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<document id="88CA952C164F80F41E8B15E6C41EFE79" ID-CLB-Dataset="24775" ID-DOI="10.11646/zootaxa.4701.4.3" ID-GBIF-Dataset="abe73cb2-b930-4796-8fcd-31d840646551" ID-ISSN="1175-5326" ID-Zenodo-Dep="3558201" ID-ZooBank="F22CEE0A-B1FF-4FBE-96C9-661F1151F5C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575044611900" checkinUser="plazi" docAuthor="Dow, Rory A." docDate="2019" docId="437D87B4FFE93247FF6C1C82FF09FB10" docLanguage="en" docName="zootaxa.4701.4.3.pdf" docOrigin="Zootaxa 4701 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphicnemis remiger Laidlaw 1912" docType="treatment" docVersion="8" lastPageNumber="381" masterDocId="BF44FFCCFFEC324DFFFB1A2CFFD5FFD7" masterDocTitle="Amphicnemis rigiketit sp. nov. from Sarawak, with notes on Amphicnemis remiger Laidlaw, 1912 (Odonata: Zygoptera: Coenagrionidae)" masterLastPageNumber="382" masterPageNumber="371" pageNumber="376" updateTime="1698769776141" updateUser="ExternalLinkService">
<mods:mods id="ACEA1CC411C82B4D42E5EABC00DDAD90" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D8EC8053FB82C5828192EE58FCB52835">
<mods:title id="B8DF6A8E0C8A1864C4A57CDD799F2552">Amphicnemis rigiketit sp. nov. from Sarawak, with notes on Amphicnemis remiger Laidlaw, 1912 (Odonata: Zygoptera: Coenagrionidae)</mods:title>
</mods:titleInfo>
<mods:name id="03689796965EE73DD62CAD7FA3BF0B40" type="personal">
<mods:role id="E491C4265EECC73B07B7435DB2BF080D">
<mods:roleTerm id="78758D59E50623628DE9B0AD38BEB867">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B553F891D678AD1552C41E847B0372C">Dow, Rory A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="ADF9131F53C1818DC229CAE32522EDE3">text</mods:typeOfResource>
<mods:relatedItem id="2E7F9E681E792E2BE59EBC157BC384A2" type="host">
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<mods:title id="C3CCBBA3DBC2E3AD5B5AF05ECDC422EF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7FCEBA743CB05A4A5E413364339A7D31">
<mods:date id="EF9E9A141510015B5198C49DF8D14C37">2019</mods:date>
<mods:detail id="D632BF947273AE685626A88148D945DC" type="pubDate">
<mods:number id="9809ED939A6E8AF96DC8CD101766318D">2019-11-29</mods:number>
</mods:detail>
<mods:detail id="DE91B4267CB17C27F8873A8CDF2AA754" type="volume">
<mods:number id="FDBF3746E214CB169FF356DDDD998356">4701</mods:number>
</mods:detail>
<mods:detail id="0F3FC52EAC94050A3FA3B82234458C91" type="issue">
<mods:number id="07C3CE2D4F5A7BF17F345F191F9023D8">4</mods:number>
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<mods:start id="2EF76CFD4F974151EF3F9F5C1923D4EC">371</mods:start>
<mods:end id="A236DECB81ACAC8EC2F28B5AF57AA0B3">382</mods:end>
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<mods:classification id="A6801E31B1F6742F7C0318CB2BB7F1CB">journal article</mods:classification>
<mods:identifier id="FAE0EBCE5B96B2C6F2F5B44D91D0CD89" type="CLB-Dataset">24775</mods:identifier>
<mods:identifier id="9DF69F6EDF4181541853C6B40752860E" type="DOI">10.11646/zootaxa.4701.4.3</mods:identifier>
<mods:identifier id="3442BB19ED6360BEA7FBCED1ACC8CC34" type="GBIF-Dataset">abe73cb2-b930-4796-8fcd-31d840646551</mods:identifier>
<mods:identifier id="BA6F951FD69E5BCF2CB661A373D452C4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6B67AD045F18B6D59F7FCFDEC0FBC83F" type="Zenodo-Dep">3558201</mods:identifier>
<mods:identifier id="235F2758143EC457233C5C032F7692B7" type="ZooBank">F22CEE0A-B1FF-4FBE-96C9-661F1151F5C7</mods:identifier>
</mods:mods>
<treatment id="437D87B4FFE93247FF6C1C82FF09FB10" ID-GBIF-Taxon="160605721" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10" httpUri="http://treatment.plazi.org/id/437D87B4FFE93247FF6C1C82FF09FB10" lastPageId="10" lastPageNumber="381" pageId="5" pageNumber="376">
<subSubSection id="83CE6529FFE93248FF6C1C82FCC4F93A" pageId="5" pageNumber="376" type="nomenclature">
<paragraph id="CB6B36A2FFE93248FF6C1C82FD90F91E" blockId="5.[151,785,1710,1773]" box="[151,581,1710,1737]" pageId="5" pageNumber="376">
<heading id="902381CEFFE93248FF6C1C82FD90F91E" bold="true" box="[151,581,1710,1737]" fontSize="11" level="1" pageId="5" pageNumber="376" reason="1">
<taxonomicName id="0CD44D21FFE93248FF6C1C82FD90F91E" ID-CoL="66F5N" authority="Laidlaw, 1912" authorityName="Laidlaw" authorityYear="1912" box="[151,581,1710,1737]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE93248FF6C1C82FD90F91E" bold="true" box="[151,581,1710,1737]" pageId="5" pageNumber="376">
<emphasis id="F9A0EAB0FFE93248FF6C1C82FE42F91E" bold="true" box="[151,407,1710,1737]" italics="true" pageId="5" pageNumber="376">Amphicnemis remiger</emphasis>
<bibRefCitation id="AF454B53FFE93248FE651C82FD90F91E" author="Laidlaw, F. F." box="[414,581,1710,1737]" pageId="5" pageNumber="376" pagination="92 - 99" refId="ref6499" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99., 1 pl." type="journal article" year="1912">Laidlaw, 1912</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="CB6B36A2FFE93248FF6C1CFFFCC4F93A" blockId="5.[151,785,1710,1773]" box="[151,785,1747,1773]" pageId="5" pageNumber="376">Figs (2, 4, 6, 8, 10, 12, 14, 16, 18, 20, 22, 24, 26, 28, 31)</paragraph>
</subSubSection>
<subSubSection id="83CE6529FFE9324AFF6C1D36FB80F945" lastPageId="7" lastPageNumber="378" pageId="5" pageNumber="376" type="materials_examined">
<paragraph id="CB6B36A2FFE9324BFF6C1D36FD29FE78" blockId="5.[151,1437,1818,2026]" lastBlockId="6.[151,1437,153,431]" lastPageId="6" lastPageNumber="377" pageId="5" pageNumber="376">
<emphasis id="F9A0EAB0FFE93248FF6C1D36FE56F8E2" bold="true" box="[151,387,1818,1845]" pageId="5" pageNumber="376">Material examined.</emphasis>
<materialsCitation id="7BBC3CFFFFE93248FE771D30FD0EF88E" ID-GBIF-Occurrence="2460219657" collectingDate="1911-05-11" collectorName="J. C. Moulton &amp; Natural History Museum &amp; London" country="Malaysia" location="Kuala Madalam" municipality="Limbang Division" pageId="5" pageNumber="376" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="F9A0EAB0FFE93248FE771D30FDD5F8E1" bold="true" box="[396,512,1820,1846]" pageId="5" pageNumber="376">
<typeStatus id="146F8800FFE93248FE771D30FE2EF8E1" box="[396,507,1820,1846]" pageId="5" pageNumber="376" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="DDD2FD2BFFE93248FDF21D30FDF5F8E1" box="[521,544,1820,1846]" pageId="5" pageNumber="376" type="male">♂</specimenCount>
,
<location id="CE0B6079FFE93248FDD51D37FD3DF8E2" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFE93248FDD51D37FD3DF8E2" box="[558,744,1819,1845]" country="Malaysia" municipality="Limbang Division" name="Kuala Madalam" pageId="5" pageNumber="376">Kuala Madalam</location>
,
<collectingMunicipality id="2B0FACD8FFE93248FD0E1D37FC13F8E2" box="[757,966,1819,1845]" pageId="5" pageNumber="376">Limbang Division</collectingMunicipality>
,
<collectingCountry id="B3C37632FFE93248FC2F1D37FBE2F8E2" box="[980,1079,1819,1845]" name="Malaysia" pageId="5" pageNumber="376">Sarawak</collectingCountry>
,
<collectingCountry id="B3C37632FFE93248FBBF1D37FB78F8E2" box="[1092,1197,1819,1845]" name="Malaysia" pageId="5" pageNumber="376">Malaysia</collectingCountry>
,
<date id="BF6A1062FFE93248FB401D37FAE5F8E2" box="[1211,1328,1819,1846]" pageId="5" pageNumber="376" value="1911-05-11">
<collectingDate id="AF2EE98AFFE93248FB401D37FAE5F8E2" box="[1211,1328,1819,1846]" pageId="5" pageNumber="376" value="1911-05-11">11 v 1911</collectingDate>
</date>
, leg.
<collectorName id="66215374FFE93248FA8A1D37FF2FF88E" pageId="5" pageNumber="376">J.C. Moulton</collectorName>
, in the
<collectorName id="66215374FFE93248FEA81D13FDA7F88D" box="[339,626,1855,1882]" pageId="5" pageNumber="376">Natural History Museum</collectorName>
,
<collectorName id="66215374FFE93248FD7B1D13FD0EF88E" box="[640,731,1855,1881]" pageId="5" pageNumber="376">London</collectorName>
</materialsCitation>
.
<emphasis id="F9A0EAB0FFE93248FD1C1D12FC71F88E" bold="true" box="[743,932,1854,1881]" pageId="5" pageNumber="376">Other material.</emphasis>
All in authors collection.
<collectingCountry id="B3C37632FFE93248FB2F1D13FAF0F88E" box="[1236,1317,1855,1881]" name="Brunei" pageId="5" pageNumber="376">Brunei</collectingCountry>
:
<materialsCitation id="7BBC3CFFFFE93248FACA1D13FC7FF8A9" ID-GBIF-Occurrence="2460219644" collectingDate="2013-03-02" country="Malaysia" location="Kuala Belai Road" pageId="5" pageNumber="376" specimenCount="1" specimenCount-male="1" stateProvince="Belait">
<specimenCount id="DDD2FD2BFFE93248FACA1D13FA8AF88D" box="[1329,1375,1855,1882]" pageId="5" pageNumber="376" type="male">1 ♂</specimenCount>
, peat swamp forest,
<location id="CE0B6079FFE93248FEC71D4FFDD2F8AA" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFE93248FEC71D4FFDD2F8AA" box="[316,519,1891,1917]" country="Malaysia" name="Kuala Belai Road" pageId="5" pageNumber="376" stateProvince="Belait">Kuala Belai Road</location>
,
<collectingRegion id="0910F840FFE93248FDE81D4FFD61F8AA" box="[531,692,1891,1917]" country="Brunei" name="Belait" pageId="5" pageNumber="376">Belait District</collectingRegion>
,
<date id="BF6A1062FFE93248FD3B1D4FFCFAF8AA" box="[704,815,1891,1917]" pageId="5" pageNumber="376" value="2013-03-02">
<collectingDate id="AF2EE98AFFE93248FD3B1D4FFCFAF8AA" box="[704,815,1891,1917]" pageId="5" pageNumber="376" value="2013-03-02">2 iii 2013</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFE93248FC4D1D4FFA4DF8A9" ID-GBIF-Occurrence="2460219655" box="[950,1432,1891,1918]" collectingDate="2013-03-04" country="Malaysia" location="Kuala Belai Road" pageId="5" pageNumber="376" specimenCount="4" specimenCount-male="4" stateProvince="Belait">
<specimenCount id="DDD2FD2BFFE93248FC4D1D4FFC22F8A9" box="[950,1015,1891,1918]" pageId="5" pageNumber="376" type="male">4 ♂♂</specimenCount>
, same location,
<date id="BF6A1062FFE93248FB551D4FFAC8F8AA" box="[1198,1309,1891,1917]" pageId="5" pageNumber="376" value="2013-03-04">
<collectingDate id="AF2EE98AFFE93248FB551D4FFAC8F8AA" box="[1198,1309,1891,1917]" pageId="5" pageNumber="376" value="2013-03-04">4 iii 2013</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFE93248FF6C1DABFA89F875" ID-GBIF-Occurrence="2460219656" box="[151,1372,1927,1954]" collectingDate="2013-03-08" country="Malaysia" location="Lumut Pipeline Road East" pageId="5" pageNumber="376" specimenCount="1" specimenCount-female="1" stateProvince="Belait">
<specimenCount id="DDD2FD2BFFE93248FF6C1DABFF16F875" box="[151,195,1927,1954]" pageId="5" pageNumber="376" type="female">1 ♀</specimenCount>
, low pH swamp forest accessed from
<location id="CE0B6079FFE93248FD851DA4FC7BF875" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFE93248FD851DA4FC7BF875" box="[638,942,1927,1954]" country="Malaysia" name="Lumut Pipeline Road East" pageId="5" pageNumber="376" stateProvince="Belait">Lumut Pipeline Road East</location>
,
<collectingRegion id="0910F840FFE93248FC401DABFB8BF876" box="[955,1118,1927,1953]" country="Brunei" name="Belait" pageId="5" pageNumber="376">Belait District</collectingRegion>
,
<date id="BF6A1062FFE93248FB901DABFB0BF876" box="[1131,1246,1927,1953]" pageId="5" pageNumber="376" value="2013-03-08">
<collectingDate id="AF2EE98AFFE93248FB901DABFB0BF876" box="[1131,1246,1927,1953]" pageId="5" pageNumber="376" value="2013-03-08">8 iii 2013</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFE93248FA921DABFB50F811" ID-GBIF-Occurrence="2460219643" collectingDate="2013-03-13" country="Malaysia" location="Lumut Pipeline Road East" pageId="5" pageNumber="376" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Belait">
<specimenCount id="DDD2FD2BFFE93248FA921DABFA42F875" box="[1385,1431,1927,1954]" pageId="5" pageNumber="376" type="male">1 ♂</specimenCount>
,
<specimenCount id="DDD2FD2BFFE93248FF6C1D87FF16F811" box="[151,195,1963,1990]" pageId="5" pageNumber="376" type="female">1 ♀</specimenCount>
(BRU13_COE103, in tandem with the male), same location,
<date id="BF6A1062FFE93248FC7C1D87FBD2F812" box="[903,1031,1963,1989]" pageId="5" pageNumber="376" value="2013-03-13">
<collectingDate id="AF2EE98AFFE93248FC7C1D87FBD2F812" box="[903,1031,1963,1989]" pageId="5" pageNumber="376" value="2013-03-13">13 iii 2013</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFE93248FB691D87FCD8F83D" ID-GBIF-Occurrence="2460219651" collectingDate="2013-03-11" country="Malaysia" location="Labi Road" pageId="5" pageNumber="376" specimenCount="9" specimenCount-male="9" stateProvince="Belait">
<specimenCount id="DDD2FD2BFFE93248FB691D87FB00F811" box="[1170,1237,1963,1990]" pageId="5" pageNumber="376" type="male">9 ♂♂</specimenCount>
, peat swamp for- est on
<location id="CE0B6079FFE93248FF191DE3FE8EF83E" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFE93248FF191DE3FE8EF83E" box="[226,347,1999,2025]" country="Malaysia" name="Labi Road" pageId="5" pageNumber="376" stateProvince="Belait">Labi Road</location>
,
<collectingRegion id="0910F840FFE93248FE9C1DE3FDDDF83E" box="[359,520,1999,2025]" country="Brunei" name="Belait" pageId="5" pageNumber="376">Belait District</collectingRegion>
,
<date id="BF6A1062FFE93248FDEF1DE3FD44F83E" box="[532,657,1999,2025]" pageId="5" pageNumber="376" value="2013-03-11">
<collectingDate id="AF2EE98AFFE93248FDEF1DE3FD44F83E" box="[532,657,1999,2025]" pageId="5" pageNumber="376" value="2013-03-11">11 iii 2013</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFE9324BFCE31DE3FE32FF64" ID-GBIF-Occurrence="2460219639" collectingDate="2013-05-22" country="Malaysia" lastPageId="6" lastPageNumber="377" location="Seria" pageId="5" pageNumber="376" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Belait">
<specimenCount id="DDD2FD2BFFE93248FCE31DE3FC8FF83D" box="[792,858,1999,2026]" pageId="5" pageNumber="376" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="DDD2FD2BFFE93248FC9C1DE3FC47F83D" box="[871,914,1999,2026]" pageId="5" pageNumber="376" type="female">1 ♀</specimenCount>
, remnant peat swamp forest near
<location id="CE0B6079FFE93248FAEA1DE3FA98F83E" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFE93248FAEA1DE3FA98F83E" box="[1297,1357,1999,2025]" country="Malaysia" name="Seria" pageId="5" pageNumber="376" stateProvince="Belait">Seria</location>
,
<collectingRegion id="0910F840FFE9324BFAA21DE3FF3BFF64" country="Brunei" lastPageId="6" lastPageNumber="377" name="Belait" pageId="5" pageNumber="376">Belait District</collectingRegion>
,
<date id="BF6A1062FFEA324BFF021AB5FEB9FF64" box="[249,364,153,179]" pageId="6" pageNumber="377" value="2013-05-22">
<collectingDate id="AF2EE98AFFEA324BFF021AB5FEB9FF64" box="[249,364,153,179]" pageId="6" pageNumber="377" value="2013-05-22">22 v 2013</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFE091AB5FE75FF00" ID-GBIF-Occurrence="2460219654" collectingDate="2013-05-31" country="Malaysia" location="Rassau area" pageId="6" pageNumber="377" specimenCount="3" specimenCount-male="3" stateProvince="Sabah">
<specimenCount id="DDD2FD2BFFEA324BFE091AB5FDE6FF64" box="[498,563,153,179]" pageId="6" pageNumber="377" type="male">3 ♂♂</specimenCount>
, peat swamp forest near highway,
<location id="CE0B6079FFEA324BFC4D1AB5FBE8FF64" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFC4D1AB5FBE8FF64" box="[950,1085,153,179]" country="Malaysia" name="Rassau area" pageId="6" pageNumber="377" stateProvince="Sabah">Rassau area</location>
, Belait District,
<date id="BF6A1062FFEA324BFB0E1AB5FABDFF64" box="[1269,1384,153,179]" pageId="6" pageNumber="377" value="2013-05-31">
<collectingDate id="AF2EE98AFFEA324BFB0E1AB5FABDFF64" box="[1269,1384,153,179]" pageId="6" pageNumber="377" value="2013-05-31">31 v 2013</collectingDate>
</date>
, leg. RAD.
<collectingCountry id="B3C37632FFEA324BFF1A1A91FE9FFF00" box="[225,330,189,215]" name="Malaysia" pageId="6" pageNumber="377">Malaysia</collectingCountry>
,
<collectingRegion id="0910F840FFEA324BFEAC1A91FE75FF00" box="[343,416,189,215]" country="Malaysia" name="Sabah" pageId="6" pageNumber="377">Sabah</collectingRegion>
</materialsCitation>
:
<materialsCitation id="7BBC3CFFFFEA324BFE501A91FC27FF00" ID-GBIF-Occurrence="2460219647" box="[427,1010,189,215]" collectingDate="2014-05-26" country="Malaysia" location="Klias Forest reserve" pageId="6" pageNumber="377" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<specimenCount id="DDD2FD2BFFEA324BFE501A91FE00FF00" box="[427,469,189,215]" pageId="6" pageNumber="377" type="female">1 ♀</specimenCount>
,
<location id="CE0B6079FFEA324BFE1F1A91FD1CFF00" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFE1F1A91FD1CFF00" box="[484,713,189,215]" country="Malaysia" name="Klias Forest reserve" pageId="6" pageNumber="377" stateProvince="Sabah">Klias Forest reserve</location>
,
<date id="BF6A1062FFEA324BFD2F1A91FC9CFF00" box="[724,841,189,215]" pageId="6" pageNumber="377" value="2014-05-26">
<collectingDate id="AF2EE98AFFEA324BFD2F1A91FC9CFF00" box="[724,841,189,215]" pageId="6" pageNumber="377" value="2014-05-26">26 v 2014</collectingDate>
</date>
, leg. unknown
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFC051A91FEA9FF2C" ID-GBIF-Occurrence="2460219653" collectingDate="2014-05-31" country="Malaysia" location="Klias Forest Reserve" pageId="6" pageNumber="377" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sabah">
<specimenCount id="DDD2FD2BFFEA324BFC051A91FBFFFF00" box="[1022,1066,189,215]" pageId="6" pageNumber="377" type="male">1 ♂</specimenCount>
,
<specimenCount id="DDD2FD2BFFEA324BFBCC1A91FBB7FF00" box="[1079,1122,189,215]" pageId="6" pageNumber="377" type="female">1 ♀</specimenCount>
,
<location id="CE0B6079FFEA324BFB8A1A91FA8BFF00" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFB8A1A91FA8BFF00" box="[1137,1374,189,215]" country="Malaysia" name="Klias Forest Reserve" pageId="6" pageNumber="377" stateProvince="Sabah">Klias Forest Reserve</location>
,
<date id="BF6A1062FFEA324BFA911A91FF07FF2C" pageId="6" pageNumber="377" value="2014-05-31">
<collectingDate id="AF2EE98AFFEA324BFA911A91FF07FF2C" pageId="6" pageNumber="377" value="2014-05-31">31 v 2014</collectingDate>
</date>
, leg. unknown
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFE731ACDFA8BFF2C" ID-GBIF-Occurrence="2460219640" box="[392,1374,225,251]" collectingDate="2018-05-28" country="Malaysia" location="Kampung Hiridian Forest Reserve" pageId="6" pageNumber="377" specimenCount="1" specimenCount-male="1">
<specimenCount id="DDD2FD2BFFEA324BFE731ACDFE63FF2C" box="[392,438,225,251]" pageId="6" pageNumber="377" type="male">1 ♂</specimenCount>
,
<location id="CE0B6079FFEA324BFE381ACDFC9EFF2C" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFE381ACDFC9EFF2C" box="[451,843,225,251]" country="Malaysia" name="Kampung Hiridian Forest Reserve" pageId="6" pageNumber="377">Kampung Hiridian Forest Reserve</location>
,
<date id="BF6A1062FFEA324BFCAC1ACDFC18FF2C" box="[855,973,225,251]" pageId="6" pageNumber="377" value="2018-05-28">
<collectingDate id="AF2EE98AFFEA324BFCAC1ACDFC18FF2C" box="[855,973,225,251]" pageId="6" pageNumber="377" value="2018-05-28">28 v 2018</collectingDate>
</date>
, leg. unknown.
<collectingCountry id="B3C37632FFEA324BFB781ACDFB3BFF2C" box="[1155,1262,225,251]" name="Malaysia" pageId="6" pageNumber="377">Malaysia</collectingCountry>
: Sarawak
</materialsCitation>
:
<materialsCitation id="7BBC3CFFFFEA324BFA911ACDFAD0FEC8" ID-GBIF-Occurrence="2460219642" collectingDate="2006-02-11" collectorName="J. Simun" country="Malaysia" county="Headhunters Trail" location="Limbang Division" municipality="Gunung Mulu National Park" pageId="6" pageNumber="377" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="DDD2FD2BFFEA324BFA911ACDFA42FF2C" box="[1386,1431,225,251]" pageId="6" pageNumber="377" type="male">1 ♂</specimenCount>
,
<specimenCount id="DDD2FD2BFFEA324BFF6C1B29FF0DFEC8" box="[151,216,261,287]" pageId="6" pageNumber="377" type="female">2 ♀♀</specimenCount>
,
<collectingCounty id="220A4E2EFFEA324BFF1D1B29FE66FEC8" box="[230,435,261,287]" pageId="6" pageNumber="377">Headhunters Trail</collectingCounty>
,
<collectingMunicipality id="2B0FACD8FFEA324BFE441B29FCD1FEC8" box="[447,772,261,287]" pageId="6" pageNumber="377">Gunung Mulu National Park</collectingMunicipality>
,
<location id="CE0B6079FFEA324BFCEB1B29FC35FEC8" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFCEB1B29FC35FEC8" box="[784,992,261,287]" country="Malaysia" county="Headhunters Trail" municipality="Gunung Mulu National Park" name="Limbang Division" pageId="6" pageNumber="377">Limbang Division</location>
,
<date id="BF6A1062FFEA324BFC171B29FBB4FEC8" box="[1004,1121,261,287]" pageId="6" pageNumber="377" value="2006-02-11">
<collectingDate id="AF2EE98AFFEA324BFC171B29FBB4FEC8" box="[1004,1121,261,287]" pageId="6" pageNumber="377" value="2006-02-11">11 ii 2006</collectingDate>
</date>
, leg.
<collectorName id="66215374FFEA324BFB651B29FAD0FEC8" box="[1182,1285,261,287]" pageId="6" pageNumber="377">J. Simun</collectorName>
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFAEA1B29FAB1FE94" ID-GBIF-Occurrence="2460219649" collectingDate="2006-02-13" collectorName="J. Simun" country="Malaysia" county="Boundary Trail" location="Gunung Mulu National Park" municipality="Mentawei" pageId="6" pageNumber="377" specimenCount="2" specimenCount-male="2">
<specimenCount id="DDD2FD2BFFEA324BFAEA1B29FA86FEC8" box="[1297,1363,261,287]" pageId="6" pageNumber="377" type="male">2 ♂♂</specimenCount>
, Men- tawei
<collectingCounty id="220A4E2EFFEA324BFF271B05FE5EFE94" box="[220,395,297,323]" pageId="6" pageNumber="377">Boundary Trail</collectingCounty>
,
<collectingMunicipality id="2B0FACD8FFEA324BFE6C1B05FDDFFE94" box="[407,522,297,323]" pageId="6" pageNumber="377">Mentawei</collectingMunicipality>
,
<location id="CE0B6079FFEA324BFDEC1B05FC88FE94" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFDEC1B05FC88FE94" box="[535,861,297,323]" country="Malaysia" county="Boundary Trail" municipality="Mentawei" name="Gunung Mulu National Park" pageId="6" pageNumber="377">Gunung Mulu National Park</location>
,
<location id="CE0B6079FFEA324BFC911B05FBEEFE94" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFC911B05FBEEFE94" box="[874,1083,297,323]" country="Malaysia" county="Boundary Trail" municipality="Mentawei" name="Limbang Division" pageId="6" pageNumber="377">Limbang Division</location>
,
<date id="BF6A1062FFEA324BFBB31B05FB6AFE94" box="[1096,1215,297,323]" pageId="6" pageNumber="377" value="2006-02-13">
<collectingDate id="AF2EE98AFFEA324BFBB31B05FB6AFE94" box="[1096,1215,297,323]" pageId="6" pageNumber="377" value="2006-02-13">13 ii 2006</collectingDate>
</date>
, leg.
<collectorName id="66215374FFEA324BFB061B05FAB1FE94" box="[1277,1380,297,323]" pageId="6" pageNumber="377">J. Simun</collectorName>
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFA8A1B05FAB3FEB0" ID-GBIF-Occurrence="2460219648" collectingDate="2011-09-05" country="Malaysia" county="Sungai Dabai" location="Miri Division" municipality="Marudi" pageId="6" pageNumber="377" specimenCount="1" specimenCount-female="1">
<specimenCount id="DDD2FD2BFFEA324BFA8A1B05FA49FE94" box="[1393,1436,297,323]" pageId="6" pageNumber="377" type="female">1 ♀</specimenCount>
(SAR11_12_COE63), dry peat swamp forest near
<collectingCounty id="220A4E2EFFEA324BFD341B61FCBFFEB0" box="[719,874,333,359]" pageId="6" pageNumber="377">Sungai Dabai</collectingCounty>
,
<collectingMunicipality id="2B0FACD8FFEA324BFC8E1B61FC1FFEB0" box="[885,970,333,359]" pageId="6" pageNumber="377">Marudi</collectingMunicipality>
,
<location id="CE0B6079FFEA324BFC2D1B61FBA7FEB0" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFC2D1B61FBA7FEB0" box="[982,1138,333,359]" country="Malaysia" county="Sungai Dabai" municipality="Marudi" name="Miri Division" pageId="6" pageNumber="377">Miri Division</location>
,
<date id="BF6A1062FFEA324BFB851B60FB3EFEB0" box="[1150,1259,332,359]" pageId="6" pageNumber="377" value="2011-09-05">
<collectingDate id="AF2EE98AFFEA324BFB851B60FB3EFEB0" box="[1150,1259,332,359]" pageId="6" pageNumber="377" value="2011-09-05">5 ix 2011</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFA891B61FB69FE5C" ID-GBIF-Occurrence="2460219641" collectingDate="2018-09-06" country="Brunei" location="Trail" pageId="6" pageNumber="377" specimenCount="1" specimenCount-male="1">
<specimenCount id="DDD2FD2BFFEA324BFA891B61FA49FEB0" box="[1394,1436,333,359]" pageId="6" pageNumber="377" type="male">1 ♂</specimenCount>
(SAR11_12_COE332), peat swamp forest on
<collectingCountry id="B3C37632FFEA324BFD641B5DFD3EFE5C" box="[671,747,369,395]" name="Brunei" pageId="6" pageNumber="377">Brunei</collectingCountry>
<location id="CE0B6079FFEA324BFD081B5DFCFEFE5C" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFD081B5DFCFEFE5C" box="[755,811,369,395]" country="Brunei" name="Trail" pageId="6" pageNumber="377">Trail</location>
, Marudi area,
<date id="BF6A1062FFEA324BFC2B1B5DFB95FE5C" box="[976,1088,369,395]" pageId="6" pageNumber="377" value="2018-09-06">
<collectingDate id="AF2EE98AFFEA324BFC2B1B5DFB95FE5C" box="[976,1088,369,395]" pageId="6" pageNumber="377" value="2018-09-06">6 ix 2018</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFB331B5DFD2CFE78" ID-GBIF-Occurrence="2460219645" collectingDate="2011-09-08" country="Brunei" location="Miri Division" municipality="Marudi" pageId="6" pageNumber="377" specimenCount="1" specimenCount-male="1">
<specimenCount id="DDD2FD2BFFEA324BFB331B5DFB23FE5C" box="[1224,1270,369,395]" pageId="6" pageNumber="377" type="male">1 ♂</specimenCount>
, old rubber on peat, near
<collectingMunicipality id="2B0FACD8FFEA324BFEF01BB9FEB5FE78" box="[267,352,405,431]" pageId="6" pageNumber="377">Marudi</collectingMunicipality>
,
<location id="CE0B6079FFEA324BFE901BB9FDD2FE78" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFE901BB9FDD2FE78" box="[363,519,405,431]" country="Brunei" municipality="Marudi" name="Miri Division" pageId="6" pageNumber="377">Miri Division</location>
,
<date id="BF6A1062FFEA324BFDE81BB9FDABFE78" box="[531,638,405,431]" pageId="6" pageNumber="377" value="2011-09-08">
<collectingDate id="AF2EE98AFFEA324BFDE81BB9FDABFE78" box="[531,638,405,431]" pageId="6" pageNumber="377" value="2011-09-08">8 ix 2011</collectingDate>
</date>
, leg. RAD
</materialsCitation>
.
</paragraph>
<caption id="9FAB662AFFEA324BFF6C1FEAFE6DF9F0" ID-DOI="http://doi.org/10.5281/zenodo.3558211" ID-Zenodo-Dep="3558211" httpUri="https://zenodo.org/record/3558211/files/figure.png" pageId="6" pageNumber="377" startId="6.[151,264,1478,1503]" targetBox="[151,1437,453,1453]" targetPageId="6">
<paragraph id="CB6B36A2FFEA324BFF6C1FEAFE6DF9F0" blockId="6.[151,1437,1478,1575]" pageId="6" pageNumber="377">
<emphasis id="F9A0EAB0FFEA324BFF6C1FEAFE80FA08" bold="true" box="[151,341,1478,1503]" pageId="6" pageNumber="377">FIGURES 17–20.</emphasis>
Lateral view of synthorax of
<taxonomicName id="0CD44D21FFEA324BFD721FEBFCC4FA08" authorityName="Selys" authorityYear="1863" box="[649,785,1479,1503]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="377" phylum="Arthropoda" rank="genus">
<emphasis id="F9A0EAB0FFEA324BFD721FEBFCC4FA08" box="[649,785,1479,1503]" italics="true" pageId="6" pageNumber="377">Amphicnemis</emphasis>
</taxonomicName>
spp. (17)
<taxonomicName id="0CD44D21FFEA324BFC801FEBFB8EFA08" authorityName="Dow" authorityYear="2019" box="[891,1115,1479,1503]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFEA324BFC801FEBFB8EFA08" box="[891,1115,1479,1503]" italics="true" pageId="6" pageNumber="377">Amphicnemis rigiketit</emphasis>
</taxonomicName>
sp. holotype male: (19)
<taxonomicName id="0CD44D21FFEA324BFAA21FEBFEEEF9D4" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFEA324BFAA21FEBFEEEF9D4" italics="true" pageId="6" pageNumber="377">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFEA324BFEB91FC7FE46F9D5" box="[322,403,1515,1538]" pageId="6" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
, female described; (18) male
<taxonomicName id="0CD44D21FFEA324BFD441FC7FC48F9D4" authorityName="Laidlaw" authorityYear="1912" box="[703,925,1515,1539]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEA324BFD441FC7FC48F9D4" box="[703,925,1515,1539]" italics="true" pageId="6" pageNumber="377">Amphicnemis remiger</emphasis>
</taxonomicName>
(SAR11_12_COE332); (20) female
<taxonomicName id="0CD44D21FFEA324BFAEF1FC7FF33F9F0" authorityName="Laidlaw" authorityYear="1912" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEA324BFAEF1FC7FF33F9F0" italics="true" pageId="6" pageNumber="377">Amphicnemis remiger</emphasis>
</taxonomicName>
(BRU13_COE103).
</paragraph>
</caption>
<paragraph id="CB6B36A2FFEA324BFF3C1C78FCBAF9B8" blockId="6.[151,1437,1620,2044]" box="[199,879,1620,1648]" pageId="6" pageNumber="377">
<emphasis id="F9A0EAB0FFEA324BFF3C1C78FE2CF9B8" bold="true" box="[199,505,1620,1647]" pageId="6" pageNumber="377">Descriptive notes on male</emphasis>
(based on SAR11_12_COE332).
</paragraph>
<paragraph id="CB6B36A2FFEA324BFF3C1C55FA9CF858" blockId="6.[151,1437,1620,2044]" pageId="6" pageNumber="377">
<emphasis id="F9A0EAB0FFEA324BFF3C1C55FED1F944" box="[199,260,1657,1683]" italics="true" pageId="6" pageNumber="377">Head</emphasis>
(
<figureCitation id="53EF2A27FFEA324BFEE81C55FEAFF944" box="[275,378,1657,1683]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1207,1232]" captionTargetBox="[151,1437,179,1179]" captionTargetId="figure@2.[151,1437,179,1179]" captionTargetPageId="2" captionText="FIGURES 1–4. Amphicnemis rigiketit sp. nov., head of holotype male: (1), dorsal-frontal view; (3) dorsal view. Head of male Amphicnemis remiger (SAR11_12_COE332); (2) dorsal-frontal view; (4) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558203" httpUri="https://zenodo.org/record/3558203/files/figure.png" pageId="6" pageNumber="377">Figs 2, 4</figureCitation>
). Labium entirely pale. Labrum black, narrowly pale centrally at free margin. Mandible bases black with cream outline except adjacent to genae. Genae pale, this extended narrowly beside compound eye to beyond level of antennae bases. Horizontal surface of clypeus shining black, vertical surface pale with intrusions of black from above (
<figureCitation id="53EF2A27FFEA324BFE771CC9FE07F928" box="[396,466,1765,1791]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1207,1232]" captionTargetBox="[151,1437,179,1179]" captionTargetId="figure@2.[151,1437,179,1179]" captionTargetPageId="2" captionText="FIGURES 1–4. Amphicnemis rigiketit sp. nov., head of holotype male: (1), dorsal-frontal view; (3) dorsal view. Head of male Amphicnemis remiger (SAR11_12_COE332); (2) dorsal-frontal view; (4) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558203" httpUri="https://zenodo.org/record/3558203/files/figure.png" pageId="6" pageNumber="377">Fig. 2</figureCitation>
). Vertical face of frons shining black becoming dark metallic green above, but largely occupied by irregular, pale transverse stripe very narrowly divided centrally, just joined to pale colour on genae. Rest of frons, vertex, occiput and underside of head bronzy metallic green. Ocelli whitish. Antenna bases with broad pale stripe on anterior face, this just extending to base of scape, scape black, pale at top, base of pedicel pale, this extended on anterior and posterior faces as stripe running almost to top, remainder brown, flagella missing.
</paragraph>
<paragraph id="CB6B36A2FFEA324AFF3C1DB5FE42FE04" blockId="6.[151,1437,1620,2044]" lastBlockId="7.[151,1437,153,467]" lastPageId="7" lastPageNumber="378" pageId="6" pageNumber="377">
<emphasis id="F9A0EAB0FFEA324BFF3C1DB5FECDF864" box="[199,280,1945,1971]" italics="true" pageId="6" pageNumber="377">Thorax</emphasis>
. Prothorax (
<figureCitation id="53EF2A27FFEA324BFE5A1DB5FDCAF864" box="[417,543,1945,1971]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1677,1702]" captionTargetBox="[151,1436,417,1652]" captionTargetId="figure@4.[151,1436,417,1652]" captionTargetPageId="4" captionText="FIGURES 9–12. Amphicnemis rigiketit sp. nov., prothorax of holotype male: (9), dorsal view; (11) lateral view. Prothorax of male Amphicnemis remiger (SAR11_12_COE332); (10) dorsal view; (12) lateral view." figureDoi="http://doi.org/10.5281/zenodo.3558207" httpUri="https://zenodo.org/record/3558207/files/figure.png" pageId="6" pageNumber="377">Figs 10, 12</figureCitation>
) with pronotum almost entirely dark metallic green apart from pair of small pale marks at rear anterior pronotal lobe, propleuron mostly cream. Posterior pronotal lobe short, collar-like centrally but with rear lateral extremities sharply angulated, directed more to rear than to sides (
<figureCitation id="53EF2A27FFEA324BFBAF1DCDFB73F82C" box="[1108,1190,2017,2043]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1677,1702]" captionTargetBox="[151,1436,417,1652]" captionTargetId="figure@4.[151,1436,417,1652]" captionTargetPageId="4" captionText="FIGURES 9–12. Amphicnemis rigiketit sp. nov., prothorax of holotype male: (9), dorsal view; (11) lateral view. Prothorax of male Amphicnemis remiger (SAR11_12_COE332); (10) dorsal view; (12) lateral view." figureDoi="http://doi.org/10.5281/zenodo.3558207" httpUri="https://zenodo.org/record/3558207/files/figure.png" pageId="6" pageNumber="377">Fig. 12</figureCitation>
). Synthorax (
<figureCitation id="53EF2A27FFEA324BFAC01DCDFA5BF82C" box="[1339,1422,2017,2043]" captionStart="FIGURES 17–20" captionStartId="6.[151,264,1478,1503]" captionTargetBox="[151,1437,453,1453]" captionTargetId="figure@6.[151,1437,453,1453]" captionTargetPageId="6" captionText="FIGURES 17–20. Lateral view of synthorax of Amphicnemis spp. (17) Amphicnemis rigiketit sp. holotype male: (19) Amphicnemis rigiketit sp. nov., female described; (18) male Amphicnemis remiger (SAR11_12_COE332); (20) female Amphicnemis remiger (BRU13_COE103)." figureDoi="http://doi.org/10.5281/zenodo.3558211" httpUri="https://zenodo.org/record/3558211/files/figure.png" pageId="6" pageNumber="377">Fig. 18</figureCitation>
): mesepisternum, mesepimeron, most of mesokatepisternum and metepisternum to top of spiracle, and extending to metepleural suture adjacent to antealar carina, metallic green, remainder of synthorax whitish except dark mark near antealar carina on metepimeron. Legs (right anterior leg absent below trochanter) cream with flexor surface femurs white, poorly defined, incomplete narrow dark stripes on extensor surface femurs, dark brown to black spines, small dark mark lowest part femur and immediately adjacent part flexor surface tibia, black rings between tarsal segments, tips of pale brown claws black. Wings: 12 Px in Fw, 10 (left) or 11 (right) in Hw. Arculus slightly distal to Ax2.
<collectionCode id="ADC5AE67FFEB324AFA7A1B61FA41FEB0" box="[1409,1428,333,359]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="378">R</collectionCode>
<subScript id="575034E7FFEB324AFA6F1B72FA49FEBB" attach="left" box="[1428,1436,350,364]" fontSize="6" pageId="7" pageNumber="378">4</subScript>
arising just distal to subnodus in all wings, IR
<subScript id="575034E7FFEB324AFD681BAEFD4EFE47" attach="left" box="[659,667,386,400]" fontSize="6" pageId="7" pageNumber="378">3</subScript>
joined to it by a short stalk. Pterostigma almost trapezoidal with costal side shorter than anal side, dark brownish grey with whitish border (poorly defined on proximal side), covering ca one underlying cell.
</paragraph>
<caption id="9FAB662AFFEB324AFF6C19DDFC37FBFA" ID-DOI="http://doi.org/10.5281/zenodo.3558213" ID-Zenodo-Dep="3558213" httpUri="https://zenodo.org/record/3558213/files/figure.png" pageId="7" pageNumber="378" startId="7.[151,264,1009,1034]" targetBox="[158,1430,489,984]" targetPageId="7">
<paragraph id="CB6B36A2FFEB324AFF6C19DDFC37FBFA" blockId="7.[151,1436,1009,1070]" pageId="7" pageNumber="378">
<emphasis id="F9A0EAB0FFEB324AFF6C19DDFE83FBDD" bold="true" box="[151,342,1009,1034]" pageId="7" pageNumber="378">FIGURES 21–22.</emphasis>
Lateral view of pattern of terminal abdominal segments of female
<taxonomicName id="0CD44D21FFEB324AFBEA19DDFB4CFBDE" authorityName="Selys" authorityYear="1863" box="[1041,1177,1009,1033]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="378" phylum="Arthropoda" rank="genus">
<emphasis id="F9A0EAB0FFEB324AFBEA19DDFB4CFBDE" box="[1041,1177,1009,1033]" italics="true" pageId="7" pageNumber="378">Amphicnemis</emphasis>
</taxonomicName>
spp.: (21),
<taxonomicName id="0CD44D21FFEB324AFAEF19DDFF32FBFA" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFEB324AFAEF19DDFF32FBFA" italics="true" pageId="7" pageNumber="378">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFEB324AFF151E3AFEEAFBFA" box="[238,319,1046,1069]" pageId="7" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
, female described; (22)
<taxonomicName id="0CD44D21FFEB324AFDC91E39FCC5FBFA" authorityName="Laidlaw" authorityYear="1912" box="[562,784,1045,1069]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEB324AFDC91E39FCC5FBFA" box="[562,784,1045,1069]" italics="true" pageId="7" pageNumber="378">Amphicnemis remiger</emphasis>
</taxonomicName>
(BRU13_COE103).
</paragraph>
</caption>
<paragraph id="CB6B36A2FFEB324AFF3C1E70FD55F9B8" blockId="7.[151,1437,1116,2042]" pageId="7" pageNumber="378">
<emphasis id="F9A0EAB0FFEB324AFF3C1E70FEE4FBA1" box="[199,305,1116,1142]" italics="true" pageId="7" pageNumber="378">Abdomen</emphasis>
. S1 cream with apical transverse dark mark extended laterally apically. S2 metallic green above, cream below. S3–6 largely very dark brown above, cream lower laterally, this becoming darker on successive segments. S7 similar but almost entirely dark brown laterally in apical ca one-third. S8 bronzy black above, dark brown below. S9 entirely bronzy black, S10 white except dorsally where mostly brown. Anal appendages (
<figureCitation id="53EF2A27FFEB324AFAE91EE4FF61FAD1" captionStart="FIGURES 23–28" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 23–28. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="7" pageNumber="378">Figs 24, 26, 28</figureCitation>
) white. Cerci in lateral view with upper branch narrow at base, directed gently upwards from base, then straight before expanding along lower margin after half-length so that apical part is rounded ventrally and at apex (
<figureCitation id="53EF2A27FFEB324AFAC71F3CFA45FAFD" box="[1340,1424,1296,1322]" captionStart="FIGURES 23–28" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 23–28. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="7" pageNumber="378">Fig. 24</figureCitation>
). Lower branch of cercus tapering from base before expanding slightly subapically where directed gently upwards to narrow but rounded tip, a small dorsal black subapical tooth just visible in this view (
<figureCitation id="53EF2A27FFEB324AFB761F74FB36FAA5" box="[1165,1251,1368,1394]" captionStart="FIGURES 23–28" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 23–28. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="7" pageNumber="378">Fig. 24</figureCitation>
). In dorsal view (
<figureCitation id="53EF2A27FFEB324AFF5B1F50FF27FA40" box="[160,242,1404,1431]" captionStart="FIGURES 23–28" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 23–28. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="7" pageNumber="378">Fig. 26</figureCitation>
) upper branch with a small subbasal subtriangular tooth after which contracted along inner margin before expanding again just after half length, the inner margin with an excised appearance between the subbasal tooth and the subapical expansion.A strong ridge runs to the apex along the outer margin, giving the expanded part the appearance of a heel-like shelf below it. In ventral view lower branch of cercus tapering gently from base, turning gently inwards at ca two-thirds length, a small, poorly defined subapical tooth on inner margin after which running straight or slightly outwards to rounded but marrow tip (
<figureCitation id="53EF2A27FFEB324AFD3F1C1CFCCEF99D" box="[708,795,1584,1610]" captionStart="FIGURES 23–28" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 23–28. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="7" pageNumber="378">Fig. 28</figureCitation>
). Paraprocts short, pale with short finger like projection directed to rear in dorsal part (lateral view).
</paragraph>
<paragraph id="CB6B36A2FFEB324AFF3C1C54FB80F945" blockId="7.[151,1437,1116,2042]" box="[199,1109,1656,1682]" pageId="7" pageNumber="378">Measurements (mm). Abdomen without anal appendages 34, cercus ca 1, Hw 20.</paragraph>
</subSubSection>
<subSubSection id="83CE6529FFEB3247FF3C1CB7FF2FFF2C" lastPageId="10" lastPageNumber="381" pageId="7" pageNumber="378" type="description">
<paragraph id="CB6B36A2FFEB324AFF3C1CB7FCC2F961" blockId="7.[151,1437,1116,2042]" box="[199,791,1691,1718]" pageId="7" pageNumber="378">
<emphasis id="F9A0EAB0FFEB324AFF3C1CB7FE13F961" bold="true" box="[199,454,1691,1718]" pageId="7" pageNumber="378">Description of female</emphasis>
(based on BRU13_COE103).
</paragraph>
<paragraph id="CB6B36A2FFEB324AFF3C1CECFE0AF90D" blockId="7.[151,1437,1116,2042]" box="[199,479,1728,1754]" pageId="7" pageNumber="378">As male except as noted.</paragraph>
<paragraph id="CB6B36A2FFEB324AFF3C1CC8FA9AF929" blockId="7.[151,1437,1116,2042]" box="[199,1359,1764,1790]" pageId="7" pageNumber="378">
<emphasis id="F9A0EAB0FFEB324AFF3C1CC8FED1F929" box="[199,260,1764,1790]" italics="true" pageId="7" pageNumber="378">Head</emphasis>
(
<figureCitation id="53EF2A27FFEB324AFEE81CC8FEA2F929" box="[275,375,1764,1790]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1370,1395]" captionTargetBox="[151,1437,345,1345]" captionTargetId="figure@3.[151,1437,345,1345]" captionTargetPageId="3" captionText="FIGURES 5–8. Amphicnemis rigiketit sp. nov., head of female described: (5), dorsal-frontal view; (7) dorsal view. Head of female Amphicnemis remiger (BRU13_COE103); (6) dorsal-frontal view; (8) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558205" httpUri="https://zenodo.org/record/3558205/files/figure.png" pageId="7" pageNumber="378">Figs 6, 8</figureCitation>
). More extensive pale area on labrum, pale stripe vertical face of frons broadly divided.
</paragraph>
<paragraph id="CB6B36A2FFEB324AFF3C1D24FDEFF82D" blockId="7.[151,1437,1116,2042]" pageId="7" pageNumber="378">
<emphasis id="F9A0EAB0FFEB324AFF3C1D24FECDF8F5" box="[199,280,1800,1826]" italics="true" pageId="7" pageNumber="378">Thorax</emphasis>
. Prothorax (
<figureCitation id="53EF2A27FFEB324AFE5E1D24FDF3F8F4" box="[421,550,1800,1827]" captionStart="FIGURES 13–16" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 13–16. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="7" pageNumber="378">Figs 14, 16</figureCitation>
) almost entirely blue, anterior carina of anterior pronotal lobe black. Posterior pronotal lobe collar-like centrally in dorsal view (
<figureCitation id="53EF2A27FFEB324AFD321D00FCC9F891" box="[713,796,1836,1862]" captionStart="FIGURES 13–16" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 13–16. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="7" pageNumber="378">Fig. 14</figureCitation>
), laterally contracted before slightly folded upwards and then extended downwards to rear as a subtriangular projection, free margin running downwards back to base (
<figureCitation id="53EF2A27FFEB324AFA941D7CFF63F858" captionStart="FIGURES 13–16" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 13–16. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="7" pageNumber="378">Fig. 16</figureCitation>
). Postepimeral strip expanded to rear just below posterior pronotal lobe (this indicated with a white line in
<figureCitation id="53EF2A27FFEB324AFA8B1D58FF63F864" captionStart="FIGURES 13–16" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 13–16. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="7" pageNumber="378">Fig. 16</figureCitation>
). Synthorax (
<figureCitation id="53EF2A27FFEB324AFEA91DB4FE70F865" box="[338,421,1944,1970]" captionStart="FIGURES 17–20" captionStartId="6.[151,264,1478,1503]" captionTargetBox="[151,1437,453,1453]" captionTargetId="figure@6.[151,1437,453,1453]" captionTargetPageId="6" captionText="FIGURES 17–20. Lateral view of synthorax of Amphicnemis spp. (17) Amphicnemis rigiketit sp. holotype male: (19) Amphicnemis rigiketit sp. nov., female described; (18) male Amphicnemis remiger (SAR11_12_COE332); (20) female Amphicnemis remiger (BRU13_COE103)." figureDoi="http://doi.org/10.5281/zenodo.3558211" httpUri="https://zenodo.org/record/3558211/files/figure.png" pageId="7" pageNumber="378">Fig. 20</figureCitation>
) bluish with yellowish cast dorsally (not apparent in life). Legs as male but marking on extensor surface femurs better defined as narrow black line. Wings with 12 Px in Fw, 11 Px in Hw, white border of pterostigma well defined on all sides.
</paragraph>
<caption id="9FAB662AFFE43245FF6C1CAAFDCEF930" ID-DOI="http://doi.org/10.5281/zenodo.3558215" ID-Zenodo-Dep="3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="8" pageNumber="379" startId="8.[151,264,1670,1695]" targetBox="[151,1436,181,1644]" targetPageId="8">
<paragraph id="CB6B36A2FFE43245FF6C1CAAFDCEF930" blockId="8.[151,1436,1670,1767]" pageId="8" pageNumber="379">
<emphasis id="F9A0EAB0FFE43245FF6C1CAAFE83F948" bold="true" box="[151,342,1670,1695]" pageId="8" pageNumber="379">FIGURES 23–28.</emphasis>
<taxonomicName id="0CD44D21FFE43245FEA61CAAFDE8F949" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" box="[349,573,1670,1694]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFE43245FEA61CAAFDE8F949" box="[349,573,1670,1694]" italics="true" pageId="8" pageNumber="379">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFE43245FDBE1CABFD4DF949" box="[581,664,1671,1694]" pageId="8" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
, anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male
<taxonomicName id="0CD44D21FFE43245FDA71C86FCEEF915" authorityName="Laidlaw" authorityYear="1912" box="[604,827,1706,1730]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE43245FDA71C86FCEEF915" box="[604,827,1706,1730]" italics="true" pageId="8" pageNumber="379">Amphicnemis remiger</emphasis>
</taxonomicName>
(SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm.
</paragraph>
</caption>
<paragraph id="CB6B36A2FFE43245FF3C1D39FCA1F84C" blockId="8.[151,1436,1813,2019]" pageId="8" pageNumber="379">
<emphasis id="F9A0EAB0FFE43245FF3C1D39FEE4F8F8" box="[199,305,1813,1839]" italics="true" pageId="8" pageNumber="379">Abdomen</emphasis>
(
<figureCitation id="53EF2A27FFE43245FEB81D39FE4FF8F8" box="[323,410,1813,1839]" captionStart="FIGURES 21–22" captionStartId="7.[151,264,1009,1034]" captionTargetBox="[158,1430,489,984]" captionTargetId="figure@7.[158,1430,489,984]" captionTargetPageId="7" captionText="FIGURES 21–22. Lateral view of pattern of terminal abdominal segments of female Amphicnemis spp.: (21), Amphicnemis rigiketit sp. nov., female described; (22) Amphicnemis remiger (BRU13_COE103)." figureDoi="http://doi.org/10.5281/zenodo.3558213" httpUri="https://zenodo.org/record/3558213/files/figure.png" pageId="8" pageNumber="379">Fig. 22</figureCitation>
). Dark marks on dorsum not metallic until apical part S7. S8 yellowish basally laterally, otherwise brown laterally, bronzy black dorsally. S9 brown laterally, bronzy black dorsally with irregular bluish white lateral apical mark. S10 and anal appendages almost entirely bluish white. Ovipositor largely cream, not quite reaching level of tips of cerci apart from styles, which are brown.
</paragraph>
<paragraph id="CB6B36A2FFE43245FF3C1D89FBA3F868" blockId="8.[151,1436,1813,2019]" box="[199,1142,1956,1983]" pageId="8" pageNumber="379">Measurements (mm). Abdomen without anal appendages or ovipositor 36, Hw 22.5.</paragraph>
<paragraph id="CB6B36A2FFE43244FF3C1DE5FD3EFE94" blockId="8.[151,1436,1813,2019]" lastBlockId="9.[151,1436,153,323]" lastPageId="9" lastPageNumber="380" pageId="8" pageNumber="379">
<emphasis id="F9A0EAB0FFE43245FF3C1DE5FE70F834" bold="true" box="[199,421,1992,2019]" pageId="8" pageNumber="379">Notes on variation</emphasis>
. As in
<taxonomicName id="0CD44D21FFE43245FE021DE5FDA6F834" authorityName="Dow" authorityYear="2019" box="[505,627,1993,2019]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFE43245FE021DE5FDA6F834" box="[505,627,1993,2019]" italics="true" pageId="8" pageNumber="379">A. rigiketit</emphasis>
</taxonomicName>
the main variation in markings in both sexes is in those of the anterior part of the head. R
<subScript id="575034E7FFE53244FE981A86FEBEFF6F" attach="left" box="[355,363,170,184]" fontSize="6" pageId="9" pageNumber="380">4</subScript>
occasionally arises at, rather than distal to, the subnodus in one or more wings. A very few males have the central part of the posterior pronotal lobe produced into a very short horn-like structure. Two males (not among those mentioned in the previous sentence) have the rear lateral extremities of the posterior pronotal lobe only slightly angulated. The subbasal spine on the upper branch of the male cercus is sometimes just visible in a typical lateral view, rarely is it clearly visible in such a view.
</paragraph>
<caption id="9FAB662AFFE53244FF6C18ECFB61FD0E" ID-DOI="http://doi.org/10.5281/zenodo.3558217" ID-Zenodo-Dep="3558217" box="[151,1204,704,729]" httpUri="https://zenodo.org/record/3558217/files/figure.png" pageId="9" pageNumber="380" startId="9.[151,264,704,729]" targetBox="[248,1340,345,679]" targetPageId="9">
<paragraph id="CB6B36A2FFE53244FF6C18ECFB61FD0E" blockId="9.[151,1204,704,729]" box="[151,1204,704,729]" pageId="9" pageNumber="380">
<emphasis id="F9A0EAB0FFE53244FF6C18ECFE81FD0E" bold="true" box="[151,340,704,729]" pageId="9" pageNumber="380">FIGURES 29–30.</emphasis>
Habitus of
<taxonomicName id="0CD44D21FFE53244FE3018ECFD7FFD0F" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" box="[459,682,704,728]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFE53244FE3018ECFD7FFD0F" box="[459,682,704,728]" italics="true" pageId="9" pageNumber="380">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFE53244FD4B18EDFCD7FD0F" box="[688,770,705,728]" pageId="9" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
: (29) holotype male; (30) female paratype.
</paragraph>
</caption>
<caption id="9FAB662AFFE53244FF6C1DA0FEC1F83A" ID-DOI="http://doi.org/10.5281/zenodo.3558219" ID-Zenodo-Dep="3558219" httpUri="https://zenodo.org/record/3558219/files/figure.png" pageId="9" pageNumber="380" startId="9.[151,250,1932,1957]" targetBox="[261,1329,751,1907]" targetPageId="9">
<paragraph id="CB6B36A2FFE53244FF6C1DA0FEC1F83A" blockId="9.[151,1436,1932,2029]" pageId="9" pageNumber="380">
<emphasis id="F9A0EAB0FFE53244FF6C1DA0FEF5F872" bold="true" box="[151,288,1932,1957]" pageId="9" pageNumber="380">FIGURE 31.</emphasis>
Distributions of
<taxonomicName id="0CD44D21FFE53244FE351DA1FD7BF872" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" box="[462,686,1933,1957]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFE53244FE351DA1FD7BF872" box="[462,686,1933,1957]" italics="true" pageId="9" pageNumber="380">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFE53244FD4E1DA1FCD1F873" box="[693,772,1933,1956]" pageId="9" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
(red circle) and
<taxonomicName id="0CD44D21FFE53244FC561DA1FB5EF872" authorityName="Laidlaw" authorityYear="1912" box="[941,1163,1933,1957]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE53244FC561DA1FB5EF872" box="[941,1163,1933,1957]" italics="true" pageId="9" pageNumber="380">Amphicnemis remiger</emphasis>
</taxonomicName>
(yellow square). Note that the problematic record of
<taxonomicName id="0CD44D21FFE53244FE5E1D9DFDC4F81E" authorityName="Laidlaw" authorityYear="1912" box="[421,529,1969,1993]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE53244FE5E1D9DFDC4F81E" box="[421,529,1969,1993]" italics="true" pageId="9" pageNumber="380">A. remiger</emphasis>
</taxonomicName>
from “Murud” (see the Remarks under
<taxonomicName id="0CD44D21FFE53244FC4A1D9DFBCBF81E" authorityName="Laidlaw" authorityYear="1912" box="[945,1054,1969,1993]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE53244FC4A1D9DFBCBF81E" box="[945,1054,1969,1993]" italics="true" pageId="9" pageNumber="380">A. remiger</emphasis>
</taxonomicName>
) is excluded from the distribution of that species.
</paragraph>
</caption>
<paragraph id="CB6B36A2FFE63247FF3C1AB5FF2FFF2C" blockId="10.[151,1437,152,1223]" pageId="10" pageNumber="381">
Measurements (mm). Males: 11–14 Px in Fw,
<quantity id="0C2C9B47FFE63247FD191AB5FC92FF64" box="[738,839,153,179]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="10" pageNumber="381" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">10–12 in</quantity>
Hw, abdomen without anal appendages 30–35, Hw 18–21. Females: 11–14 Px in Fw,
<quantity id="0C2C9B47FFE63247FDD21A91FD45FF00" box="[553,656,189,215]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="10" pageNumber="381" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">10–12 in</quantity>
Hw, abdomen without anal appendages or ovipositor 33–35.5, Hw 21–21.5.
</paragraph>
</subSubSection>
<subSubSection id="83CE6529FFE63247FF3C1B29FF09FB10" pageId="10" pageNumber="381" type="discussion">
<paragraph id="CB6B36A2FFE63247FF3C1B29FB30FE20" blockId="10.[151,1437,152,1223]" pageId="10" pageNumber="381">
<emphasis id="F9A0EAB0FFE63247FF3C1B29FEE1FEC8" bold="true" box="[199,308,261,287]" pageId="10" pageNumber="381">Remarks</emphasis>
. The
<typeStatus id="146F8800FFE63247FE8C1B29FE0CFEC8" box="[375,473,261,287]" pageId="10" pageNumber="381" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0CD44D21FFE63247FDFB1B29FDACFEC8" authorityName="Laidlaw" authorityYear="1912" box="[512,633,261,287]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE63247FDFB1B29FDACFEC8" box="[512,633,261,287]" italics="true" pageId="10" pageNumber="381">A. remiger</emphasis>
</taxonomicName>
is from somewhere in the vicinity of the mouth of the Madalam River (Kuala Madalam), a tributary of the Limbang River, in Sarawak’s Limbang Division (
<bibRefCitation id="AF454B53FFE63247FBA51B05FAD5FE94" author="Laidlaw, F. F." box="[1118,1280,297,323]" pageId="10" pageNumber="381" pagination="92 - 99" refId="ref6499" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99., 1 pl." type="journal article" year="1912">Laidlaw 1912</bibRefCitation>
), collected on the
<collectingCountry id="B3C37632FFE63247FF3B1B61FEF5FEB0" box="[192,288,333,359]" name="Malaysia" pageId="10" pageNumber="381">Sarawak</collectingCountry>
Museum’s 1911 expedition to Mount Batu Lawi (
<bibRefCitation id="AF454B53FFE63247FCB01B61FC25FEB0" author="Moulton, J. C." box="[843,1008,333,359]" pageId="10" pageNumber="381" pagination="1 - 58" refId="ref6783" refString="Moulton, J. C. (1912) An expedition to Mount Batu Lawi. Journal of the Straits Branch of the Royal Asiatic Society, 63, 1 - 58." type="journal article" year="1912">Moulton 1912</bibRefCitation>
). Later
<bibRefCitation id="AF454B53FFE63247FBBF1B61FACEFEB0" author="Laidlaw, F. F." box="[1092,1307,333,359]" pageId="10" pageNumber="381" pagination="63 - 80" refId="ref6564" refString="Laidlaw, F. F. (1913) Contributions to a study of the dragonfly fauna of Borneo. Part I. The Corduliinae: The genus Amphicnemis: The legion Protoneura. Proceedings of the Zoological Society of London, 1913, 63 - 80, pl. 4 excl. https: // doi. org / 10.1111 / j. 1096 - 3642.1913. tb 01984. x" type="journal article" year="1913">Laidlaw (1913: 72)</bibRefCitation>
mistakenly gave the type locality as Batu Lawi, possibly believing that Kuala Madalam is actually in the vicinity of the mountain (actually it is more than
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distant from it). I have compared recently collected specimens of
<taxonomicName id="0CD44D21FFE63247FADF1BB9FA48FE78" authorityName="Laidlaw" authorityYear="1912" box="[1316,1437,405,431]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE63247FADF1BB9FA48FE78" box="[1316,1437,405,431]" italics="true" pageId="10" pageNumber="381">A. remiger</emphasis>
</taxonomicName>
directly against the
<typeStatus id="146F8800FFE63247FE8E1B95FE02FE04" box="[373,471,441,467]" pageId="10" pageNumber="381" type="holotype">holotype</typeStatus>
(in the Natural History Museum, London) to confirm their identity. The
<typeStatus id="146F8800FFE63247FAF71B95FABBFE04" box="[1292,1390,441,467]" pageId="10" pageNumber="381" type="holotype">holotype</typeStatus>
is a slightly immature individual, so a fresh mature male specimen was used for the description above.
</paragraph>
<paragraph id="CB6B36A2FFE63247FF3C182DFE11FD50" blockId="10.[151,1437,152,1223]" pageId="10" pageNumber="381">
The type locality is at low altitude and it is likely that the
<typeStatus id="146F8800FFE63247FCBC182DFC7CFDCC" box="[839,937,513,539]" pageId="10" pageNumber="381" type="holotype">holotype</typeStatus>
was collected in swamp forest of some type. Most subsequent records, where the habitat is recorded, are also from swamp forest, often peat swamp forest, or streams in swamp or alluvial forest; my own specimens are from such habitats.
<bibRefCitation id="AF454B53FFE63247FBE81865FB11FDB4" author="Orr, A. G." box="[1043,1220,585,611]" pageId="10" pageNumber="381" pagination="167 - 220" refId="ref6844" refString="Orr, A. G. (2001) An annotated checklist of the Odonata of Brunei with ecological notes and descriptions of hitherto unknown males and larvae. International Journal of Odonatology, 4 (2), 167 - 220. https: // doi. org / 10.1080 / 13887890.2001.9748168" type="journal article" year="2001">Orr (2001: 183)</bibRefCitation>
states “common in swampy areas” for
<collectingCountry id="B3C37632FFE63247FE8B1841FE15FD50" box="[368,448,621,647]" name="Brunei" pageId="10" pageNumber="381">Brunei</collectingCountry>
.
</paragraph>
<paragraph id="CB6B36A2FFE63247FF3C18BDFDC9FC7F" blockId="10.[151,1437,152,1223]" pageId="10" pageNumber="381">
There is a rather puzzling record of this species in
<bibRefCitation id="AF454B53FFE63247FD0B18BDFC01FD7B" author="Laidlaw, F. F." box="[752,980,657,684]" pageId="10" pageNumber="381" pagination="311 - 342" refId="ref6636" refString="Laidlaw, F. F. (1920) Contributions to a study of the dragonfly fauna of Borneo. Part IV. A list of species known to occur in the island. Proceedings of the Zoological Society of London, 1920, 311 - 342. https: // doi. org / 10.1111 / j. 1469 - 7998.1920. tb 07074. x" type="journal article" year="1920">Laidlaw (1920: 336)</bibRefCitation>
where as well as again incorrectly giving the
<typeStatus id="146F8800FFE63247FF3A1899FF24FD18" box="[193,241,693,719]" pageId="10" pageNumber="381">type</typeStatus>
locality as Batu Lawi, he lists two males, stated to be “imperfect”, from “Murud”, collected on the 20
<superScript id="3CA19BEAFFE63247FA8A1898FAABFD15" attach="left" box="[1393,1406,692,706]" fontSize="6" pageId="10" pageNumber="381">th</superScript>
of
<date id="BF6A1062FFE63247FF6C18F5FE9EFD24" box="[151,331,729,755]" pageId="10" pageNumber="381" value="1914-12">December 1914</date>
. Murud presumably refers to Gunung Murud, the highest mountain in
<collectingCountry id="B3C37632FFE63247FBAC18F5FB6EFD24" box="[1111,1211,729,755]" name="Malaysia" pageId="10" pageNumber="381">Sarawak</collectingCountry>
. Collectors from the
<collectingCountry id="B3C37632FFE63247FF6C18D1FF22FCC0" box="[151,247,765,791]" name="Malaysia" pageId="10" pageNumber="381">Sarawak</collectingCountry>
Museum went to Gunung Murud, in late 1914 (
<bibRefCitation id="AF454B53FFE63247FCF418D1FC66FCC0" author="Moulton, J. C." box="[783,947,764,791]" pageId="10" pageNumber="381" refId="ref6816" refString="Moulton, J. C. (1915) Twelfth report on the Sarawak Museum, 1914. Sarawak Government Printing Office, Kuching, 48 pp." type="book" year="1915">Moulton 1915</bibRefCitation>
), but it is not clear if they were still there in December and these specimens may have been collected somewhere on their route back. If these specimens were actually collected in the highlands it seems doubtful that they were actually
<taxonomicName id="0CD44D21FFE63247FC001969FBA6FC88" authorityName="Laidlaw" authorityYear="1912" box="[1019,1139,837,863]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE63247FC001969FBA6FC88" box="[1019,1139,837,863]" italics="true" pageId="10" pageNumber="381">A. remiger</emphasis>
</taxonomicName>
and might actually be the unnamed
<taxonomicName id="0CD44D21FFE63247FEFE1945FE49FC54" authorityName="Selys" authorityYear="1863" box="[261,412,873,899]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="F9A0EAB0FFE63247FEFE1945FE49FC54" box="[261,412,873,899]" italics="true" pageId="10" pageNumber="381">Amphicnemis</emphasis>
</taxonomicName>
species recorded in
<bibRefCitation id="AF454B53FFE63247FD841945FC90FC54" author="Dow, R. A. &amp; Reels, G. T. &amp; Ngiam, R. W. J." box="[639,837,872,899]" pageId="10" pageNumber="381" pagination="1 - 17" refId="ref6386" refString="Dow, R. A., Reels, G. T. &amp; Ngiam, R. W. J. (2015) Odonata collected at Usun Apau National Park, Miri Division, Sarawak, Malaysia in April and May 2012. International Dragonfly Fund Report, 79, 1 - 17." type="journal article" year="2015">
Dow
<emphasis id="F9A0EAB0FFE63247FD471946FD38FC54" box="[700,749,873,899]" italics="true" pageId="10" pageNumber="381">et al</emphasis>
. (2015)
</bibRefCitation>
from Usun Apau and the Tama Abu Range (the range to which Gunung Murud belongs).
</paragraph>
<paragraph id="CB6B36A2FFE63247FF3C199DFC7EFB8C" blockId="10.[151,1437,152,1223]" pageId="10" pageNumber="381">
Laidlaw’s description of the male anal appendages of
<taxonomicName id="0CD44D21FFE63247FCD3199DFC75FC1C" authorityName="Laidlaw" authorityYear="1912" box="[808,928,945,971]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE63247FCD3199DFC75FC1C" box="[808,928,945,971]" italics="true" pageId="10" pageNumber="381">A. remiger</emphasis>
</taxonomicName>
is somewhat misleading. He states, referring to the upper branch of the cercus as the upper pair of anal appendages, “The shaft a little bowed with a small tooth at its middle” (
<bibRefCitation id="AF454B53FFE63247FEC619D5FDDDFBC3" author="Laidlaw, F. F." box="[317,520,1017,1044]" pageId="10" pageNumber="381" pagination="92 - 99" refId="ref6499" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99., 1 pl." type="journal article" year="1912">Laidlaw 1912: 96</bibRefCitation>
). However, the illustration of the anal appendages of the
<typeStatus id="146F8800FFE63247FB7C19D5FB3CFBC4" box="[1159,1257,1017,1043]" pageId="10" pageNumber="381" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="AF454B53FFE63247FB0319D5FA4DFBC4" author="Laidlaw, F. F." box="[1272,1432,1017,1043]" pageId="10" pageNumber="381" pagination="92 - 99" refId="ref6499" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99., 1 pl." type="journal article" year="1912">Laidlaw 1912</bibRefCitation>
:
<figureCitation id="53EF2A27FFE63247FF6C1E31FF0FFBE0" box="[151,218,1053,1079]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1207,1232]" captionTargetBox="[151,1437,179,1179]" captionTargetId="figure@2.[151,1437,179,1179]" captionTargetPageId="2" captionText="FIGURES 1–4. Amphicnemis rigiketit sp. nov., head of holotype male: (1), dorsal-frontal view; (3) dorsal view. Head of male Amphicnemis remiger (SAR11_12_COE332); (2) dorsal-frontal view; (4) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558203" httpUri="https://zenodo.org/record/3558203/files/figure.png" pageId="10" pageNumber="381">Fig. 4</figureCitation>
) correctly shows the tooth in a subbasal position, as in all specimens that I have examined (including that illustrated here), although the tooth is only rarely visible in lateral view.
</paragraph>
<paragraph id="CB6B36A2FFE63247FF3C1E49FF09FB10" blockId="10.[151,1437,152,1223]" pageId="10" pageNumber="381">
<taxonomicName id="0CD44D21FFE63247FF3C1E49FE6BFBA8" authorityName="Laidlaw" authorityYear="1912" box="[199,446,1125,1151]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE63247FF3C1E49FE6BFBA8" box="[199,446,1125,1151]" italics="true" pageId="10" pageNumber="381">Amphicnemis remiger</emphasis>
</taxonomicName>
appears to have a rather restricted distribution in
<collectingCountry id="B3C37632FFE63247FC091E49FB94FBA8" box="[1010,1089,1125,1151]" name="Brunei" pageId="10" pageNumber="381">Brunei</collectingCountry>
, immediately adjacent parts of
<collectingCountry id="B3C37632FFE63247FF6C1EA5FF22FB74" box="[151,247,1161,1187]" name="Malaysia" pageId="10" pageNumber="381">Sarawak</collectingCountry>
and extending to the south-west of
<collectingRegion id="0910F840FFE63247FD741EA5FD06FB74" box="[655,723,1161,1187]" country="Malaysia" name="Sabah" pageId="10" pageNumber="381">Sabah</collectingRegion>
(
<figureCitation id="53EF2A27FFE63247FD1F1EA5FCECFB74" box="[740,825,1161,1187]" captionStart="FIGURE 31" captionStartId="9.[151,250,1932,1957]" captionTargetBox="[261,1329,751,1907]" captionTargetId="figure@9.[259,1329,750,1907]" captionTargetPageId="9" captionText="FIGURE 31. Distributions of Amphicnemis rigiketit sp. nov. (red circle) and Amphicnemis remiger (yellow square). Note that the problematic record of A. remiger from “Murud” (see the Remarks under A. remiger) is excluded from the distribution of that species." figureDoi="http://doi.org/10.5281/zenodo.3558219" httpUri="https://zenodo.org/record/3558219/files/figure.png" pageId="10" pageNumber="381">Fig. 31</figureCitation>
). In
<collectingCountry id="B3C37632FFE63247FC961EA5FC18FB74" box="[877,973,1161,1187]" name="Malaysia" pageId="10" pageNumber="381">Sarawak</collectingCountry>
it has not been found west of the Baram River.
</paragraph>
</subSubSection>
</treatment>
</document>
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