,1838]" box="[181,1423,727,748]" pageId="21" pageNumber="94">Anaepisternum and metaventrite short. Wings absent..........................................................................(7)</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FD22FA379A3A" blockId="21.[181,1425,215,1838]" box="[181,1424,757,776]" pageId="21" pageNumber="94">
7. Anterior tentorial pits deep, clearly visible as circular depressions (
<figureCitation id="5C9452C81520BB4CFC45FD22FBA79A3A" box="[952,1024,757,776]" captionStart="Figures 410" captionStartId="22.[144,236,1361,1380]" captionTargetBox="[369,1198,199,1328]" captionTargetId="figure-0@22.[367,1200,196,1331]" captionTargetPageId="22" captionText="Figures 410. Head ventral view (4, 6), mentum (5, 7, 8), and anterior tentorial pits (9, 10). Anchophthalmus (Anchophthalmus) algoensis (4); Anchophthalmus (Kochogaster) impressicollis (5); Angolositus lidiae (8); Nesopatrum josephi (6); Selinus planus (10); Upembarus (Pseudoselinus) punctatostriatus (7, 9). Arrows indicate particular states of the characters used in the phylogenetic analysis, autapomorphies (ap), and diagnostic characters (d)." figureDoi="http://doi.org/10.5281/zenodo.10831736" httpUri="https://zenodo.org/record/10831736/files/figure.png" pageId="21" pageNumber="94">Fig. 10</figureCitation>
) ............................................. (8)
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FCD8FA379A16" blockId="21.[181,1425,215,1838]" box="[181,1424,783,804]" pageId="21" pageNumber="94">Anterior tentorial pits invisible or visible as longitudinal depressions ..................................................... (9)</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FCFAFA379A6E" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
8. Pronotum widest at the base, sides evenly narrowing towards apex. Fifth abdominal ventrite bordered. Penis narrow; clavae developing from two-thirds of the aedeagal tegmen length (cf. Kamin´ ski, 2014) .............
<taxonomicName id="03AF35CE1520BB4CFABFFC9EFA379A6E" authorityName="Mulsant &amp; Rey" authorityYear="1853" box="[1346,1424,841,860]" class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFABFFC9EFA379A6E" box="[1346,1424,841,860]" italics="true" pageId="21" pageNumber="94">Selinus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FCB4FA379AA6" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
Pronotum widest in the middle, sides rounded. Fifth abdominal without bordering. Penis wide; clavae developing from half of the aedeagal tegmen length (cf. Kamin´ ski, 2014) ......................................................
<taxonomicName id="03AF35CE1520BB4CFAD4FC56FA379AA6" authorityName="Koch" authorityYear="1956" box="[1321,1424,897,916]" class="Insecta" family="Tenebrionidae" genus="Monodius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFAD4FC56FA379AA6" box="[1321,1424,897,916]" italics="true" pageId="21" pageNumber="94">Monodius</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FC4AFA379AF9" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
9. Dorsal side of frons with conspicuous depressions (
<figureCitation id="5C9452C81520BB4CFCCCFC4BFCD59A9D" box="[817,882,924,943]" captionStart="Figures 410" captionStartId="22.[144,236,1361,1380]" captionTargetBox="[369,1198,199,1328]" captionTargetId="figure-0@22.[367,1200,196,1331]" captionTargetPageId="22" captionText="Figures 410. Head ventral view (4, 6), mentum (5, 7, 8), and anterior tentorial pits (9, 10). Anchophthalmus (Anchophthalmus) algoensis (4); Anchophthalmus (Kochogaster) impressicollis (5); Angolositus lidiae (8); Nesopatrum josephi (6); Selinus planus (10); Upembarus (Pseudoselinus) punctatostriatus (7, 9). Arrows indicate particular states of the characters used in the phylogenetic analysis, autapomorphies (ap), and diagnostic characters (d)." figureDoi="http://doi.org/10.5281/zenodo.10831736" httpUri="https://zenodo.org/record/10831736/files/figure.png" pageId="21" pageNumber="94">Fig. 6</figureCitation>
). Bursa copulatrix with several small sclerites (
<figureCitation id="5C9452C81520BB4CFF14FC6FFE959AFE" box="[233,306,952,972]" captionStart="Figures 2729" captionStartId="25.[164,252,1765,1784]" captionTargetBox="[231,714,1432,1733]" captionTargetId="figure-51@25.[229,716,1430,1735]" captionTargetPageId="25" captionText="Figures 2729. Bursa copulatrix (27) and aedeagal tegmen (28, 29). Nesopatrum josephi (27); Anchophthalmops platessa (28); Phallocentrion edentatum (29). Arrows indicate particular states of the characters used in the phylogenetic analysis and autapomorphies (ap)." figureDoi="http://doi.org/10.5281/zenodo.10831744" httpUri="https://zenodo.org/record/10831744/files/figure.png" pageId="21" pageNumber="94">Fig. 27</figureCitation>
)...........................................................................................................................
<taxonomicName id="03AF35CE1520BB4CFAEDFC6EFA379AF9" authorityName="Gebien" authorityYear="1920" box="[1296,1424,953,971]" class="Insecta" family="Tenebrionidae" genus="Nesopatrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFAEDFC6EFA379AF9" box="[1296,1424,953,971]" italics="true" pageId="21" pageNumber="94">Nesopatrum</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FC04FA289AD5" blockId="21.[181,1425,215,1838]" box="[181,1423,979,1000]" pageId="21" pageNumber="94">Dorsal side of frons without depressions. Bursa copulatrix with a single sclerite or without sclerites.......... (10)</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FC27FA289D2D" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
10. Submentum pentagonal (
<figureCitation id="5C9452C81520BB4CFE0DFC27FD8A9D31" box="[496,557,1008,1027]" captionStart="Figures 410" captionStartId="22.[144,236,1361,1380]" captionTargetBox="[369,1198,199,1328]" captionTargetId="figure-0@22.[367,1200,196,1331]" captionTargetPageId="22" captionText="Figures 410. Head ventral view (4, 6), mentum (5, 7, 8), and anterior tentorial pits (9, 10). Anchophthalmus (Anchophthalmus) algoensis (4); Anchophthalmus (Kochogaster) impressicollis (5); Angolositus lidiae (8); Nesopatrum josephi (6); Selinus planus (10); Upembarus (Pseudoselinus) punctatostriatus (7, 9). Arrows indicate particular states of the characters used in the phylogenetic analysis, autapomorphies (ap), and diagnostic characters (d)." figureDoi="http://doi.org/10.5281/zenodo.10831736" httpUri="https://zenodo.org/record/10831736/files/figure.png" pageId="21" pageNumber="94">Fig. 5</figureCitation>
). Last segment of maxillary palpi strongly widened (
<figureCitation id="5C9452C81520BB4CFB94FC27FB009D31" box="[1129,1191,1008,1027]" captionStart="Figures 410" captionStartId="22.[144,236,1361,1380]" captionTargetBox="[369,1198,199,1328]" captionTargetId="figure-0@22.[367,1200,196,1331]" captionTargetPageId="22" captionText="Figures 410. Head ventral view (4, 6), mentum (5, 7, 8), and anterior tentorial pits (9, 10). Anchophthalmus (Anchophthalmus) algoensis (4); Anchophthalmus (Kochogaster) impressicollis (5); Angolositus lidiae (8); Nesopatrum josephi (6); Selinus planus (10); Upembarus (Pseudoselinus) punctatostriatus (7, 9). Arrows indicate particular states of the characters used in the phylogenetic analysis, autapomorphies (ap), and diagnostic characters (d)." figureDoi="http://doi.org/10.5281/zenodo.10831736" httpUri="https://zenodo.org/record/10831736/files/figure.png" pageId="21" pageNumber="94">Fig. 4</figureCitation>
)...
<taxonomicName id="03AF35CE1520BB4CFB38FC27FAD09D31" authorityName="Gerstaecker" authorityYear="1854" box="[1221,1399,1008,1027]" class="Insecta" family="Tenebrionidae" genus="Anchophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFB38FC27FAD09D31" box="[1221,1399,1008,1027]" italics="true" pageId="21" pageNumber="94">Anchophthalmus</emphasis>
</taxonomicName>
... .................................................................................................................................................(11)
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FBF1FA289D09" blockId="21.[181,1425,215,1838]" box="[181,1423,1062,1084]" pageId="21" pageNumber="94">Submentum triangular. Last segment of maxillary palpi narrow...........................................................(12)</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FB93FA289D41" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
11. Lateral wings of mentum developing at the level of two-thirds of mentum length (
<figureCitation id="5C9452C81520BB4CFBC4FB93FBD29D65" box="[1081,1141,1092,1111]" captionStart="Figures 410" captionStartId="22.[144,236,1361,1380]" captionTargetBox="[369,1198,199,1328]" captionTargetId="figure-0@22.[367,1200,196,1331]" captionTargetPageId="22" captionText="Figures 410. Head ventral view (4, 6), mentum (5, 7, 8), and anterior tentorial pits (9, 10). Anchophthalmus (Anchophthalmus) algoensis (4); Anchophthalmus (Kochogaster) impressicollis (5); Angolositus lidiae (8); Nesopatrum josephi (6); Selinus planus (10); Upembarus (Pseudoselinus) punctatostriatus (7, 9). Arrows indicate particular states of the characters used in the phylogenetic analysis, autapomorphies (ap), and diagnostic characters (d)." figureDoi="http://doi.org/10.5281/zenodo.10831736" httpUri="https://zenodo.org/record/10831736/files/figure.png" pageId="21" pageNumber="94">Fig. 5</figureCitation>
). Base of the fourth elytral interval protruding towards the front of the body (cf. Kamin´ ski &amp;
<bibRefCitation id="A03E33BC1520BB4CFC52FBB7FB949D41" author="Iwan D" box="[943,1075,1120,1139]" pageId="21" pageNumber="94" pagination="3669" refId="ref23029" refString="Kamin' ski MJ, Iwan D. 2013 a. Taxonomy of the genus Kochogaster Kamin'ski et Ras', 2011 (Coleoptera: Tenebrionidae: Pedinini), with description of a second known species. Zootaxa 3669: 027 - 036." type="journal article" year="2013">Iwan, 2013a</bibRefCitation>
) ............. subgenus
<taxonomicName id="03AF35CE1520BB4CFAECFBB7FA289D41" authorityName="Kamiński" authorityYear="2015" box="[1297,1423,1120,1139]" class="Insecta" family="Tenebrionidae" genus="Anchophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="subGenus" subGenus="Kochogaster">
<emphasis id="F6DB925F1520BB4CFAECFBB7FA289D41" box="[1297,1423,1120,1139]" italics="true" pageId="21" pageNumber="94">Kochogaster</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FBADFA289D99" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
Lateral wings of mentum visible from the base. Base of the fourth elytral interval not protruding towards the front of the body............................................................................................subgenus
<taxonomicName id="03AF35CE1520BB4CFB20FB4FFA289D99" authorityName="Gerstaecker" authorityYear="1854" box="[1245,1423,1176,1195]" class="Insecta" family="Tenebrionidae" genus="Anchophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFB20FB4FFA289D99" box="[1245,1423,1176,1195]" italics="true" pageId="21" pageNumber="94">Anchophthalmus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FB63FA379DD1" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
12. Pronotal disc and elytra densely covered with setae. Elytral humeri strongly protruding sideways (cf.
<bibRefCitation id="A03E33BC1520BB4CFAFEFB63FA219DF5" author="Iwan D" box="[1283,1414,1204,1223]" pageId="21" pageNumber="94" pagination="53 - 63" refId="ref22208" refString="Iwan D. 2001 a. A revision of Phallocentrion Koch, 1956 from Africa (Coleoptera: Tenebrionidae: Platynotini). Annales Zoologici 51: 53 - 63." type="journal article" year="2001">Iwan, 2001a</bibRefCitation>
). Clavae developing from half of the aedeagal tegmen length (
<figureCitation id="5C9452C81520BB4CFC93FB07FC109DD1" box="[878,951,1232,1251]" captionStart="Figures 2729" captionStartId="25.[164,252,1765,1784]" captionTargetBox="[231,714,1432,1733]" captionTargetId="figure-51@25.[229,716,1430,1735]" captionTargetPageId="25" captionText="Figures 2729. Bursa copulatrix (27) and aedeagal tegmen (28, 29). Nesopatrum josephi (27); Anchophthalmops platessa (28); Phallocentrion edentatum (29). Arrows indicate particular states of the characters used in the phylogenetic analysis and autapomorphies (ap)." figureDoi="http://doi.org/10.5281/zenodo.10831744" httpUri="https://zenodo.org/record/10831744/files/figure.png" pageId="21" pageNumber="94">Fig. 29</figureCitation>
).......................................
<taxonomicName id="03AF35CE1520BB4CFB0AFB07FA379DD1" authorityName="Koch" authorityYear="1956" box="[1271,1424,1232,1251]" class="Insecta" family="Tenebrionidae" genus="Phallocentrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFB0AFB07FA379DD1" box="[1271,1424,1232,1251]" italics="true" pageId="21" pageNumber="94">Phallocentrion</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FB3DFA379C05" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">Pronotal disc and elytra densely glabrous or covered with fine and sparsely distributed setae. Elytral humeri not protruding outwards. Clavae developing from two-thirds of the aedeagal tegmen length ................................ .................................................................................................................................................(13)</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FA97FA289C61" blockId="21.[181,1425,215,1838]" box="[181,1423,1344,1363]" pageId="21" pageNumber="94">
13. Apical part of mentum with a median indentation (
<figureCitation id="5C9452C81520BB4CFCF0FA97FCEF9C61" box="[781,840,1344,1363]" captionStart="Figures 410" captionStartId="22.[144,236,1361,1380]" captionTargetBox="[369,1198,199,1328]" captionTargetId="figure-0@22.[367,1200,196,1331]" captionTargetPageId="22" captionText="Figures 410. Head ventral view (4, 6), mentum (5, 7, 8), and anterior tentorial pits (9, 10). Anchophthalmus (Anchophthalmus) algoensis (4); Anchophthalmus (Kochogaster) impressicollis (5); Angolositus lidiae (8); Nesopatrum josephi (6); Selinus planus (10); Upembarus (Pseudoselinus) punctatostriatus (7, 9). Arrows indicate particular states of the characters used in the phylogenetic analysis, autapomorphies (ap), and diagnostic characters (d)." figureDoi="http://doi.org/10.5281/zenodo.10831736" httpUri="https://zenodo.org/record/10831736/files/figure.png" pageId="21" pageNumber="94">Fig. 8</figureCitation>
).........................................................
<taxonomicName id="03AF35CE1520BB4CFAEBFA97FA289C61" authorityName="Koch" authorityYear="1954" box="[1302,1423,1344,1363]" class="Insecta" family="Tenebrionidae" genus="Angolositus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFAEBFA97FA289C61" box="[1302,1423,1344,1363]" italics="true" pageId="21" pageNumber="94">Angolositus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FA8DFA289C5D" blockId="21.[181,1425,215,1838]" box="[181,1423,1370,1391]" pageId="21" pageNumber="94">Apical part of mentum straight.......................................................................................................(14)</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FAAFFA289C95" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
14. Third antennomere short (ratio of its length to the length of second antennomere ≥ 2.0). Elytral humeri strongly protruding towards body apex (
<figureCitation id="5C9452C81520BB4CFDCEFA43FDDB9C95" box="[563,636,1428,1447]" captionStart="Figures 1621" captionStartId="24.[144,235,1318,1337]" captionTargetBox="[369,1198,197,1287]" captionTargetId="figure-0@24.[367,1200,196,1289]" captionTargetPageId="24" captionText="Figures 1621. Intercoxal process of prosternum (17), hypomeron (16), scutellum (18), and base of elytron (1921). Anchophthalmus (Kochogaster) impressicollis (16); Upembarus (Pseudoselinus) punctatostriatus (17); Anchophthalmops platessa (18); Ectateus modestus (19); Glyptopteryx lesnei (20); Eleoselinus villiersi (21). Arrows indicate particular states of the characters used in the phylogenetic analysis." figureDoi="http://doi.org/10.5281/zenodo.10831740" httpUri="https://zenodo.org/record/10831740/files/figure.png" pageId="21" pageNumber="94">Fig. 20</figureCitation>
).................................................................................
<taxonomicName id="03AF35CE1520BB4CFAF6FA43FA289C95" authorityName="Gebien" authorityYear="1910" box="[1291,1423,1428,1447]" class="Insecta" family="Tenebrionidae" genus="Glyptopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFAF6FA43FA289C95" box="[1291,1423,1428,1447]" italics="true" pageId="21" pageNumber="94">Glyptopteryx</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FA79FA289CED" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
Third antennomere long (ratio of its length to the length of second antennomere &lt;3.0). Elytral humeri not protruding towards body apex (
<figureCitation id="5C9452C81520BB4CFDECFA1CFDFE9CED" box="[529,601,1483,1503]" captionStart="Figures 1621" captionStartId="24.[144,235,1318,1337]" captionTargetBox="[369,1198,197,1287]" captionTargetId="figure-0@24.[367,1200,196,1289]" captionTargetPageId="24" captionText="Figures 1621. Intercoxal process of prosternum (17), hypomeron (16), scutellum (18), and base of elytron (1921). Anchophthalmus (Kochogaster) impressicollis (16); Upembarus (Pseudoselinus) punctatostriatus (17); Anchophthalmops platessa (18); Ectateus modestus (19); Glyptopteryx lesnei (20); Eleoselinus villiersi (21). Arrows indicate particular states of the characters used in the phylogenetic analysis." figureDoi="http://doi.org/10.5281/zenodo.10831740" httpUri="https://zenodo.org/record/10831740/files/figure.png" pageId="21" pageNumber="94">Fig. 19</figureCitation>
).................................................................................................(10)
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48FA30FA289F24" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
15. Bases of the first four elytral rows connected in the following way: 12, 34 (
<figureCitation id="5C9452C81520BB4CFBE3FA30FBCF9CC8" box="[1054,1128,1511,1530]" captionStart="Figures 1621" captionStartId="24.[144,235,1318,1337]" captionTargetBox="[369,1198,197,1287]" captionTargetId="figure-0@24.[367,1200,196,1289]" captionTargetPageId="24" captionText="Figures 1621. Intercoxal process of prosternum (17), hypomeron (16), scutellum (18), and base of elytron (1921). Anchophthalmus (Kochogaster) impressicollis (16); Upembarus (Pseudoselinus) punctatostriatus (17); Anchophthalmops platessa (18); Ectateus modestus (19); Glyptopteryx lesnei (20); Eleoselinus villiersi (21). Arrows indicate particular states of the characters used in the phylogenetic analysis." figureDoi="http://doi.org/10.5281/zenodo.10831740" httpUri="https://zenodo.org/record/10831740/files/figure.png" pageId="21" pageNumber="94">Fig. 19</figureCitation>
). Scutellum dented. Elytral epipleuron flat over its whole surface. Apex of the elytral epipleuron emarginated.............................
<emphasis id="F6DB925F1520BB4CFACBF9D3FA289F24" box="[1334,1423,1540,1558]" italics="true" pageId="21" pageNumber="94">Ectateus</emphasis>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48F9CAFA379F7C" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">Bases of the first four elytral rows not connected. Scutellum situated at the level of elytra. Elytral epipleuron curved at the level of the base of fifth ventrite. Apex of the elytral epipleuron not emarginated.................(16)</paragraph>
<paragraph id="C4104E4D1520BB4CFF48F980FA289FB4" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
16. Antennomeres from seventh to 11th elongated (
<figureCitation id="5C9452C81520BB4CFD2AF980FC959F58" box="[727,818,1623,1642]" captionStart="Figures 1115" captionStartId="23.[164,254,1326,1345]" captionTargetBox="[390,1218,197,1294]" captionTargetId="figure-0@23.[387,1220,195,1296]" captionTargetPageId="23" captionText="Figures 1115. Head dorsal view (11), antenna (12), and pronotum (1315). Anchophthalmops gridellii (12A), Ectateus ghesquierei (15); Glyptopteryx interioris (13); Lechius abacoides (11); Phymatoplata halsteadi (12B); Selinus planus (14). Arrows indicate particular states of the characters used in the phylogenetic analysis and autapomorphies (ap)." figureDoi="http://doi.org/10.5281/zenodo.10831738" httpUri="https://zenodo.org/record/10831738/files/figure.png" pageId="21" pageNumber="94">Fig. 12B</figureCitation>
). Parallel basal halves of the lateral elytral sides. Elytral intervals covered with coarse tubercles (
<figureCitation id="5C9452C81520BB4CFD6EF9A4FD7B9FB4" box="[659,732,1651,1670]" captionStart="Figures 2226" captionStartId="25.[164,254,1238,1257]" captionTargetBox="[431,1174,199,1206]" captionTargetId="figure-0@25.[428,1177,196,1209]" captionTargetPageId="25" captionText="Figures 2226. Apex of elytra (22), epipleuron (23, 26), abdominal ventrites (24), and anaepisternum (25). Phymatoplata halsteadi (22); Anchophthalmus (Kochogaster) impressicollis (23); Anchophthalmops gridellii (24); Zidalus latipes (25); Selinus planus (26). Arrows indicate particular states of the characters used in the phylogenetic analysis and autapomorphies (ap)." figureDoi="http://doi.org/10.5281/zenodo.10831742" httpUri="https://zenodo.org/record/10831742/files/figure.png" pageId="21" pageNumber="94">Fig. 22</figureCitation>
) ...................................................................
<taxonomicName id="03AF35CE1520BB4CFB01F9A4FA289FB4" authorityName="Koch" authorityYear="1956" box="[1276,1423,1651,1670]" class="Insecta" family="Tenebrionidae" genus="Phymatoplata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFB01F9A4FA289FB4" box="[1276,1423,1651,1670]" italics="true" pageId="21" pageNumber="94">Phymatoplata</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48F95AFA379F8C" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
Antennomeres from seventh to 11th transverse (
<figureCitation id="5C9452C81520BB4CFD24F958FC949F90" box="[729,819,1679,1698]" captionStart="Figures 1115" captionStartId="23.[164,254,1326,1345]" captionTargetBox="[390,1218,197,1294]" captionTargetId="figure-0@23.[387,1220,195,1296]" captionTargetPageId="23" captionText="Figures 1115. Head dorsal view (11), antenna (12), and pronotum (1315). Anchophthalmops gridellii (12A), Ectateus ghesquierei (15); Glyptopteryx interioris (13); Lechius abacoides (11); Phymatoplata halsteadi (12B); Selinus planus (14). Arrows indicate particular states of the characters used in the phylogenetic analysis and autapomorphies (ap)." figureDoi="http://doi.org/10.5281/zenodo.10831738" httpUri="https://zenodo.org/record/10831738/files/figure.png" pageId="21" pageNumber="94">Fig. 12A</figureCitation>
). Rounded basal halves of the lateral elytral sides. Elytral intervals without tubercles.............................................................................................................(17)
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48F910FA289FC4" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
17. Antennomeres from seventh to 11th triangular. Scutellum small (ratio of the width of a single elytra to the base of the scutellum&gt; 15.0;
<figureCitation id="5C9452C81520BB4CFE1EF934FD8A9FC4" box="[483,557,1763,1782]" captionStart="Figures 1621" captionStartId="24.[144,235,1318,1337]" captionTargetBox="[369,1198,197,1287]" captionTargetId="figure-0@24.[367,1200,196,1289]" captionTargetPageId="24" captionText="Figures 1621. Intercoxal process of prosternum (17), hypomeron (16), scutellum (18), and base of elytron (1921). Anchophthalmus (Kochogaster) impressicollis (16); Upembarus (Pseudoselinus) punctatostriatus (17); Anchophthalmops platessa (18); Ectateus modestus (19); Glyptopteryx lesnei (20); Eleoselinus villiersi (21). Arrows indicate particular states of the characters used in the phylogenetic analysis." figureDoi="http://doi.org/10.5281/zenodo.10831740" httpUri="https://zenodo.org/record/10831740/files/figure.png" pageId="21" pageNumber="94">Fig. 18</figureCitation>
). Base of parameres strongly widened (
<figureCitation id="5C9452C81520BB4CFC25F934FB869FC4" box="[984,1057,1763,1782]" captionStart="Figures 2729" captionStartId="25.[164,252,1765,1784]" captionTargetBox="[231,714,1432,1733]" captionTargetId="figure-51@25.[229,716,1430,1735]" captionTargetPageId="25" captionText="Figures 2729. Bursa copulatrix (27) and aedeagal tegmen (28, 29). Nesopatrum josephi (27); Anchophthalmops platessa (28); Phallocentrion edentatum (29). Arrows indicate particular states of the characters used in the phylogenetic analysis and autapomorphies (ap)." figureDoi="http://doi.org/10.5281/zenodo.10831744" httpUri="https://zenodo.org/record/10831744/files/figure.png" pageId="21" pageNumber="94">Fig. 28</figureCitation>
) .....................
<taxonomicName id="03AF35CE1520BB4CFB2EF934FA289FC4" authorityName="Koch" authorityYear="1956" box="[1235,1423,1763,1782]" class="Insecta" family="Tenebrionidae" genus="Anchophthalmops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFB2EF934FA289FC4" box="[1235,1423,1763,1782]" italics="true" pageId="21" pageNumber="94">Anchophthalmops</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4104E4D1520BB4CFF48F92AFA379E1C" blockId="21.[181,1425,215,1838]" pageId="21" pageNumber="94">
Antennomeres from seventh to 11th rounded. Scutellum large (ratio of the width of a single elytra to the base of the scutellum &lt;12.0). Base of parameres slightly widened (cf. Kamin´ ski, 2014) .............................
<taxonomicName id="03AF35CE1520BB4CFAE7F8CCFA379E1C" authorityName="KAMINSKI" authorityYear="2014" box="[1306,1424,1819,1838]" class="Insecta" family="Tenebrionidae" genus="Eleoselinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F6DB925F1520BB4CFAE7F8CCFA379E1C" box="[1306,1424,1819,1838]" italics="true" pageId="21" pageNumber="94">Eleoselinus</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>